Milada Bocakova

Milada Bocakova
Palacký University Olomouc · Department of Biology

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84
Publications
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Publications

Publications (84)
Article
Cladocalon gen. nov. is proposed within the tribe Eurrhacini in the subfamily Lycinae including five species: C. chiriquense (Gorham, 1884) comb. nov. and C. histrionicum (Gorham, 1884) comb. nov. (transferred from Calocladon), C. bugabense sp. nov., C. mexicanus sp. nov. and C. guatemalense sp. nov. Lectotypes and paralectotypes of the two previou...
Article
Phylogenetic relationships of Vestini with extensive sampling were inferred using 37 taxa scored for 66 morphological characters. Consistent with recent DNA analyses, Vesta is found to be sister to Photurinae rendering Vestini monotypic. Conversely, our parsimony and model-based analyses robustly support the new subfamily Cladodinae established for...
Article
A new genus of Neotropical Lycidae, Currhaeus gen. nov., is herein proposed as the second Eurrhacini genus lacking parameres in male genitalia. Seven new species are described: Currhaeus striatus sp. nov., C. nigroapicalis sp. nov., C. championi sp. nov., C. tabascensis sp. nov., C. ruschii s . nov., C. polegattoi sp. nov., and C. paranaensis sp. n...
Article
A new subgenus of Tenebroides, Polynesibroides subgen. nov., is established for six potentially extinct beetles, Tenebroides atiu, T. moorea, T. raivavae, T. rimatara, T. tubuai and T. mihiura spp. nov., described from the Cook, Austral and Society Islands based on subfossil fragments, and the extant T. tahiti sp. nov. from the island of Tahiti, So...
Article
A new subgenus of Tenebroides, Polynesibroides subgen. nov., is established for six potentially extinct beetles, Tenebroides atiu, T. moorea, T. raivavae, T. rimatara, T. tubuai and T. mihiura spp. nov., described from the Cook, Austral and Society Islands based on subfossil fragments, and the extant T. tahiti sp. nov. from the island of Tahiti, So...
Article
The South American genus Lycoplateros Pic, 1922 is revised. This genus is a member of the Eurrhacini clade, placed as a sister-group of Neolinoptes Nascimento & Bocakova, 2017. Lycoplateros can be differentiated from other Eurrhacini based on its characteristic pronotum and complex male genitalia, characterized by phallus presenting internal sac wi...
Article
Full-text available
A new species of Lycidae, Falsocaenia santatereza sp. nov. is described and illustrated. This species is a member of a Müllerian mimicry ring from the Atlantic Forest in Brazil, presenting color pattern also found in other sites of Neotropical region. There are now 15 species from Falsocaenia Pic, 1922, all restricted to Neotropical region. Additio...
Article
The first thorough molecular phylogeny of the superfamily Cleroidea, represented by 377 taxa, and the first with an emphasis on Trogossitidae, was undertaken. Maximum likelihood and Bayesian analyses were performed on a four‐gene dataset (18S, 28S, cox1, cytb) of 395 taxa (along with 18 outgroups), including all 16 currently recognized families of...
Article
Full-text available
Background Rhinorhipidae Lawrence, 1988 is an enigmatic beetle family represented by a single species, Rhinorhipus tamborinensis Lawrence, 1988, from Australia, with poorly established affinities near the superfamily Elateroidea (click beetles, soldier beetles and fireflies) or the more inclusive series (infraorder) Elateriformia. Its evolutionary...
Preprint
Full-text available
Background: Rhinorhipidae Lawrence, 1988 is an enigmatic beetle family represented by a single species, Rhinorhipus tamborinensis Lawrence, 1988, from Australia, with poorly established affinities near the superfamily Elateroidea (click beetles, soldier beetles and fireflies) or the more inclusive series (infraorder) Elateriformia. Its evolutionary...
Article
Full-text available
The relationships of the monogeneric family Plastoceridae Crowson, 1972 (Coleoptera: Elateroidea) have remained contentious due to its modified morphology, incorrect information on incomplete metamorphosis of females and the absence of molecular data. We produced the sequences for P. angulosus (Germar, 1844) (the type-species of Plastocerus Schaum,...
Article
Here we present a revision of the Neotropical genus Eurrhacus Waterhouse, 1879 that is placed in Eurrhacini tribe, strongly related to Calopterini, being the only genus from these tribes or even among exclusive Neotropical lycids known so far to present reduced or absent parameres. Six species are placed in the genus, two of them, E. tristis (Water...
Article
The genus-level molecular phylogeny of Omalisidae Lacordaire is presented for six of seven currently recognised genera. The monophyly and internal relationships are well-supported including the taxa which were placed in other elateroid families. We conducted molecular analyses using maximum-likelihood optimality criterion and Bayesian inference and...
Article
Neolinoptes gen. n. is erected to replace Linoptes Gorham, 1884, preoccupied by Linoptes Menge, 1854 (Arachnida: Araneae). Consequently, Neolinoptes imbrex (Gorham, 1884) comb. n., N. amazonicus (Pic, 1923) comb. n., N. atronotatus (Pic, 1922) comb. n., N. atripennis (Pic, 1932) comb. n. are proposed. Calocladon rubidum Gorham, 1884 is transferred...
Article
Full-text available
Neolyrium iquitosense sp. nov., a new species of Lycidae, is described and illustrated. This second species of the genus represents the fi rst record of Neolyrium from Peru, keeping its distribution restricted to tropical areas of South America.
Article
Full-text available
Neolyrium iquitosense sp. nov., a new species of Lycidae, is described and illustrated. This second species of the genus represents the fi rst record of Neolyrium from Peru, keeping its distribution restricted to tropical areas of South America.
Article
A new genus and four new species of Indian endemic beetles belonging to the subfamily Ototretinae (Lampyridae) are described: Flabellopalpodes gen. nov., F. flavus sp. Nov., F. brancuccii sp. Nov., F. atricornis sp. Nov., and F. tamilnaduensis sp. Nov. One species formerly placed in Drilidae, Selasia maindroni Bourgeois, 1903, is transferred to the...
Article
Full-text available
The Amydetinae genus Scissicauda McDermott, 1964 is reviewed and redescribed. We describe S. balena sp. n. from Brazil as new, and provide illustrations of the structural features and a key to species of both sexes.
Article
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Cladodes illigeri (Kirby, 1818) is redescribed, and can be distinguished by the following features: color pattern overall black, paired spots and elytral margins pale yellow; pygidium bisinuate, posterior angles rounded, as long as median margin; and aedeagus with phallus 1/3 shorter than the parameres, which are sinuated apically. Cladodes lamelli...
Article
Full-text available
The evolution of animal life strategies is among the main themes of current evolutionary biology. Checkered beetles, soft-winged flower beetles and their allies (superfamily Cleroidea), exhibit well-known aposematic colour patterns, particularly in the family Cleridae, which participate in mimicry complexes mostly with unpalatable beetles, ants and...
Article
A new extant species of Electribius Crowson, E. llamae sp.n., is described from Cusuco National Park, Cortés Province, Honduras. This new species lacks one of the supposed autapomorphies of the genus; therefore the definition of the genus requires modification. A revised key to the known extant species is presented, and their known distributions ar...
Article
Full-text available
The Ototretinae genus Lamellipalpodes Maulik, 1921 is reviewed. Four species are proposed as new to science, L. bajhangensis Bocakova sp. nov., L. holzschuhi Bocakova sp. nov. and L. godawarensis Bocakova sp. nov., all from Nepal, and L. yunnanensis Bocakova sp. nov. from China (Yunnan) and northern Laos. Six species are redescribed. Illustrations...
Article
Full-text available
A new genus Lolodorfus is proposed within the subfamily Dexorinae and Lolodorfus flavus sp. nov. from Cameroon is described. Illustrations of diagnostic characters are given. The genus Mimolibnetis Pic, 1936 is transferred to the Dexorinae Bocak et Bocakova, 1989, and consequently the subfamily Mimolibnetinae Kazantsev, 2013 is considered to be a y...
Article
Full-text available
The Artematopodidae is a species-poor beetle family with con- tentious relationships to byrrhoid and elateroid families. Recent molecular phylogenetic analyses brought ambiguous results based on a single sequenced species. We investigated the taxonomic placement of Artematopodidae within Elateriformia using ribo- somal (18S, 28S) and mitochondrial...
Article
Brasilycus peruanus sp. nov. is proposed as new to science. Photographs of the habitus, antenna, pronotum, and aedeagus of the single specimen are presented. This new species represents the first record of the genus Brasilycus from Peru and is the fourth species in the genus recorded from South America.
Article
The Ototretinae are revised using comparative morphology of adults. All recognized genus-group taxa are redescribed and illustrated and a key to genera is provided. The concept of Ototretinae within Lampyridae is clarified and the taxonomic placement with the phylogenetic relationships of several taxa formerly associated as cantharoids is discussed...
Article
Several nomenclatural acts are proposed in this paper. Plateros plaumanni n. nov. is proposed for Sculptocalleros discithorax Pic, 1949 as a replacement name. Following 3 varieties are elevated to species rank and transferred to the genus Plateros from Calleros: Plateros curtebilineatus (Pic, 1934) comb. et stat. nov. for Calleros brasiliensis var....
Article
Full-text available
The Neotropical genus Falsocaenia Pic, 1922 is revised. Two species Falsocaenia ecuadorensis sp. nov. and F. melanopteron sp. nov. are proposed as new to science. Idiopteron (F.) apicicorne (Pic, 1922) and Idiopteron (F.) aurantiacocostatum (Pic, 1932) are placed in the genus Falsocaenia, and consequently, Falsocaenia apicicornis (Pic, 1922) comb....
Article
Full-text available
A new species Picomicrolycus ireneae sp. nov. from Brazil is described and Calleros pilosus Pic, 1934 is transferred to Picomicrolycus, recording this genus for the first time from South America. Diagnostic characters are illustrated and a key to known Picomicrolycus species is given.
Article
The melyrid lineage of beetles form a distinct group of the superfamily Cleroidea with a high level of soft-bodiedness. Here we present the first molecular phylogenetic analysis of this group. The data matrix included partial sequences of the small and large subunits of rRNA, the mitochondrial large subunit rRNA, and cytochrome oxidase subunit I of...
Article
The genus Hyperstoma Wittmer, 1979 is reviewed and Hyperstoma wittmeri sp. nov. from Sri Lanka is proposed as a new species. A key to Hyperstoma and illustrations of diagnostic characters are given.
Article
The Neotropical genus Ceratopriomorphus Pic, 1922 is reviewed. Three species are currently placed in the genus, one of which, C. bolivianus sp. nov., is described here. All diagnostic characters are illustrated, and a key to known Ceratopriomorphus species is given.
Article
The genus group taxa of the tribe Calopterini are revised. The genus Callanganum Pic, 1922 is proposed as a junior synonym of Ceratopriomorphus Pic, 1922, the genus Pseudolinoptes Pic, 1922 as a junior synonym of Emplectus Erichson, 1847. The genera Broxylus Waterhouse, 1879 and Ceratoprion Gorham, 1884 are transferred to the subfamilies Metriorrhy...
Article
Full-text available
The new genus Brasilycus is proposed within the tribe Calopterini. Following new species are described in the genus: Brasilycus capixabus n. sp., B. rondonensis n. sp., and B. bolivianus n. sp., all from South America. Illustrations of diagnostic characters and key to Brasilycus are given.
Article
We review the Neotropical genus Cartagonum Pic, redescribing C. bernardi Pic and describing two new species, Cartagonum apiuba sp. nov. and Cartagonum mayense sp. nov. New collection records, a key to species, and illustrations of somatic and genitalic characters are provided. Species of Cartagonum are now known to be distributed from Colombia to s...
Article
Emasia n. gen. is proposed within the Ototretinae together with one new species Emasia dentata n. sp. from Malaysia (Borneo). Principal apomorphic features are listed and illustrations of diagnostic characters provided.
Article
The Neotropical genus Lycomorphon Pic, 1922 is revised. Eleven species are classified in the genus, six of them are redescribed, five species (Lycomorphon brasiliense sp. n., L. amazonicum sp. n., L. bimaculatum sp. n. L. bolivianum sp. n., L. fulvohumeralis sp. n.) are proposed as new to science. Lycomorphon elongaticolle v. diversicolle Pic, 1926...
Article
The Neotropical genus Lycomorphon Pic, 1922 is revised. Eleven species are classified in the genus, six of them are redescribed, five species (Lycomorphon brasiliense sp. n., L. amazonicum sp. n., L. bimaculatum sp. n. L. bolivianum sp. n., L. fulvohumeralis sp. n.) are proposed as new to science. Lycomorphon elongaticolle v. diversicolle Pic, 1926...
Data
FIGURES 38 – 52. Male genitalia: (38 – 40) Lycomorphon fulvohumeralis; (41 – 43) L. amazonicum; (44 – 46) L. (Spinolycus) brasiliense; (47 – 49) L. (Spinolycus) bolivianum; (50 – 52) L. (Spinolycus) irregularis. (38, 41, 44, 47, 50) ventral view; (39, 42, 45, 48, 51) dorsal view; (40, 43, 46, 49, 52) lateral view. Scales 0.5 mm.
Data
FIGURES 1 – 9. (1 – 3) Pronotum in dorsal view: (1) Lycomorphon diversicolle; (2) L. opacum; (3) L. brasiliense. (4 – 6) Abdominal sternum 8: (4) L. bimaculatum; (5) L. bolivianum; (6) L. irregularis. (7 – 8) Terminal abdominal segments in ventral view: (7) L. nitidum; (8) L. opacum. (9) elytron of L. opacum. Scales 0.5 mm.
Data
FIGURES 10 – 19. Basal antennomeres: (10) Lycomorphon elongaticolle; (11) L. opacum; (12) L. nitidum; (13) L. diversicolle; (14) L. bimaculatum; (15) L. fulvohumaralis; (16) L. amazonicum; (17) L. brasiliense; (18) L. bolivianum; (19) L. irregularis. Scales 0.5 mm.
Data
FIGURES 20 – 37. Male genitalia: (20 – 22) Lycomorphon elongaticolle; (23 – 25) L. angusticolle; (26 – 28) L. opacum; (29 – 31) L. nitidum; (32 – 34) L. diversicolle; (35 – 37) L. bimaculatum. (20, 23, 26, 29, 32, 35) ventral view; (21, 24, 27, 30, 33, 36) dorsal view; (22, 25, 28, 31, 34, 37) lateral view. Scales 0.5 mm.
Article
Full-text available
Lycidae, net winged beetles, have proved difficult to classify using morphological characters. Here, using a previously published molecular phylogeny, comparing the results with morphological data and re-analyzing previously published morphological data set, we propose a revised classification of Lycidae. All analyses support the monophyly of Lycid...
Article
Full-text available
Neoteny, the maintenance of larval features in sexually mature adults, is a radical way of generating evolutionary novelty through shifts in relative timing of developmental programmes. While controlled by the environment in facultative neotenics, retention of larval features is obligatory in many species of Lycidae (net-winged beetles). They are s...
Article
Genus-group taxa of Libnetini are revised and valid taxa are redescribed. Microlyropaeus Pic, 1929 formerly classified with Libnetini (Bocakova 2001a) is transferred to the subfamily Leptolycinae. New genus Prolibnetis gen. n. is proposed within Libnetini, and two new Libnetisia species from China are described: Libnetisia yunnanesis sp. n. and L....
Article
Review of Ceylonese species of the tribe Cladophorini and key to Cautires species are given. New species Cautires ceylonicus sp. n. and C. krombeini sp. n. are described. Cautires dolosus Kln. is proposed to be a synonym to C. astutus F. Walk. The new genus Tapromenoeus gen. n. is described, it contains the only species Tapromenoeus hirtus (Kln.) —...
Article
Full-text available
Beetles represent almost one-fourth of all described species, and knowledge about their relationships and evolution adds to our understanding of biodiversity. We performed a comprehensive phylogenetic analysis of Coleoptera inferred from three genes and nearly 1900 species, representing more than 80% of the world's recognized beetle families. We de...
Article
Phylogenetic relationships in the coleopteran Series Elateriformia (click beetles, jewel beetles, fireflies and allies) were investigated using > 3800 nucleotides of partial nuclear (small and large subunit rRNA genes) and mitochondrial (large subunit rRNA and cytochrome oxidase subunit I) gene sequences. The Elateriformia includes several soft-bod...
Article
really a basal neopteran lineage – a reply to Kazantsev. — Acta Zoologica (Stockholm) 88 : 153–158 In a study on the mandible and mandibular articulation of larvae of the lycid genus Platerodrilus Kazantsev proposes a phylogenetic scheme with Polyphaga as a basal group of Neoptera and Lycidae as a basal group within Polyphaga. Here we point out dif...
Article
Full-text available
A review of genera within the tribe Lyropaeini is given. Three new genera - Lyrolib gen. n., Horakiella gen. n., and Ambangia gen. n., and a subgenus Macroambangia subgen. n. within Ambangia gen. n. are described. The following twelve new species are proposed as new to science: Ambangia nigra sp. n. (Sulawesi), A. wallacei sp. n. (Sulawesi), Ambang...
Article
The phylogeny of lycid tribe Calopterini is examined in the light of adult morphological evidence. Representatives from each of the Calopterini genus - group taxa and genera Dictyoptera, Lygistopterus, Conderis, Plateros, and Lycus are examined. An investigation of Calopterini phylogeny is presented here, based on 33 characters and 24 supraspecific...
Article
The genus Dihammatus Waterhouse, 1879 is recorded from China for the first time. Descriptions of two Chinese Dihammatus are presented with new Dihammatus from Nepal and Laos, and a new Libnetis Waterhouse, 1878 from central China. The following new species are described: Dihammatus chinensis, D. xunyanbanensis, D. holzschuhi, D. kubani, and Libneti...
Article
Full-text available
Genus-group taxa of Platerodinae are revised and valid taxa are redescribed. The validity of Plateros Bourgeois, 1879 is reinstated. Libnetomimus Kleine, 1927 is made a junior synonym of Libnetis Waterhouse, 1878. Calleros Gorham, 1881, Calloplateros Pic, 1923, Costatoplateros Pic, 1949, Ditoneces Waterhouse, 1879, Libnetomorphus Pic, 1921, Micropl...
Article
Libnetis species from Laos are treated. Eight new species are described: Libnetis laosensis sp. n., L. merkli sp. n., L. viti sp. n., L. jendeki sp. n., L. holzschuhi sp. n., L. dentifer sp. n., L. montanus sp. n., and L. sausai sp. n.. Species L. opacus PIC, 1921, L. thai KASANTSEV, 1993, and L. vietnamicus (KASANTSEV, 1993) are recorded from Laos...
Article
Following eight species are described: Melaneros inermis sp. n., M. oculeus sp. n., M. kejvali sp. n., M. cardamomensis sp. n., M. pacholatkoi sp. n., M. uzeli sp. n., M. bicoloratus sp. n. and M. parvus sp. n. Male copulatory organs of all species and other important diagnostic features are figured.
Article
Full-text available
A new genus Chinotaphes gen. n. with new species Chinotaphes weibaoshanensis sp. n. from the tribe Erotini is proposed. Following species are classified in the genus Lobatang Bocák, 1998: Lobatang flavohu-meralis sp. n. is described from Japen island, Indonesia and Lobatang dubitabilis (Kleine, 1925), comb. n. (Philippines) is transferred from the...
Article
1. vyd. Nad názvem: Univerzita Palackého v Olomouci, Pedagogická fakulta

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Projects (2)
Project
This project aims to establish the knowledge about taxonomy, spacial distribution and ecology of Neotropical Lycidae through taxonomic reviews, descriptions of new taxa and study of mimetic relations involving lycid beetles.
Project
I initiated research on beetle diversity and communities in Peru in 2007. The project has focused on inventories along an elevational transect along the Madre de Dios Watershed (Kosñipata Valley to Puerto Maldonado). Annual collections have yielded 1000s of specimens, labeled and digitized at the University of Kansas. These collections are the basis of phase 1 that developed species lists with validated names, relevant literature, and a network of collaborating family experts. Phase 2 will describe many new species my teams have collected, continue beetle surveys around Peru, and continue study of beetle communities, environment, and elevation along the Madre de Dios transect.