Norman E. Woodley

Smithsonian Institution · Department of Entomology

The genus Pseudocyphomyia Kertész (Diptera: Stratiomyidae: Pachygastrinae) is reviewed. Two new species are described, P. hansoni spec. nov. and P. thompsoni spec. nov., both known only from Costa Rica. A neotype is designated for P. mimetica Kertész. Zusammenfassung Es wird eine Übersicht zu den Arten der Gattung Pseudocyphomyia Kertész (Diptera:...

This revision is part of a continuing series of taxonomic work aimed at the description of new taxa and the redescription of known taxa of the Tachinidae of Area de Conservación Guanacaste in northwestern Costa Rica. Here we describe 33 new species in the genus Belvosia Robineau-Desvoidy, 1830 (Diptera: Tachinidae). All species described here were...

Antissella White, 1914 is a small genus of soldier flies endemic to Australia, previously thought to comprise only two described species and suspected to have a role in pollination. Six new species of Antissella are named, described and illustrated herein, and an updated identification key to all known species is provided. The new species include:...

Many eye‐catching and undescribed species of Australian soldier flies have been collected but left undiagnosed in entomological collections. We describe Opaluma Lessard & Woodley gen. nov., a new genus of iridescent soldier fly endemic to Australia from the subfamily Antissinae, and provide an updated key to the five Australian antissine genera. Se...

A taxonomic treatment to genera is provided for the Australian members of the soldier fly subfamily Sarginae. This includes an updated identification key for the Australian genera, along with the diagnosis and illustration of Ptecticus Loew, 1855 and Sargus Fabricius, 1798, and three newly recorded genera: Cephalochrysa Kertész, 1912 (Cephalochrysa...

We review the generic limits and classification of the little studied Australian fauna of the soldier fly subfamily Chiromyzinae. Among chiromyzines are many species that are confirmed agricultural pests of sugar cane, maize and grasses, including Inopus rubriceps (Macquart, 1847) that has been introduced in New Zealand and the United States. A tax...

A taxonomic treatment for the genera of the soldier fly subfamily Stratiomyinae is provided for the Australian fauna, including the first key to the Australian genera and updated key for the genus Acanthasargus White, 1914. Each genus is diagnosed and illustrated, including: Acanthasargus , Odontomyia Meigen, 1803, the newly recorded genus Prosopoc...

Study of all flies (Diptera) collected for one year from a four-hectare (150 x 266 meter) patch of cloud forest at 1,600 meters above sea level at Zurquí de Moravia, San José Province, Costa Rica (hereafter referred to as Zurquí), revealed an astounding 4,332 species. This amounts to more than half the number of named species of flies for all of Ce...

Estimations of tropical insect diversity generally suffer from lack of known groups or faunas against which extrapolations can be made, and have seriously underestimated the diversity of some taxa. Here we report the intensive inventory of a four-hectare tropical cloud forest in Costa Rica for one year, which yielded 4332 species of Diptera, provid...

A taxonomic treatment is provided for the Australian fauna of the soldier fly subfamily Hermetiinae, including a diagnosis of the subfamily based on the Australian fauna, a key to the two genera found in Australia, and a key to Australian species of Hermetia Latreille, 1804. The Australian fauna is illustrated and revised, including the redescripti...

Thirteen new generic synonyms, nineteen species synonyms and forty-eight new combinations of African Stratiomyidae are proposed (senior synonym in parentheses): Arthronemina Lindner in James, 1980 syn. nov. (=Argyrobrithes Grünberg, 1915), Arthronema Lindner, 1966b syn. nov. (=Argyrobrithes Grünberg, 1915), Brachyphleps Lindner, 1965 syn. nov. (=Ps...

Vermileonidae

A new species of Paraberismyia Woodley (Diptera, Stratiomyidae, Beridinae), P. imitator, n. sp., is described from Chiapas, Mexico. The new species is compared to previously described species and diagnostic notes are presented for separation of Paraberismyia from Berismyia Giglio-Tos.

A new species of Cyphomyia Wiedemann, Cyphomyia baoruco sp. n., is described from the Dominican Republic. A key to the species of Cyphomyia known from the Caribbean islands is provided.

Two new species of Brachyodina Lindner from the Caribbean are described, B. janestanleyae Wood ley, new species, from the Dominican Republic and B. caymanensis Wood ley, new species from Grand Cayman Island. A key to the species known from the Caribbean islands is provided.

The Neotropical genus Paraberismyia Woodley, 1995, is revised. Three new species, P. chiapas sp. n., P. mathisi sp. n., and P. triunfo sp. n. are described, all having type localities in Chiapas, Mexico. A key to the four known species is provided.

A new species of Lixophaga Townsend (Diptera: Tachinidae) from Colombia, Lixophaga puscolulo Carrejo & Woodley, sp. nov., is described and illustrated. It is a parasitoid of the tomato fruit borer, Neoleucinodes elegantalis (Guenée) (Lepidoptera: Crambidae), an insect pest of Solanum quitoense Lam., in Colombia. Aspects of its biology are briefly d...

The genus Parastratiosphecomyia Brunetti is revised with the description of two new species: Parastratiosphecomyia freidbergi sp. n. from India and Parastratiosphecomyia rozkosnyi sp. n. from Laos and Thailand. All four species in the genus are illustrated and a key to species is provided. Type localities of previously described taxa are briefly di...

Adult bot flies of the genus Cephenemyia Latreille are not commonly collected in spite of the abundance of their cervid host. The recent discovery of a live bot puparium (Figure 1) and emergence of the adult fly (Figures 2 and 3), identified as Cephenemyia phobifer (Clark) (Diptera: Oestridae) (sometimes incorrectly called C. phobifera), inspired t...

A new species of Culcua Walker (Diptera: Stratiomyidae) from Vietnam Insecta Mundi 0243: 1–4 Insecta Mundi is a journal primarily devoted to insect systematics, but articles can be published on any non-marine arthropod. Topics considered for publication include systematics, taxonomy, nomenclature, checklists, faunal works, and natural history. Inse...

A new genus and species of Stratiomyidae, Brianmyia stuckenbergi gen. et sp. n., are described from the Drakensberg, KwaZulu-Natal, South Africa. The new genus is placed in the tribe Prosopochrysini of the sub family Stratiomyinae and is the first genus and species of this tribe known from southern Africa.

Evidence is presented that the subspecies Chrysobothris thoracica guadeloupensis Descarpentries, 1981 (Coleoptera: Buprestidae) should be recognized at the species level. Character evidence is provided to separate C. guadeloupensis, new status, from C. thoracica Fabricius, 1798. Both species are illustrated with habitus photographs and images of th...

We present an overview of a 3 year project between the International Programs of the USDA Forest Service (FS) and the Agriculture Research Service (ARS) with some of the preliminary research results of the first 4 months. The ultimate research goals of this project are to define the species group to which Emerald Ash Borer (EAB), Agrilus planipenni...

Background/Question/Methods: A biodiversity inventory of the 120,000 terrestrial ha Area de Conservacion Guanacaste (ACG) in northwestern Costa Rica has been running for 33 years and has documented at least 9,000 species of Lepidoptera (and their parasitoids and food plants) of the estimated 15,000 species. In 2003 the inventory began routinely DNA...

A new genus and species of Stratiomyidae (Diptera), Vitilevumyia bobwoodleyi gen. et sp. nov. is described from the island of Viti Levu, Fiji. It exhibits an unusual combination of character states, but is tentatively placed in the tribe Proso-pochrysini of the subfamily Stratiomyinae.

New host records are given for Zelia tricolor (Coquillett), a tachinid fly that is a parasitoid of stem-boring beetles. New host records are Cerambycidae: Hippopsis lemniscata (F.) and Oberea tripunctata (Swederus); Curculionidae: Rhodobaenus quinquepunctatus (Say); and Languriidae: Languria bicolor (F.). The few previously reported hosts of Z. tri...

Oxycerina gen.n. (Stratiomyidae: Stratiomyinae), including three new species, O. hauseri sp.n., O. merzi sp.n. and O. sabaha sp.n., is described from the Oriental Region and compared with related genera of Stratiomyinae and Raphiocerinae. The monotypic genus Scapanocnema Enderlein, 1914 is considered to be a synonym of Odontomyia Meigen, 1803; S. s...

A new genus and species, Parameristomerinx copelandi gen. et sp. n. (Diptera: Stratiomyidae: Pachygastrinae) is described from material from Kenya. This genus is similar to the Afrotropical genera Meristomerinx Enderlein, 1914 and Meristomeringina James, 1952. A new generic synonym is also proposed, Nemodema Lindner, 1958 = Dolichodema Kertész, 191...

A new genus and species, Parameristomerinx copelandi gen. et sp. n. (Diptera: Stratiomyidae: Pachygastrinae) is described from material from Kenya. This genus is similar to the Afrotropical genera Meristomerinx Enderlein, 1914 and Meristomeringina James, 1952. A new generic synonym is also proposed, Nemodema Lindner, 1958 = Dolichodema Kertesz, 191...

Microchrysa flaviventris (Wiedemann) (Diptera: Stratiomyidae: Sarginae), a species widespread in the Old World, is reported from North America for the first time. Apparently it is established in the eastern United States. Specimens were taken in Alexandria, Virginia in August 2007.

Two new species of the cosmopolitan genus Metatrichia Coquillett are described. Metatrichia dhimurru sp. nov. is described from Arnhem Land (Northern Territory), Australia and represents the third species of the genus to be described from the Australasian region. Metatrichia venezuelensis sp. nov. from Venezuela is the third extant species of the g...

Two new species of the cosmopolitan genus Metatrichia Coquillett are described. Metatrichia dhimurru sp. nov. is described from Arnhem Land (Northern Territory), Australia and represents the third species of the genus to be described from the Australasian region. Metatrichia venezuelensis sp. nov. from Venezuela is the third extant species of the g...

Inventory of the caterpillars, their food plants and parasitoids began in 1978 for today's Area de Conservacion Guanacaste (ACG), in northwestern Costa Rica. This complex mosaic of 120 000 ha of conserved and regenerating dry, cloud and rain forest over 0-2000 m elevation contains at least 10 000 species of non-leaf-mining caterpillars used by more...

The genus Ditylometopa Kertész (Diptera: Stratiomyidae, Clitellariinae), which occurs only in the Neotropical Region, is reviewed. Euryneura rufifrons Curran, 1934 is synonymized with Ditylometopa elegans Kertész, 1923. A neotype is designated for D. elegans. Two species are recognized, including Ditylometopa centralensis sp. nov., which is describ...

The genus Ditylometopa Kertesz (Diptera: Stratiomyidae, Clitellariinae), which occurs only in the Neotropical Region, is reviewed. Euryneura rufifrons Curran, 1934 is synonymized with Ditylometopa elegans Kertesz, 1923. A neotype is designated for D. elegans. Two species are recognized, including Ditylometopa centralensis sp. nov., which is describ...

FIGURES 1 – 4. Habitus images of Ditylometopa species. 1, D. elegans, male. 2, D. centralensis, male. 3, D. elegans, female. 4, D. centralensis, female.

FIGURES 5 – 7. Morphological details of Ditylometopa species. 5, D. elegans, dorsal view of female abdomen. 6, D. centralensis, dorsal view of female abdomen; note: this image makes the abdominal hairs appear denser than they actually are. 7, D. elegans, posterior view of front femur of male.

FIGURES 8 – 11. Male terminalia of Ditylometopa elegans. 8, genital capsule, dorsal view. 9, phallic complex, dorsal view. 10, epandrium and post-genital segments, dorsal view. 11, phallic complex, left lateral view.

FIGURES 12 – 15. Male terminalia of Ditylometopa centralensis. 12, genital capsule, dorsal view. 13, phallic complex, dorsal view. 14, epandrium and post-genital segments, dorsal view. 15, phallic complex, left lateral view.

FIGURE 1. Metatrichia dhimurru sp. nov., male, Arnhem Land, Northern Territory (Body length = 3.5 mm) (Photo: S. L. Winterton).

FIGURE 4. Metatrichia venezuelensis sp. nov.: Holotype male, dorsal. Body length = 4.0 mm.

FIGURE 8. Metatrichia dhimurru sp. nov.: A, male genitalia, dorsal view; B, same, lateral view. Metatrichia venezuelensis sp. nov.: C. male genitalia, dorsal view; D, same, lateral view. Abbreviations: c, cercus; d, distiphallus; e, epandrium; ea, ejaculatory apodeme; ga, gonocoxal apodeme; gs, gonostylus; gx, gonocoxite; hy, hypandrium; va, ventra...

FIGURE 6. Metatrichia venezuelensis sp. nov.: Paratype female, dorsal. Body length = 4.5 mm.

FIGURE 2. Metatrichia dhimurru sp. nov.: Holotype male, dorsal. Body length = 3.5 mm.

FIGURE 3. Metatrichia dhimurru sp. nov.: Holotype male, lateral. Body length = 3.5 mm.

FIGURE 5. Metatrichia venezuelensis sp. nov.: Holotype male, lateral. Body length = 4.0 mm.

FIGURE 7. Metatrichia venezuelensis sp. nov.: Paratype female, lateral. Body length = 4.5 mm.

A new genus, Lobomyia Woodley & Arnaud (Diptera: Tachinidae), from the Neotropical Region was recently described by us (Woodley & Arnaud, 2008: 32; type species, Lobomyia neotropica Woodley & Arnaud; published 6 June 2008). At the time the manuscript was being prepared, one of us (NEW) checked available nomenclatural resources to ensure the generic...

A new genus and species, Lobomyia neotropica gen. nov., sp. nov., (Diptera: Tachinidae) is described from material from Colombia (type locality), Brazil, Costa Rica, Mexico, and Trinidad. The Colombian material was reared from Glena bisulca Rindge (Lepidoptera: Geometridae), a defoliator of introduced cypress trees. Costa Rican material was reared...

A new genus and species, Lobomyia neotropica gen. nov., sp. nov., (Diptera: Tachinidae) is described from material from Colombia (type locality), Brazil, Costa Rica, Mexico, and Trinidad. The Colombian material was reared from Glena bisulca Rindge (Lepidoptera: Geometridae), a defoliator of introduced cypress trees. Costa Rican material was reared...

A new genus and species, Kerteszmyia ecuadora gen. nov., sp. nov., (Diptera: Stratiomyidae, Pachygastrinae) is described from material from Ecuador (type locality), Venezuela, and Costa Rica. A key to the known Neotropical genera of Pachygastrinae with two or more scutellar spines is presented.

A new species of Hoplitimyia James, H. inbioensis sp. nov., is described from Costa Rica. A new genus, Panamamyia gen. nov. (type species P. silbergliedi sp. nov.) is described from Panama. Both taxa are very rare in collections, not having been taken during extensive Malaise trap surveys in Costa Rica.

A new subfamily of Stratiomyidae is proposed for Parhadrestia James and Cretaceogaster Teskey (fossil from Upper Cretaceous Canadian amber). Evidence is delimited that indicates that this subfamily is the sister-group to all other known stratiomyids. Taxa in the subfamily are systematically described, including a new species, Parhadrestia curico, f...

FIGURE 3. Lobomyia neotropica, gen. nov., sp. nov., dorsal view of female showing scutellar setation and abdominal color pattern.

FIGURES 4 – 6. Lobomyia neotropica, gen. nov., sp. nov., male terminalia. 3, Fifth sternite, ventral view. 4, Genital complex, posterior view. 5, Genital complex, lateral view. Abbreviations: c, cercus. dp, distiphallus. ej ap, ejaculatory apodeme. epi, epiphallus. gon, pregonite. s, surstylus.

FIGURES 1 – 2. Lobomyia neotropica, gen. nov., sp. nov. 1, Male, lateral view. 2, Female, lateral view.

FIGURE 7. Distribution map of Lobomyia neotropica, gen. nov., sp. nov. Filled circles indicate known localities (except more localities are known from Costa Rica but are not indicated due to size constraints).

Today’s knowledge of worldwide species diversity of 19 families of aquatic Diptera in Continental Waters is presented. Nevertheless, we have to face for certain in most groups a restricted knowledge about distribution, ecology and systematic, particularly in the tropical environments. At the same time we realize a dramatically decline or even lack...

Habitus photographs of holotypes and syntypes of Buprestidae (Coleoptera) in the Fernando de Zayas Collection, Havana, Cuba, are presented. Notes on the identities of the species are given. Peronaemis viridithorax Zayas 1988 is synonymized with Peronaemis monticola Fisher 1936.

Jurinella baoruco, n. sp., (Diptera: Tachinidae) is described and illustrated from the Sierra de Baoruco, Dominican Republic. A key to the genera of Tachinini from the Caribbean is presented. Two new generic synonyms are proposed: Hystriciella Townsend, 1915 and Parajurinia Townsend, 1928 both = Jurinella Brauer & Bergenstamm, 1889, resulting in Ju...

Derancistrus hovorei Lingafelter & Woodley, new species, is described from the Dominican Republic. A diagnosis is provided to discriminate it from morphologically similar taxa, in particular Derancistrodes vittatus (Olivier) and Derancistrus elegans (Palisot de Beauvois). Biological, distributional, and/or taxonomic notes are provided for other His...

A new species of Dasyomma Macquart, D. chrysopilum sp. nov., is described from Chile (Malleco and Ñuble Provinces). This species differs from all other known species in having dense golden pilosity on the abdominal tergites of both sexes. Dasyomma basale Malloch is redescribed, with the female described for the first time.

A new species of Dasyomma Macquart, D. chrysopilum sp. nov., is described from Chile (Malleco and Ñuble Provinces). This species differs from all other known species in having dense golden pilosity on the abdominal tergites of both sexes. Dasyomma basale Malloch is redescribed, with the female described for the first time.

Insect parasitoids are a major component of global biodiversity and affect the population dynamics of their hosts. However, identification of insect parasitoids is often difficult, and they are suspected to contain many cryptic species. Here, we ask whether the cytochrome c oxidase I DNA barcode could function as a tool for species identification a...

Populations of the cottonwood leaf beetle, Chrysomela scripta Fabricius (Coleoptera: Chrysomelidae), in Minnesota were examined for the presence of parasitoids over a period of two years. Field samples of larvae and pre-pupae yielded two parasitoids: a tachinid fly, Cleonice setosa (Reinhard), and a pteromalid wasp, Schizonotus sieboldi (Ratzeburg)...