Richard SpellenbergNew Mexico State University | NMSU · Department of Biology
Richard Spellenberg
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Publications (58)
Eleven angiosperm plant species are added to the flora of New Mexico that were revealed through herbarium curation associated with a project to digitize and unify the two herbaria of New Mexico State University (NMC & NMCR) and through ongoing field work. One, Dimorphocarpa candicans, is restored to the state’s flora, after it was earlier placed in...
Astragalus pilosior Spellenb. & E.W. Anderson is described and illustrated as a new species. It is known to occur in the pine forests of southwestern Chihuahua, Mexico, on the western slopes of the Sierra Madre Occidental between approximately 1900 m and 2500 m elevation. The new species is placed in the section Strigulosi. It is compared with the...
Observations of the Sierra Madrean oak Quercus viminea in ten Sky Island mountain ranges 15 to 85 km south of the Arizona and New Mexico border in Sonora and Chihuahua help define the northern and western limits of its distribution. We also discovered it in the Patagonia Mountains of southern Arizona as far north as 15.3 km from Sonora. It is sympa...
Quercus undata Trel. (Fagaceae, Quercus, Section Quercus) has a complex taxonomic and nomenclatural history. Intensive sampling of oaks at the type locality of Q. undata Trel. in Durango, Mexico and evaluation of herbarium specimens and plants in the field indicate that Q. undata represents variation in Quercus chihuahuensis Trel. in white oak comm...
Quercus undata Trel. (Fagaceae, Quercus, Section Quercus) has a complex taxonomic and nomenclatural history. Intensive sampling of oaks at the type locality of Q. undata Trel. in Durango, Mexico and evaluation of herbarium specimens and plants in the field indicate that Q. undata represents variation in Quercus chihuahuensis Trel. in white oak comm...
A hybrid between Oryzopsis hendersonii and Stipa lemmonii is reported and formally described. In a population in central Washington where the two parent species are sympatric 12 hybrid plants were found. The hybrids are morphologically and anatomically intermediate; O. hendersonii, which occupies drier sites, has xeromorphic modifications in the le...
Miscellaneous comments regarding nomenclature and typification are made for the Nyctaginaceae of the Intermountain region of the western United States. Tripterocalyx carneus (Greene) Galloway var. pedunculatus (M E. Jones) Spellenb. is presented as a new combination. Abroma crux-maltac Kellogg is neotypified. Boerhavia spkata Choisy var torreyana S...
Recent phylogenetic work shows that existing tribal concepts within Nyctaginaceae are incompatible with the principle of recognizing monophyletic taxa. We review the history of supergeneric classification in Nyctaginaceae, clarify issues pertaining to priority of certain generic names, and discuss the application of the conserved family name to the...
Recent phylogenetic work shows that existing tribal concepts within Nyctaginaceae are incompatible with the principle of recognizing monophyletic taxa. We review the history of supergeneric classification in Nyctaginaceae, clarify issues pertaining to priority of certain generic names, and discuss the application of the conserved family name to the...
Se describe la especie nueva Dalea analiliana de los alrededores de Yécora en la Sierra Madre Occidental al oriente de Sonora, México. Esta planta perenne está relacionada con D. nelsonii, pero se diferencia por sus flores blancas y la presencia de glándulas en ambas superficies de las hojuelas y su rango geográfico más al norte. Dalea analiliana o...
The name for the poorly known, narrowly distributed, Boerhavia triquetra S. Wats, predates the name B. intermedia M.E. Jones, a well-known widespread species of the arid American Southwest. A survey of herbarium specimens and recently collected specimens from the type locality of B. triquetra at Bahia de los Angeles in Baja California, shows the tw...
For the upcoming treatment of the genus Mirabilis for the Flora of North America North of Mexico, M. linearis (Pursh) Heimerl is treated as having three thoroughly intergrading varieties: M. linearis var. linearis, M. linearis var. decipiens (Standley) S. L. Welsh, and M. linearis var. subhispida (Heimerl) Spellenberg, a new combination published h...
A new combination is made for A Flora of the Chihuahuan Desert Region and for Flora of North America North of Mexico in the now more broadly constructed Acleisanthes (Nyctaginaceae): Acleisanthes lanceolata (Wooton) R.A. Levin var. megaphylla (Fowler & B.L. Turner) Spdlenb. & J. Poole, comb. nov. This is based on evidence published recently by Levi...
Strobilanthinae comprise approximately 350 species from south and southeast Asia and Melanesia. Three main classifications of Strobilanthinae have been advocated, differing markedly in the number of groups recognised, their circumscription, and the rank assigned to them. But there remains no consensus concerning the best approach to the classificat...
Boerhavia coulteri (Hook. f.) S. Wats. var. palmeri (S. Wats.) Spellenb., comb. nov., based on B. spicata Choisy var. palmeri S. Wats., from the vicinity of Guaymas, Sonora, Mexico, is proposed for a Boerhavia that differs from the var. coulteri in its small fruits and flowers, and its with sparsely-fruited spicate inflorescences. The variety range...
A revision of Mirabilis, section Oxybaphoides, Nyctaginaceae, in western North America is presented. Mirabilis oligantha (Standl.) J.F. Macbride, M. oxybaphoides (A. Gray) A. Gray, and M. tenuiloba S. Wats remain as traditionally classified. Mirabilis bigelovii A. Gray, M. californica A. Gray ex Torr., M. laevis (Benth.) Curran, and M. retrorsa Hel...
Familia de 26 a 31 géneros con 300 a 400 especies distribuidas sobre todo en regiones tropicales y subtropicales de América, pero bien representada asimismo en el Antiguo Mundo. El género Boerhavia incluye malezas de repartición pantropical. Su importancia económica no es muy grande. Varios representantes se usan localmente en medicina popular para...
Observations are reported on anthesis, stamen and style movement, and insect visitation in five taxa of Boerhavia, one a pantropical perennial (B. coccinea Mill.), and five that are North American annuals (B. intermedia M.E. Jones, B. spicata Choisy, B. torreyana S. Wats., and B. wrightii A. Gray). Observations were made in natural situations in La...
A survey of the flora on a hitherto unexplored gypsum outcrop on the west face of the Guadalupe Mountains in southern New Mexico has revealed a new variety of Anulocaulis (Nyctaginaceae), A. leiosolenus (Tort.) Standl.var. howardii Spellenb. and Wootten (var. nov.). This variety has a combination of characteristics in various degrees intermediate t...
Quercus gambelii Nutt. es un encino blanco (Quercus, subgénero y sección Quercus, subsección Gambelieae) común en la parte sur de la región de las Montañas Rocosas en los Estados Unidos y distribuido hasta la parte central de la porción chihuahuense de la Sierra Madre Occidental. Quercus arizonica Sarg., lejanamente relacionado con el anterior (sub...
A taxonomy of a small section of Mexican black oaks (Quercus, section Lobatae, subsection Racemiflorae) is presented. Four species are recognized; Q. conzattii, Q. radiata, Q. tarahumara, Q. urbanii. All are annual-fruited, the acorns in racemose inflorescences ranging from condensed or reduced to elongate, the species differing by characteristics...
Populations of hybrids of distantly related black oaks (Fagaceae: Quercus: section Lobatae), Quercus conzatti Trelease and Quercus eduardii Trelease, are examined throughout the southern Sierra Madre Occidental of Mexico, where the distributions of the two parental species overlap. Morphometric analyses are presented to show the structure of a hybr...
Se encontraron 20 nuevos registros repartidos en 13 familias para las montañas del norte de México. La mayor parte de estos hallazgos se localizaron en la zona del Parque Nacional ¿Cascada de Basaseachic¿, Chihuahua, y son el resultado de los estudios florísticos llevados a cabo por los autores en los últimos siete años.
Quercus mcvaughii is described as a new species of black oak (subgenus Erythrobalanus) from the Sierra Madre Occidental, in northern Durango and western Chihuahua, Mexico. It is most closely related to the more southern Q. crassifolia. Sessile or subsessile hairs, usually of different sizes, on the abaxial leaf surface distinguish Q. mcvaughii from...
Se dan a conocer los números cromosómicos de 18 taxa de Astragalus (Fabaceae) del norte de México y el sureste de los Estados Unidos. Los datos correspondientes a las 12 especies siguientes se revelan por primera vez: CONTEOS PROVENIENTES DE BOTONES FLORALES: (n=11) A. cynaeus, A. eremiticus, A. insulares var. harwoodii, A. proximus; (n=14) A. hart...
We examined patterns of density and species diversity for leaf-mining Lepidopterans and gall-forming Hymenopterans in two oak (Quercus spp.) hybrid zones: Quercus depressipes x Q. rugosa and Q. emoryi x Q. coccolobifolia. In both species complexes, hybrid hosts typically supported significantly lower densities and species diversity of parasites tha...
Cuando Trelease describió la variedad Quercus oblongifolia Torr. var. pallidinervis Trel. Indicó que el tipo se había fotografiado en el herbario “Washington”, pero la muestra que ilustró se descubrió recientemente en el Herbario Gray. Por consiguiente esta muestra representa el holotipo; un ejemplar diferente de la misma colección se encuentra en...
Perityle vigilans is described from the Sierra Madre Occidental of southwestern Chihuahua. It differs from other Perityle by the combination of its white ligules, sparsely setosehispid achenes, finely grayish-villosulous tomentum, and absence of pappus bristles. The new species is believed to be most closely related to P. rosei and P. trichodonta,...
The name Eriogonum densum Greene, as presently applied, refers to one of the rarest species in New Mexico. It is known from only two collections consisting of 16 plants. The name also has been applied in a much broader sense to plants that are now included in the common Eriogonum palmerianum Reveal in Munz, a species somewhat similar in habit but d...
Astragalus chuskanus Barneby & Spellenberg, a new species of ser. Humistratae, is described, figured, discussed, and inserted into current monographic keys to the genus.
Chaetopappa elegans, a narrow endemic from the White Mountains of south-central New Mexico, is unusual in Chaetopappa because of its double pappus of numerous bristles and narrow scales, comparatively long, lanceolate stylar appendages, crinkled stem pubescence, alveo- late receptacle, faint cypsela nerves, long ligules, and occurrence on granitic...
Dalea formosa consists of diploids (n = 7), tetraploids (n = 14), and hexaploids (n = 21), the polyploids restricted to the Chihuahuan Desert region or its immediate borders. There is very little morphological
differentiation between the three chromosome races and, therefore, the polyploids are assumed to be primarily autoploid. Tetraploids
discove...
The complete floras of three Chihuahuan desert plant communities were surveyed to determine the relative numbers of species that possess either the C<sub>3</sub> C<sub>4</sub> or CAM carbon reduction pathway. There were more C<sub>3</sub> species in all three communities. Spring-flowering species were predominantly C<sub>3</sub> and CAM; C<sub>4</s...
This study presents 126 original reports of chromosome numbers for 101 taxa of North American Astragalus; 46 are the first records for species. Chromosome numbers are now known from at least one count for slightly more than 40% of the North American species; on a world wide basis about 80% of the species are still unknown from a single count. This...
A predominantly autogamous breeding system is described forP. occidentale Scribn.,P. pacificum Hitchc. & Chase, andP. thermale Boland. Five synthetic hybrids and three generations of progeny are also described for crosses among these three species.
All parents haven = 9 chromosomes and normal meiosis. Hybrids exhibit various irregularities during m...
Chromosome numbers of n = 1 are reported for six species of Astragalus, section Strigulosi. The chromosome number allies the section with New World groups of Astragalus, (mostly n = 11 or 12) and not Old World groups (x = 8). Descriptive notes are added for the heretofore rare A. altus.
Natural and synthetic hybrids are reported betweenPanicum, scribnerianum Nash and two other species,P. pacificum Hitchc. & Chase andP. occidentale Scribn. & Merr. The hybrids are morphologically intermediate between the parents, and those in whichP. pacificum is one parent strongly resemble the rareP. shastense Scribn. & Merr. All the parent plants...
Adenophyllum porophyllum (Cav.) Hemsl. var. porophyllum is reported for Chihuahua, its presence there overlooked in a recent revision. Adenophyllum wrightii A. Gray var. wrightii is reported from two new locations, confirming a single century old record for Chihuahua. Astragalus ervoides Hook. & Arn. var. maysillesii Barneby is reported as a new re...
Thesis (Ph. D.)--University of Washington, 1968. Includes bibliographical references (leaves 161-165). Microfilm. s
A new species of ricegrass,Oryzopsis swallenii, from the sagebrush hills at the east end of the Lost River and Lemhi ranges in east central Idaho is described. The plant
combines features ofStipa andOryzopsis, but is set apart fromStipa by virtue of a quickly deciduous awn and a blunt callus, and from other species inOryzopsis by its relatively lon...