Olof Ryding

• PhD
• Manager at Natural History Museum of Denmark

The name Nepeta nepetellae has been accepted in Carl Christensen's standard index to Forsskål's Flora Aegyptiaco-Arabica, listed in Index Kewensis and IPNI, listed as an accepted name in The Plant List and accepted in Hepper and Friis's revision of the plants of Forsskål's Flora Aegyptiaco-Arabica. If accepted, N. nepetellae may threaten names in c...

We have investigated the phylogenetic position of Stachys tibetica by use of both plastid (rps16 intron, trnL-F region and matK) and nuclear ribosomal (ITS) DNA sequence data and both parsimony and Bayesian phylogenetic methods. All data corroborate that S. tibetica is extraneous in Stachys (tribe Stachydeae) and suggest a phylogenetic affiliation...

This is the first comprehensive molecular investigation of the genus Lamium L. We have addressed phylogenetic relationships and presumed allopolyploid speciation by use of nuclear (NRPA2, 5S-NTS) and chloroplast (matK, psbA-trnH, rps16, trnL, trnL-F, trnS-G) DNA sequence data. Nuclear and chloroplast data were incongruent, and nuclear data showed b...

Lamioideae comprise the second-largest subfamily in Lamiaceae. Although considerable progress has recently been made in Lamioideae phylogenetics, the subfamily remains one of the most poorly investigated subfamilies in Lamiaceae. Here we present a taxonomic update of the subfamily based on earlier published data as well as 71 new DNA extracts from...

The diversity of crystal types is studied in fruiting calyces of 435 out of the c. 7,200 species, and 184 out of the c. 236 genera of Lamiaceae, mainly in the subfamilies Nepetoideae and Lamioideae. Calcium oxalate crystals are found in about half of the studied calyces, and belong to the following main types: prismatics in the inner epidermis, pri...

This paper presents a phylogenetic analysis of Lamiaceae subfam. Lamioideae (including subfamily Pogostemonoideae) based on sequences of the trnL intron, trnL-trnF intergenic spacer, and rps16 intron of the plastid genome. It is the first analysis that includes all major lamioid and pogostemonoid genera. Monophyly of Lamioideae s.l. (i.e., includin...

In 1776, Rottb0ll published 12 species from Rolander's collection of plants of Suriname. Here, we update the taxonomic identity of those species, discussing, whenever pertinent, their nomenclatural status. Six of the 12 names and also two of their synonyms, one the name of a species described by Linné filius and another described by Kunth, are lect...

Pericarp structure is investigated in 143 specimens belonging to 59 out of ca. 65 genera and 134 out of ca. 2,000 species of Lamiaceae subfamily Nepetoideae tribe Mentheae. Some of the data presented by earlier authors are believed to be incorrect. The variation in pericarp structure has been found to be strongly correlated to the variations in DNA...

Pericarp and seed structure was investigated in 13 out of the 16 species of the old Caryopteris s.l. and in a few species of the related Rubiteucris, Amethystea, Trichostema and Ajuga. The pericarp structure was found to be very variable, and the variation largely matches the most recent classification where Caryopteris s.l. is divided into six gen...

Pericarp structure has been investigated in 10 out of the c. 20 species of the genus Monarda in Lamiaceae subfamily Nepetoideae tribe Mentheae subtribe Menthinae. Old data showing that the pericarp structure of Monarda is extremely variable and partly matches the pericarp of the genus Salvia (in subtribe Salviinae) is found to be incorrect. Instead...

No comprehensive revision of Micromeria is available and uncertainties about the taxonomy of the genus have lasted for a long time. Since the last synopsis many new data from both morphological and molecular genetic studies have been accumulated and, consequently, the number of accepted taxa and the delimitation of the genus have changed considerab...

Pericarp structure has been investigated in 52 species belonging to the closely related genera Phlomis s. lat. (incl. Phlomoides), Eremostachys, Lamiophlomis and Notochaete. The new data supports the division of the Phlomis group into two main groups: 1) a group of Phlomis sect. Phlomis; 2) a group of Eremostachys, Notochaete, Lamiophlomis and Phlo...

A new species from gypsum plains in northern Somalia, Otostegia ericoidea Ryding is described and illustrated.

The material of Micromeria s.s. (M. sect. Micromeria) from tropical to southern Africa and the Arabian Peninsula is revised. The endemic Ethiopian M. unguentaria Schweinf. is retained as a species, while the other African and Arabian material of Micromeria is included in M. imbricata (Forssk.) C. Chr. M. imbricata is divided into the following thre...

Calyces of 306 species representing 181 out of the 236 genera of Lamiaceae have been studied with particular emphasis on the amount of fibres and similar xylem cells. A characteristic calyx tissue called ``mesophyll fibres'', reported by earlier authors, is found to belong to the xylem. Most species of the subfamilies Lamioideae and Scutellarioidea...

A tropical African group of species commonly included in the genus Satureja is revised and moved to the genus Clinopodium. Satureja abyssinica (Benth.) Briq. ssp. abyssinica and ssp. condensata (Hedberg) Seybold, S. paradoxa (Vatke) Engl. ex Seybold, S. robusta (Hook.f) Brenan and S. vernayana Brenan should be known under the following names: Clino...

An afromontane to afroalpine group of species commonly included in the genus Satureja is studied by biometrical methods and revised. Satureja simensis (Benth.) Briq., S. kilimandschari (Gürke) Hedberg, S. uhligii Gürke and S. pseudosimense Brenan should be known under the following names: Clinopodium simense (Benth.) Kuntze, C. kilimandschari (Gürk...

Plectranthus igniarioides Ryding, a new species from northern Somalia, is described and illustrated. Comparison is made with P. argentifolius Ryding, P. bariensis Ryding, P. cuneatus (E. G. Baker) Ryding, P. gillettii, J. K. Morton, P. igniarius (Schweinfurth) Agnew, and P. puberulentus J. K. Morton. The circumscription of Plectranthus gillettii, w...

Trees, shrubs, subshrubs or perennial or annual herbs, rarely climbers, aromatic or not. Roots rarely tuberous. Stems often quadrangular, erect to prostrate, somtimes forming usually present, of glandular, rarely scale-like, usually multicellular-uniseriate, simple, branched, dendroid or stellate, sometimes gland-tipped, large-headed subsessile gla...

The interrelationship of the genus Lamium is investigated using cladistic analysis. The result suggests that the European to West Asiatic species of Lamium (excl. L. galeobdolon (L.) L.) form a monophyletic group together with the genus Wiedemannia, and that Lamium is paraphyletic with Wiedemannia as a subgroup. In agreement with this result, Wiede...

Several new combinations and new synonomies are made affecting the following genera of Labiatae: Keiskea, Collinsonia, Chingyungia, Perrierastrum, Plectranthus, Nepeta and Pitardia. In addition, subtribe Hanceolinae (tribe Ocimeae) is recognized. This includes Isodon, Hanceola and Siphocranion.

The endemic and monotypic Somali genus Puntia is included in Endostemon sect. Diffusi. The new combination E. stenocaulis (Hedge) Ryding, A. J. Paton & Thulin is made, and a new species of the same section, E. racemosus Ryding, A. J. Paton & Thulin, is described. A detailed description of pericarp structures in E. racemosus is also given.

This article provides a short life history of the Danish count and amateur botanist Frederik Christian Raben, giving a short account of his journey in Brazil, a history of his Brazilian herbaria, and explanations of the annotations on his specimens. A few years after his death, F. C. Raben's Brazilian herbarium was divided into different sets. Spec...

The variable tropical African Aeollanthus myrianthus Baker is here divided into two subspecies mainly on the basis of differences in the length of the corolla-tube. Subsp. myrianthus represents the plants with a short corolla tube. The new combination, subsp. gamwelliae (G. Taylor) Ryding, represents the plants with a long corolla tube. The identit...

Data on the amount of mucilage in mericarps of 400 species of Lamiaceae subfam. Nepetoideae has been obtained by tests and gathered from literature. The relative frequency of myxodiaspory (the condition of having mucilaginous diaspores) has been calculated for groups of species with similar biological characteristics, and the differences tested for...

Ocimum α Zatarhendi Forssk., O. aegyptiacum Forssk. and Plectranthus crassifolius Vahl have been identified as conspecific with P. tenuiflorus (Vatke) Agnew. Ocimum a Zatarhendi Forssk. is regarded as not validly published, and P. aegyptiacus (Forssk.) C. Chr. is regarded as the correct name of the taxon. A neotype of O. aegyptiacum and a lectotype...

A new species from Ethiopia and the northern Kenya, Plectranthus minutiflorus is described and illustrated.

Pycnostachys recurvata, a new species from Ethiopia, is described and illustrated. Comparisons are made to P. pseudospeciosa and P. chevalieri, which it resembles by having recurved leaves, and to P. niamniamensis, P. reticulata, and P. deflexifolia, which it resembles in calyx shape.

The single species of the genus Bovonia, B. diphylla Chiov. from Congo-Kinshasa is identified; it is found to be conspecific to Aeollanthus homblei De Wild. The latter name has to be applied for this taxon. /// L'unique espèce du genre Bovonia, B. diphylla Chiov. du Congo(-Kinshasa) est identifiée; elle s'avère être la même espèce que Aeollanthus h...

The names Plectranthus barbatus Andrews and P. comosus Sims have been applied to different species (or different forms of the same species) and are here epitypified by collections from Eritrea and Nepal respectively. A collection assigned to P. barbatus sensu lato is selected as epitype for Coleus coerulescens Gürke. The material from Somalia and Y...

Teucrium (Lamiaceae) from NE tropical Africa is revised. Of the six species recognized, three new species are described and illustrated: T. clementiae Ryding from E Ethiopia and N Somalia, T. somalense Ryding from NW Somalia, and T. polioides Ryding from NE and C Somalia. Maps and a key to the species are also provided.

Interrelationships within the Leucas group have been investigated using cladistic analysis of gross morphological, anatomical and cytological data. Fifty species were sampled from Acrotome, Ballota, Isoleucas, Leonotis, Leucus, Moluccella and Otostegia. Different analyses were made, with and without continuous characters, and with and without chara...

The subtribes of the Ocimeae are discussed and new definitions of Plectranthinae and Ociminae are provided. The position of Hanceola, Siphocranion and Isodon within the tribe Ocimeae is discussed and the three genera are placed as incertae sedis. A key to the subtribes of the Ocimeae and genera incertae sedis are presented. The genus Skapanthus is...

Description d'une nouvelle espèce, Plectranthus globosus Ryding (Lamiaceae) du Zaïre. L'anatomie du péricarpe est incluse dans la description.

Pericarp structure was investigated in 158 species of the familiesLamiaceae andVerbenaceae. Data from 221 out of 262 genera ofLamiaceae s.l. and a few ofVerbenaceae s.str. were collected in a table. A cladistic analysis was performed on the basis of pericarp characters only. The abandonment of subfam.Pogostemonoideae as a taxonomic unit is consider...

Artificial crossing experiments were made with three taxa of Ocimum. Plants ofO. forskolei Benth. andO. basilicum L. var. purpurascens Benth. were found to be interfertile, butO. americanum L. var.pilosum (Willd.) Paton was found to be reproductively isolated from the other two taxa. The new chromosome number 2n = 48 was counted inO. forskolei. The...

A new species from southern and Eastern Ethiopia, Plantago annua is described and illustrated.

The pericarp structure has been investigated in 11 species representing the genera of the subfamily Pogostemonoideae (emend. Cantino et al. 1992): Anisomeles, Colebrookea, Comanthosphace, Leucosceptrum, Pogostemon and Rostrinucula. The pericarp structure of Anisomeles, Colebrookea and Pogostemon have been found to resemble that of the subfamilies L...

The two generaPlectranthus andIsodon are compared and found to be very dissimilar.Isodon ist considered to be misplaced inOcimeae subtribePlectranthinae and apparently is more closely related to subtribeHyptidinae. The disjunct genusRabdosiella is compared to these two genera and regarded to be polyphyletic. The AfricanR. calycina (Benth.)Codd is r...

Herbarium material from Ethiopia, Somalia and Kenya, previously included under Endostemon tereticaulis (Poir.) Ashby (Labiatae) have been found to belong to a separate species, earlier described as Orthosiphon kelleri Briq. The new combination, Endostemon kelleri (Briq.) O. Ryding is made. Synonymy, morphological description, habitat, comparisons w...

The phylogeny of the tribe Coreopsideae was investigated by cladistic analysis of generic interrelationships using a set of 37 morphological characters. The cladograms display three main groups: 1) a rather vaguely characterized assemblage of mainly American genera, viz. Coreopsis, Cosmos, Heterosperma, Henricksonia, Dahlia, Thelesperma, Coreocarpu...

A study of the variation within the tropical African taxon Trifolium burchellianum ssp. johnstonii, including var. oblongum, is made on the basis of herbarium material and cultivated material. The chromosome numbers 2n = 48 and 2n = 96 were counted in the collections of var. johnstonii and var. oblongum respectively. In most other material, the chr...

Thesis (doctoral)--Uppsala University, 1986. Includes bibliographical references (p. [146]-149) and index.

A revision is presented for the Aeollanthus abyssinicus group. Two new species, A. caudatus Ryding and A. namibiensis Ryding are described. A. lobatus N. E. Br., A. stuhlmannii Giirke, A. abyssinicus Hochst. ex Benth. and A. pinnatifidus Hochst. ex Benth. are also treated. Typifications, descriptions, notes on distribution and habitat, maps and key...