Nicolas Goudemand

Nicolas Goudemand
Ecole normale supérieure de Lyon | ENS Lyon ·  UMR 5242 - Institut de Génomique Fonctionnelle de Lyon (IGFL)

Prof ENS Lyon, PhD tech. sci. ETH Zurich, PhD nat. sci. Univ. Zurich

About

98
Publications
28,523
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2,780
Citations
Additional affiliations
September 2017 - present
Ecole normale supérieure de Lyon
Position
  • Professor (Full)
October 2015 - present
Ecole normale supérieure de Lyon
Position
  • Group Leader
November 2014 - October 2015
University of Zurich
Position
  • Oberassistent

Publications

Publications (98)
Article
Full-text available
Shark populations that are distributed alongside a latitudinal gradient often display body size differences at sexual maturity and vicariance patterns related to their number of tooth files. Previous works have demonstrated that Scyliorhinus canicula populations differ between the northeastern Atlantic Ocean and the Mediterranean Sea based on biolo...
Article
Full-text available
The present dataset contains the 3D models analyzed in Berio, F., Bayle, Y., Baum, D., Goudemand, N., and Debiais-Thibaud, M. 2022. Hide and seek shark teeth in Random Forests: Machine learning applied to Scyliorhinus canicula. It contains the head surfaces of 56 North Atlantic and Mediterranean small-spotted catsharks Scyliorhinus canicula, from w...
Article
Taxonomy is the very first step of most biodiversity studies, but how confident can we be in the taxa delineation? One may hypothesize that the more abundant the material, the more accurate the description of morphological variability and hence the better the taxonomic delineation. Yet, as we shall see, in the case of numerous transitional forms, t...
Conference Paper
Full-text available
Ammonoids constitute an invaluable fossil record for documenting macroevolutionary patterns. Our study investigates the morphological evolution of ammonoids during their initial radiation through the Early and Middle Devonian. Previous morphological disparity analyses were restricted to conch geometry; here we have also explored the evolutionary pa...
Article
Full-text available
The mass extinction characterizing the Permian/Triassic boundary (PTB; ~ 252 Ma) corresponds to a major faunal shift between the Palaeozoic and the Modern evolutionary fauna. The temporal, spatial, environmental, and ecological dynamics of the associated biotic recovery remain highly debated, partly due to the scarce, or poorly-known, Early Triassi...
Article
Free download link for 50 days : https://authors.elsevier.com/a/1bsT13HcE1cwvI Conodonts, one of the longest-lived early groups of vertebrates, have a very complete fossil record ranging from the late Cambrian to the end of the Triassic and persisted through many global climatic and biotic events. In this paper, we analyse a large dataset harveste...
Conference Paper
Full-text available
The ammonoids appeared during the Lower Devonian (c. 405 Ma); they derived from the Bactritida which root in the Orthocerida in the latest Silurian or earliest Devonian. Earliest forms are found in Morocco (e.g. Praechebbites debaetsi) in early Emsian deposits. In the Anti-Atlas of Morocco, the Emsian ammonoids are abundant and well preserved. Thes...
Article
A key challenge regarding development of carbonate platforms is predicting the temporal pattern of platform-margin progradation, aggradation, retrogradation, and drowning. Numerical forward models of carbonate sedimentation have been widely applied to this problem, shedding substantial light on the roles of sediment production, transport, tectonic...
Article
Full-text available
Teeth in sharks are shed and replaced throughout their lifetime. Morphological dental changes through ontogeny have been identified in several species and have been correlated with shifts in diet and the acquisition of sexual maturity. However, these changes were rarely quantified in detail along multiple ontogenetic stages, which makes it difficul...
Article
Full-text available
The dynamics of the carbon cycle across different timescales is crucial for understanding past and present global climate changes. Following the Permian–Triassic boundary mass extinction (PTBME), the carbon cycle changed profoundly during the following 5.4 Myr, with magnitudes of changes comparable to those of the Precambrian. In pace with the succ...
Article
New Smithian (Early Triassic) ammonoid assemblages were sampled near the Utah/Arizona border. They provide several spatiotemporal constraints on the regional Sinbad Formation showing that the extent of the Smithian sea in the southwestern-most part of the western USA basin is larger than previously expected, reaching northern Arizona and an area ju...
Conference Paper
Full-text available
The Devonian ammonoids are particularly abundant and well preserved in the Anti-Atlas of Morocco. These rich faunas are well documented in the literature, and can provide an interesting record of the morphological disparity changes that characterize the macroevolution of the early Ammonoidea during the Devonian period. The present study aims to inv...
Article
Full-text available
Background: Many pathologies that modify the shell geometry and ornamentation of ammonoids are known from the fossil record. Since they may reflect the developmental response of the organism to a perturbation (usually a sublethal injury), their study is essential for exploring the developmental mechanisms of these extinct animals. Ammonoid patholo...
Article
In the aftermath of the Permian-Triassic boundary mass extinction (~252 Ma) ― the most dramatic biotic crisis of the Phanerozoic ― changes in climate, the carbon cycle, and biodiversity patterns remained extremely variable for several million years. In particular, the Smithian-Spathian boundary crisis, which occurred ca. 1.5 Ma after the Permian-Tr...
Article
Full-text available
Conodonts are among the first vertebrates to have evolved mineralized tooth-like structures. Among these, the so-called P 1 elements are known to have been used to break down food and display a wide variety of morphologies. In particular, the repeated independent evolution of platform-like P 1 elements have been suggested to correspond to similar f...
Article
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To assess evolutionary processes in deep time, it is essential to understand the roles of development and environment, both recorded through the morphological variability of fossil assemblages. Thanks to their great abundance and the high temporal resolution of their fossil record, conodont elements are ideal to address this issue. In this paper, w...
Article
A new, diverse and complex Early Triassic assemblage was recently discovered west of the town of Paris, Idaho (Bear Lake County), USA. This assemblage has been coined the Paris Biota. Dated earliest Spathian (Olenekian), the Paris Biota provides further evidence that the biotic recovery from the end-Permian mass extinction was well underway ca. 1.5...
Article
Conodont elements are composed of two main parts: the crown and the basal body. The basal body of a conodont element is a laminated structure that is less densely mineralized than the crown and, as a result, is rarely preserved. Elements with preserved basal bodies are found only in exceptional cases in the fossil record. To date, no S or M element...
Article
In contrast to the benthos whose taxonomic recovery was lagging way behind that of the nekton, highly diverse Smithian conodont and ammonoid faunas were profoundly decimated during the late Smithian, ca. 2 Myr after the Permian-Triassic boundary mass extinction. As body size reduction is a common evolutionary response to heavy environmental stress,...
Article
Full-text available
A new Early Triassic marine fauna is described from an exotic block (olistolith) from the Ad Daffah conglomerate in eastern Oman (Batain), which provides new insights into the ecology and diversity during the early aftermath of the Permian–Triassic Boundary mass extinction. Based on conodont quantitative biochronology, we assign a middle Griesbachi...
Article
Full-text available
We describe an Olenekian (Early Triassic) “fossil squid” belonging to the oldest complex Mesozoic marine biota collected in the Lower Shale unit of the Lower Triassic Thaynes Group in Idaho, USA. The studied specimen shows a tapered structure embedded in a cylindrical soft body. Morphological, ultrastructural and geochemical features of the specime...
Article
Recurrent microbialite proliferations during the Early Triassic are usually explained by ecological relaxation and abnormal oceanic conditions. Most Early Triassic microbialites are described as single or multiple lithological units without detailed ecological information about lateral and coeval fossiliferous deposits. Exposed rocks along Workman...
Article
The Guryul Ravine section (Kashmir, India) exposes one the world's most continuous carbonate rock successions throughout the Permian-Triassic boundary and beyond. Due to political instability in this region, the biostratigraphy of this section has not been updated for nearly three decades. Following new high-resolution sampling, we reassess here th...
Conference Paper
Full-text available
Following the Permian-Triassic mass extinction (PTME), the Early Triassic is characterized by large short-lived perturbations of the global carbon cycle associated with radiation and extinction pulses of the biota. More stable conditions resumed in the Middle Triassic (Anisian). The exact ages and duration of these short-lived but intense radiation...
Article
Full-text available
In the wake of the end-Permian mass extinction, the Early Triassic (~251.9 to 247 million years ago) is portrayed as an environmentally unstable interval characterized by several biotic crises and heavily depauperate marine benthic ecosystems. We describe a new fossil assemblage —the Paris Biota— from the earliest Spathian (middle Olenekian, ~250.6...
Data
Supplementary material (text, figures and tables) to: Brayard et al. 2017 - Science Advances 3, e1602159.
Article
1. Introduction In their Comment, Jiang et al. (2016) claim that the discordance between our zonation (Brosse et al., 2016) and the interval zones does not rest on the use of the Unitary Association method (Guex, 1991 and Guex et al., 2015) per se but on our “failure to use the most recent published conodont ranges from some key Chinese sections”....
Chapter
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The core of Evo-Devo lies in the intuition that the way tissues grow during embryonic development, the way they sustain their structure and function throughout lifetime, and the way they evolve are closely linked. Epithelial tissues are ubiquitous in metazoans, covering the gut and internal branched organs, as well as the skin and its derivatives (...
Article
The biochronological characterization of the Permian–Triassic boundary (PTB) is here improved by means of conodont unitary associations zones (UAZs). The selected data set comprises the six best documented sections in South China, including the Meishan global stratotype section and point of the PTB. This new biochronological zonation has a much hig...
Conference Paper
Full-text available
In the Batain region of Oman, basal Triassic crinoid limestones occur as exotic boulders within the Jurassic Guweiza Formation near Asselah. Some of these boulders contain the conodont association Hindeodus parvus-Isarcicella isarcica, indicating a Griesbachian age. Disarticulated crinoid columnal segments are the main component of these calcarenit...
Article
Full-text available
Changes of community structure in response to competition usually take place on timescales that are much too short to be visible in the geological record. Here we report the notable exception of a benthic marine community in the wake of the end-Permian mass extinction, which is associated with the microbial limestone facies of the earliest Triassic...
Article
Full-text available
We describe a new Early Triassic (Griesbachian) succession of conodont faunas from a high-resolution sampling of the basal Early Triassic microbial limestone and the base of the overlying unit at the Wuzhuan section (Nanpanjiang Basin, Guangxi, South China). The microbial limestone records the earliest phase of the Early Triassic biotic recovery af...
Conference Paper
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At the classical Guryul ravine section of Kashmir, the Permo-Triassic (P-T) boundary is located about 3 m above the base of the Khunamuh Formation. Brookfield et al. (2013) proposed that the deposits straddling the boundary between the Khunamuh Formation and the underlying, Permian Zewan Formation are Siberian Traps-induced seismites overlain by ts...
Conference Paper
Full-text available
Basal Triassic carbonate sediments are cropping out on the Gondwana margin in Oman, which are well dated (Hindeodus parvus to Isarcicella isarcica zones). These sediments reveal the contrasted oceanic conditions that prevailed in the immediate aftermath of the end-Permian "great dying". The Griesbachian shallow water carbonate platform consists of...
Conference Paper
Full-text available
A new sampling of Early Triassic (Griesbachian) conodonts is obtained from the microbialite overlying the latest Permian peri-reefal shallow water limestone in Wuzhuan section (Nanpanjiang Basin, Guangxi, South China). High resolution sam-pling in the lower twelve meters of the Luolou Formation provides rather diversified conodont faunas and allows...
Conference Paper
Full-text available
The importance of the Early-Middle Triassic boundary (EMTB) lays in the fact that this boundary marks the end of the time interval characterized by the biotic recovery after the end-Permian mass extinction. Therefore, to gain highest possible time accuracy and precision for quantification of biotic processes, we undertook a detailed calibration of...
Article
The end-Permian mass extinction is marked by pronounced terrestrial ecosystem turnover and a severe loss of marine invertebrate biodiversity. This extinction event is accompanied by a prominent negative carbon-isotope excursion indicating massive changes in the global carbon cycle across the Permian-Triassic boundary. In this study, we present orga...
Article
Full-text available
Sun et al. (Reports, 19 October 2012, p. 366) reconstructed Permian to Middle Triassic equatorial seawater temperatures. After correct temporal positioning of their data points, their presumed trends of temperature changes, and hence their assumption of a one-to-one relationship between putative "lethally hot" seawater temperatures and a disputable...
Article
Full-text available
Recovery from the end-Permian mass extinction is frequently described as delayed, with complex ecological communities typically not found in the fossil record until the Middle Triassic epoch. However, the taxonomic diversity of a number of marine groups, ranging from ammonoids to benthic foraminifera, peaked rapidly in the Early Triassic. These var...
Data
Recovery from the end-Permian mass extinction is frequently described as delayed, with complex ecological communities typically not found in the fossil record until the Middle Triassic epoch. However, the taxonomic diversity of a number of marine groups, ranging from ammonoids to benthic foraminifera, peaked rapidly in the Early Triassic. These var...
Data
Recovery from the end-Permian mass extinction is frequently described as delayed, with complex ecological communities typically not found in the fossil record until the Middle Triassic epoch. However, the taxonomic diversity of a number of marine groups, ranging from ammonoids to benthic foraminifera, peaked rapidly in the Early Triassic. These var...
Article
Several fused clusters of conodont elements of the genera Neospathodus and Novispathodus were recovered from limestone beds at the Dienerian–Smithian and Smithian–Spathian boundaries, respectively, from several localities in Guangxi province, South China. Conodont clusters are otherwise extremely rare in the Triassic, and these are first described...
Article
The thermally altered but otherwise well preserved Early Triassic palynomorphs from the Tulong area (South Tibet) allowed for qualitative and quantitative analyses and the differentiation of two distinct palynofloras, a Smithian lycopod-dominated flora and a Spathian gymnosperm-dominated flora. This suggests that gymnosperm vegetation recovered abo...
Article
The First Appearance Datum (FAD) of the conodont Chiosella timorensis has been recently proposed as an index for the worldwide recognition of the Olenekian-Anisian Boundary (OAB, Early-Middle Triassic boundary). We here report the co-occurrence of C. timorensis with the ammonoids Neopopanoceras haugi (Hyatt and Smith), Keyserlingites pacificus (Hya...
Article
Full-text available
Ammonoid-rich exotic bocks of Hallstatt facies in Jebel Safra, in Wadi Musjah and at Baid (Oman Mountains) have yielded several well preserved and highly diversified Smithian (Early Triassic) ammonoid faunas (Baidites hermanni fauna, Rohillites omanensis fauna, Flemingites rursiradiatus fauna, Nammalites pilatoides fauna, Owenites koeneni fauna and...
Article
Full-text available
Recovery from the devastating Permian-Triassic mass extinction about 252 million years ago is usually assumed to have spanned the entire 5 million years of the Early Triassic epoch. The post-crisis interval was characterized by large-scale fluctuations of the global carbon cycle and harsh marine conditions, including a combination of ocean acidific...
Article
Lower to Middle Triassic ostracods from the Tulong section, south Tibet, are described here for the first time. Samples from the first two stages of the Early Triassic (Griesbachian and Dienerian) are barren of ostracods; the following stage (Smithian) revealed low diversity ostracod faunas; a substantial diversification in taxa began at the base o...
Article
Full-text available
The origin of jaws remains largely an enigma that is best addressed by studying fossil and living jawless vertebrates. Conodonts were eel-shaped jawless animals, whose vertebrate affinity is still disputed. The geometrical analysis of exceptional three-dimensionally preserved clusters of oro-pharyngeal elements of the Early Triassic Novispathodus,...
Article
Full-text available
Monnet, C., Klug, C., Goudemand, N., De Baets, K. & Bucher, H. 2011: Quantitative biochronology of Devonian ammonoids from Morocco and proposals for a refined unitary association method. Lethaia, Vol. 44, pp. 469–489. Based on a rich dataset, the biostratigraphy of the late Emsian and the Eifelian (Early–Middle Devonian) ammonoids from the Moroccan...
Article
Full-text available
Based on new collections of abundant and well preserved material from the Salt Range (Pakistan), Spiti (Northern India) and Tulong (South Tibet), several recent studies focused on the taxonomic revision and detailed biostratigraphy of Smithian ammonoids. In this work, biochronological data for these three welldocumented basins are analyzed by means...