Nathaniel F. Shoobs

Nathaniel F. Shoobs
The Ohio State University | OSU · Museum of Biological Diversity - Division of Mollusks & Crustaceans

M.S. Environmental Science; B.A. Biology; Ecology
Collecting, curating, fighting entropy and imposing order on nature, etc.

About

17
Publications
7,184
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46
Citations
Introduction
I am a museum-based invertebrate zoologist and evolutionary biologist interested in the taxonomy, biogeography, and morphological evolution of land snails on oceanic islands. I use an integrative taxonomic approach grounded in collection-based revisionary systematics, comparative anatomy, and phylogenomics in order to understand the generation and maintenance of species-level diversity in the non-marine Mollusca through evolutionary time. Though my taxonomic expertise is in the Orthalicoid snails of the Neotropics (specifically the Galápagos Archipelago, northern South America, and the Caribbean Basin), I have broad interests in documenting, describing, and conserving terrestrial and freshwater invertebrate biodiversity across the globe.
Additional affiliations
March 2017 - March 2021
Academy of Natural Sciences of Drexel University
Position
  • Research Associate
August 2016 - December 2019
Drexel University
Position
  • PhD Student
June 2016 - July 2016
Delaware Museum of Natural History
Position
  • Mollusk Inventory Specialist
Education
September 2016 - December 2019
Drexel University
Field of study
  • Biodiversity, Earth, and Environmental Science
August 2012 - May 2016
Bard College at Simon's Rock
Field of study
  • Biology; Ecology

Publications

Publications (17)
Article
Full-text available
The presence of the freshwater polychaete, Namanereis hummelincki (Augener), on Montserrat is documented for the first time. Although collected in the sediment of a freshwater stream, this subterranean species most likely lives in groundwater aquifers. A mitochondrial cytochrome c oxidase subunit I (COI) sequence obtained from this material support...
Article
Full-text available
Four populations of the large freshwater ostracod, Chlamydotheca unispinosa (Baird, 1862), were discovered on the Caribbean island of Montserrat. These are the first records of the species on Montserrat and extend its known distribution approximately 113 km northwest and 63 km southeast of the closest known populations on Îles des Saintes (Guadelou...
Article
Full-text available
A new species of cave snail (Littorinimorpha: Cochliopidae) in the genus Antrorbis is described from the dark zone of two caves in the Appalachian Valley and Ridge province in eastern Tennessee, United States. The Tennessee Cavesnail, Antrorbis tennesseensis Perez, Shoobs, Gladstone, & Niemiller, sp. nov. is distinguished from its only known congen...
Article
Full-text available
Newly arrived species on young or remote islands are likely to encounter less predation and competition than source populations on continental landmasses. The associated ecological release might facilitate divergence and speciation as colonizing lineages fill previously unoccupied niche space. Characterizing the sequence and timing of colonization...
Article
Full-text available
Aim The accumulation of functional diversity in communities is poorly understood. Conveniently, the general dynamic model of island biogeography (GDM) makes predictions for how such diversity might accumulate over time. In this multiscale study of land snail communities on 10 oceanic archipelagos located in various regions of the globe, we test hyp...
Article
Full-text available
Freshwater crayfish are amongst the largest macroinvertebrates and play a keystone role in the ecosystems they occupy. Understanding the global distribution of these animals is often hindered due to a paucity of distributional data. Additionally, non-native crayfish introductions are becoming more frequent, which can cause severe environmental and...
Data
Records count per crayfish taxa integrated in WoC platform.
Data
Crayfish and A. astaci records integrated in WoC platform count per major hydrographic basins.
Data
Crayfish and A. astaci records integrated in WoC platform count per country.
Article
Full-text available
Four populations of the large freshwater ostracod, Chlamydotheca unispinosa (Baird, 1862), were discovered on the Caribbean island of Montserrat. These are the first records of the species on Montserrat and extend its known distribution approximately 113 km northwest and 63 km southeast of the closest known populations on Îles des Saintes (Guadelou...
Conference Paper
The presence of morphological anomalies in the radula and jaw of Naesiotus snails from Pinzón Island, Galapagos, is reported, and their potential evolutionary implications are discussed.
Thesis
In this thesis, I comprehensively assess the systematics, ecology, and distribution of the terrestrial and freshwater mollusks (Mollusca: Gastropoda) of Montserrat, a small volcanic island in the Leeward Islands of the Caribbean Lesser Antilles. This is the first systematic evaluation of Montserrat’s terrestrial and freshwater mollusk biodiversity,...
Conference Paper
Full-text available
A presentation regarding my preliminary work on the taxonomy, anatomy, ecology, and distribution of two rare and poorly known genera of Lesser Antillean semi-slugs.
Conference Paper
Full-text available
Montserrat is a small, volcanic island in the Caribbean Lesser Antilles, which is home to an unknown number of Neotropical gastropod taxa. We conducted the most extensive survey of the island to date, sampling over 30 sites across most of the inhabited portion of the island, and collecting specimens in order to better understand the biodiversity an...
Conference Paper
Full-text available
Gastropods in the Muricidae family secrete a white fluid from their hypobranchial glands, which becomes the Tyrian purple dye upon exposure to sunlight and air. This dye and its precursors may play a defensive role in both adult snails and egg masses. One muricid species, Plicopurpura patula (Linnaeus, 1758), is found along the Atlantic coast from...

Questions

Questions (3)
Question
Apologies for the confusing wording of the question.
To elaborate:
Smith describes a species, Aus bus Smith in 1875, and in his description, three things are clear:
1. there is only one specimen (that is, the specimen he describes is the holotype by monotypy),
2. Smith, not the most prodigious of scholars, only describes part of the animal, and his description is not sufficiently detailed to allow for identification between sister species, and
3. his good friend Jones, a talented anatomist, has  generously illustrated the anatomy of the new Aus bus Smith in a separate paper, from the same monotypic holotype specimen. This illustration is of a different part than Smith described (soft parts vs hard parts, for example). 
To make matters worse for recent authors,  exhaustive searching has yielded no parts of the holotype among the former collections of either taxonomist. Because it was a description of a monotypic specimen, there is no other contemporary material that can be referred to for reference, and contemporary workers of Jones and Smith expressed confusion with the name.
Recently collected topotypic material is clearly different from Jones' excellent illustrations, but Smith's poor description and lack of illustration makes it impossible to be sure of an ID. 
To repeat the principal question: is the fact that there was mutual acknowledgement that the illustrations of Jones were made from the holotype, can Jones' illustrations be used to solidify the concept of the name with comparisons to modern material in the absence of type material?
Do Articles 73.1.2  and 73.1.3 apply in this case? Furthermore, following Article 73.1.2, if it was possible for an author to designate an illustration is a holotype, could an illustration be lectotypified by a later worker? (Understanding that such illustrations are verifiably illustrations of the holotype).
Question
I am writing my undergraduate thesis as a taxonomic monograph. What is the best way to treat undescribed taxa, previously unpublished synonymies, or any other taxonomic acts in an undergraduate thesis? Is it best to use n.sp abbreviations and to wait until formal publication to fully describe new taxa?
The thesis will technically be available via a university digital commons, and a non-circulating but publically accessible physical copy will also remain in the library of my college. I also plan on scientifically publishing the monograph after I graduate. 
Any advice or technicalities to be aware of would be greatly appreciated. 
Question
Here's a question for the ICZN buffs out there:
If a species is described from a whole specimen that is split into separate museum lots, is the holotype both of the lots, or just one?
For example: I describe a novel species of snail from a specimen, and the description includes a description of the shell, the soft parts, and the radular anatomy. When depositing the specimen, I separate the shell and it gets a dry catalogue number. I separate the radula as well and it gets a separate catalogue number. The soft tissues get their own number and go into the wet collection. Are all of these lots still the holotype? Are they syntypes? They are all parts of the same organism but are obviously separate museum objects if they are prepared this way. 
Best, 
Nate

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