Maxim Nabozhenko

• Doctor of Sciences
• Researcher at Daghestan Federal Research Centre of the Russian Academy of Sciences

Comb-clawed beetles of the genus Allecula Fabricius, 1801 (Coleoptera: Tenebrionidae) of Russia are briefly reviewed. A new species, A. sundukovi sp. n., is described from the Russian Far East (Khabarovsky Krai). This new species differs from all congeners by the shape of the pronotum, very similar to those in tenebrionid of the tribe Lagriini, ver...

A review of darkling beetles of the genus Microdera Eschscholtz, 1831 from Uzbekistan is presented. In total, 12 species are known in this country. Two new species are described: M. abdullohi N. Bekchanov et Nabozhenko, sp. nov. (Sulton Uvays Mts.) and M. uzbekistanica N. Bekchanovet Nabozhenko,sp. nov. (Kuldzhuktau Mts.). The following new synonym...

Five new tenebrionid species of the genus Calyptopsis Solier, 1835 are described from Iran: C. bakhtiarii n. sp., C. muilwijki n. sp., C. granulimentum n. sp., C. seyedabadii n. sp. and C. mokhtariani n. sp. A new synonymy is proposed: Calyptopsis sulcimargo Reitter, 1897, n. syn. of Calyptopsis harpaloides Baudi di Selve, 1874. The following speci...

The darkling beetles of the genus Penthicus Faldermann, 1836 (Tenebrionidae: Blaptinae: Opatrini) from Iraq, Iran and Pakistan are revised. We used male and female genitalia for the first time for taxonomy of Penthicus. The subgenus Stonavus Reitter, 1904, subg. resurr. is resurrected from the synonymy with Penthicus and incudes three species-group...

Sixteen species of darkling beetles (Coleoptera: Tenebrionidae) in mountainous Ingushetia (Russia, the North Caucasus) were collected during two years in different seasons and habitats. The new species Mycetochara (Ernocharis) ingushetica sp. n. (close to M. abschasica Pic, 1925) is described from an oak grove in the Assa River valley. The followin...

The composition and distribution of darkling beetles of the genus Uloma are briefly reviewed for the Russian fauna. In total, four species are known from Russia, one of which is distributed in the European part and the Russian Caucasus, one is transpalaearctic, two sympatric species are known only in Primorsky krai of the Russian Far East. The spec...

The invertebrate fauna of the Hissar State Reserve in Uzbekistan remains largely unstudied. In this reserve, we collected and examined darkling beetles, which are among the most diverse and abundant groups of beetles in Middle Asia. All species were found at altitudes ranging from 1,800 to 2,500 meters, spanning the zone of Juniperus seravschanica...

The lectotype is designated for Anisocerus funebris Reitter, 1898 (now in the genus Ceratanisus Gemminger, 1870). This species is distributed in central and eastern Turkey, and specimens from Artvin Province (northeastern Turkey) in previous paper of the first author were erroneously identified as С. funebris. The new species Ceratanisus arankae Na...

The fundamental value of universal nomenclatural systems in biology is that they enable unambiguous scientific communication. However, the stability of these systems is threatened by recent discussions asking for a fairer nomenclature, raising the possibility of bulk revision processes for "inappropriate" names. It is evident that such proposals co...

List of the 1543 additional coauthors. Co-authors who contributed revising translations are listed first. Then, the rest of the coauthors are listed according alphabetic order of countries/territories.

The Central Asian genus Dengitha Reitter, 1887 is briefly reviewed. The following new synonymy is proposed: Dengitha crystallina Semenow, 1896 = Dengitha symmetrica G.S. Medvedev, 1964 syn. nov.; Dengitha lutea Reitter, 1887 = Dengitha hyalina Semenow, 1896 resyn. Lectotype is designated for D. crystallina. In total, two species of the genus are di...

The genus Sternodes Fischer von Waldheim, 1837 currently consists of one species with two subspecies from Karakum, Sundukli deserts and several surrounding areas. Sternodes pallasi Semenov-Tian-Shanskij et Bogatchev, 1940 is resurrected from a junior synonym of Sternodes caspicus (Pallas, 1781). The following new synonymy is established: Sternodes...

The third fossil species of the tribe Bacaniini (Coleoptera, Histeridae), Bacanius caterinoi Sokolov et Perkovsky, sp. nov., is described from the Eocene Rovno amber. Previous two extinct species were known from the Rovno and Baltic amber. We compared the new species with them and extant representatives of related taxa. The new species belongs to t...

New data on distribution of rare and little-known species of comb-clawed beetles (Tenebrionidae: Alleculinae) in the European part of Russia and the Caucasus are presented. Allecula morio (Fabricius, 1787) is recorded for Russia for the first time; Crimean Peninsula is the easternmost border of its range. Information about three very rare species,...

The subgenus Turcmenicola Bogatchev, 1952 (genus Colposcelis Dejean, 1834) is reviewed. Representatives of the studied taxon occur in the most lifeless rocky, gravelly and clayey areas of the deserts of Middle Asia (Ustyurt plateau, Kyzylkum desert, foothills of southern Kugitang, Betpak-Dala desert). Turcmenicola comprises five species from which...

The subgenus Turcmenicola Bogatchev, 1952 (genus Colposcelis Dejean, 1834) is reviewed. Representatives of the studied taxon occur in the most lifeless rocky, gravelly and clayey areas of the deserts of Middle Asia (Ustyurt plateau, Kyzylkum desert, foothills of southern Kugitang, Betpak-Dala desert). Turcmenicola comprises five species from which...

Several fossil beetles from the Late Pliocene (Piacenzian) of Willershausen (Lower Saxony, Germany) previously identified as Tenebrionidae and Staphylinidae are reviewed. One species, formerly determined as "Staphylinoidea" is identified as Euboeus mimonti Boieldieu, 1865 (Tenebrionidae: Helopini). This species is the oldest record of an extant spe...

A new species of darkling beetle of the tribe Helopini is described from the Eastern Caucasus (Russia): Nalassus magomedrasuli Nabozhenko sp. n. It was originally described as the Dagestan population of N. kalashiani Nabozhenko, 2001 based on the similar structure of the male aedeagus. The new species is similar to N. kalashiani in that it differs...

Although the complex evolutionary history of lichen-forming fungi has gained considerable attention, particularly regarding the long-debated role of these organisms in shaping early terrestrial ecosystems, the evolution of lichenivory and its potential impact on the diversification of lichenophages have been largely neglected. With > 800 described...

Пересмотрено таксономическое положение некоторых видов и подвидов рода Pedinus Latreille, 1796. Два подвида вида Pedinus cimmerius G.S. Medvedev, 1968, обитающие на нижнеднепровских песках и песчаных морских побережьях Кавказа, отнесены к виду Pedinus femoralis (Linnaeus, 1767): P. femoralis caucasicus G.S. Medvedev, 1968 и P. femoralis znoikoi G.S...

Several fossil beetles from the Late Pliocene (Piacenzian) of Willershausen (Lower Saxony, Germany) previously identified as Tenebrionidae and Staphylinidae are reviewed. One species, formerly determined as “Staphylinoidea” is identified as Euboeus mimonti Boieldieu, 1865 (Tenebrionidae: Helopini). This species is the oldest record of an extant spe...

A new species of darkling beetles of the genus Philhammus Fairmaire, 1871 is described from Amudarya bank, Khorezm Region, Uzbekistan: Ph. dilaramae Nabozhenko et N. Bekchanov, sp. n. This first representative of the genus in the country belongs to the nominotypical subgenus and most similar to Pre-Caspian Ph. zaitsevi G.S. Medvedev, 1979. The new...

Results on study of darkling beetles after expeditions 2021–2023 in western Uzbekistan (Khorezm Region, the Republic of Karakalpakstan) are summarized. In total, 79 species of tenebrionid beetles were collected, from which four taxa are new for science: Diaphanidus mamuni Nabozhenko et N. Bekchanov, sp. n., Diaphanidus crassiantennatus Nabozhenko e...

A new extinct monotypic genus of the ironclad beetles, Zopheromimus auriborussiensis gen. et sp. nov., is described and illustrated based on an inclusion in Baltic amber. The new Eocene fossil taxon is placed in Zopheromimini trib. nov., a new tribe within the subfamily Zopherinae, and compared with related extant beetles of the subfamily. The genu...

Two darkling beetles from the subfamily Lagriinae from the late Pliocene of Japan, Kabutoiwa Member (early Piacenzian, ~3.5 Ma) of the Motojuku Formation are discussed: Lagria koshimizui sp. nov. and Lagria aff. formosensis Borchmann, 1912. These are the first members of the family Tenebrionidae recorded from Pliocene and the first clearly document...

New data on the distribution, taxonomy, and nomenclature of fi ve tenebrionid species of the genus Microdera from the Caucasus and/or the south of the European part of Russia are presented. A new species, M. urartu sp. n., is described from Armenia and the Nakhchivan Autonomous Republic of Azerbaijan; this species is similar to the Central Asian M....

A new genus and species are described from the Eocene Baltic amber: Archaeoluprops groehni gen. et sp. nov. This fossil darkling beetle of the subfamily Lagriinae (Tenebrionidae) is externally similar to species of the tribes Lupropini and Goniaderini having the robust pubescent body and estriate elytra, but differs in the absence of intersegmental...

A new species of the currently Oriental alleculine genus Oracula Novák, 2019 is described from the Eocene Rovno amber, Ukraine: O. campbelli sp. n. This is the first report of a representative of the family Tenebrionidae from this Lagerstätte. Extant species of Oracula occur in Nepal, southern China (Yunnan), Thailand, Laos and Indonesia. The newly...

The present state of the invasive population of Odocnemis aurichalcea (Adams, 1817) (Coleoptera: Tenebrionidae) in North Caucasus is studied. This species is widespread in Transcaucasia and has entered Nalchik and Nartkala towns (North Cau-casus, Kabardino-Balkaria, Russia) possibly during urban greening where it is mentioned in these localities si...

В мае 2022 года были проведены исследования по фауне жуков-чернотелок территории Рицинского реликтового национального парка и Пицундо-Мюссерского государственного заповедника. Было зарегистрировано 15 видов, из которых 3 впервые указаны для фауны Абхазии: Mycetochara gracilicornis, Gonodera macrophthalma, Scaphidema metallica. Составлен каталог чер...

Caucasian and Ciscaucasian representatives of the genus Opatrum Fabricius, 1775 are briefly reviewed. In total, four species and one subspecies occur in this area. Opatrum sabulosum sabulosum (Linnaeus, 1758) is widely distributed on all studied territory of the Caucasus up to 3000 m in Transcaucasia; O. sabulosum amauropterum Reichardt, 1936 occur...

A well preserved print of darkling beetle, belonging to the genus Promethis Pascoe, 1869 (Stenochiinae: Cnodalonini) has been collected in late Miocene Isarizawa Formation (Messinian, 7.246-5.333 Ma), at the shore of Toyosawa reservoir in Minamitoyosawa National Forest, Hanamaki city, Iwate Prefecture, Japan. The extant species of this genus are wi...

New data on the distribution, taxonomy and nomenclature of five Caucasian and/or southern European Russian tenebrionid species of the genus Microdera are presented. A new species, M. urartu sp. n., is described from Armenia and the Nakhchivan Autonomous Republic of Azerbaijan; this species is similar to the Central Asian M. globulicollis Ménétriés...

Aim. To reveal mutual relations between species of the genus Nalassus in taxocenes and the division of ecological niches during cohabitation. Material and Methods. Observations and collection of adult beetles and food objects were carried out in 2020–2022 in Dagestan, North Ossetia, Kabardino‐Balkaria, Karachay‐Cherkessia, Krasnodar Region (Russia)...

A taxonomic review of the tenebrionid genus Psammocryptus Kraatz, 1865 (Coleoptera: Tenebrionidae: Pimeliinae: Tentyriini) is given. Species of this genus are distributed in the south of Russia (Caspian depression), Transcaucasia, Kazakhstan and Central Asia, Afghanistan and Pakistan, occurring in areas with salty soil, often in tugay forests of ri...

A taxonomic review of the genus Euboeus from the Caucasus, Iran and Turkmenistan is given. All revised species belong to the subgenus Pelorinus Vauloger de Beaupré, 1900 and can be divided into two groups: the tenebricosus species-group and the huedepohli species-group. The first one consists of mainly forest low mountain species, while the second...

The taxonomic review of the genus Dendarus Dejean, 1821 from Iran and Turkmenistan is given. In total, five species are distributed in Iran and four species in South Turkmenistan; three species occur in both countries. The new species D. matthewsi sp. n. is described from Iran and compared with D. simplex Seidlitz, 1893. One taxon is resurrected fr...

A single species of the genus Odocnemis Allard, 1876 occurs in Cyprus and it is considered endemic to the island: O. intrusicollis (Seidlitz, 1895). The species was known only by the original description of one female (holotype) from the type locality “Cyprus”. We found two populations of this darkling beetle across the Troodos mountain range, at h...

A brief well-illustrated review of the tenebrionid genus Nalassus Mulsant, 1854 from the Greater Caucasus is presented. A new species, N. (Caucasonotus) negrobovi Nabozhenko, sp. n. is described from the alpine zone of Abkhazia. This species is similar to N. dombaicus (Nabozhenko, 2000), N. adriani (Reitter, 1922) and N. alanicus (Nabozhenko, 2000)...

This study is based on the material of darkling beetles (Tenebrionidae) collected from different localities of Turkey between 1966 and 2020 and deposited in Biodiversity Science Museum, Atatürk University (Erzurum, Turkey). In total, 80 species and subspecies of the family Tenebrionidae were listed. The following five species are recorded for Turke...

A taxonomic review of tenebrionid platyopoid genera of the subfamily Pimeliinae from Eastern Europe, Central Asia, Afghanistan, Iran and Pakistan is given. This group of taxa was known before 1994 as the tribe Platyopini Motschulsky, 1849, which is now interpreted as a junior synonym of Pimeliini Latreille, 1802. The group is different from other P...

A new species is described from the Central part of the North Caucasus (Russia): Blaps caucasica M. Nabozhenko et I. Chigray sp. n. This taxon was interpreted earlier as B. scabriuscula subalpina Ménétriés, 1832, but both our morphological and molecular genetic analyses showed that it is in fact a separate new species. The following new synonymies...

We present the life history and trophic relations of the lichenophagous darkling beetle Helops glabriventris glabriventris Reitter, 1885 (Coleoptera: Tenebrionidae: Helopini), which is widespread in Cyprus. Adults of H. glabriventris feed primarily on Pseudevernia furfuracea (L.) Zopf (Parmeliaceae), which is the most widespread fruticose corticolo...

A new species of comb-clawed beetles of the genus Asiomira Dubrovina, 1973 (A. dubrovinae sp. n.) is described from the Eocene Baltic amber. This newly descovered fossil species displays typical generic characters and is the most similar to the extant Asiomira ophtalmica (Seidlitz, 1896). Both species share a similar shape of the pronotum and the f...

The faunistic review of Tenebrionidae (excluding Alleculinae) from Armenia is presented. The work is based on longterm collections of one of the authors and materials from museum collections, deposited in Yerevan (the most of material), St Petersburg and Budapest. In total, 123 species of Tenebrionidae, excluding Alleculinae are reliably known from...

The new genus Doyenellus Nabozhenko and Steiner (Coleoptera: Tenebrionidae, Helopini) is recognised and described, and includes three North American species native to southeastern USA. The type-species, Doyenellus cisteloides (Germar), new combination, is diagnosed and two new species, Doyenellus worthleyorum Steiner, new species and Doyenellus the...

Turkonalassus mavi M. Nabozhenko & B. Keskin, sp. n. and Odocnemis rufocruralis M. Nabozhenko & B. Keskin, sp. n., two distinctive darkling beetle species of the tribe Helopini, are described from the Southeastern Anatolia region of Turkey, based on both their external morphology and genital structures. Turkonalassus mavi sp. n. differs from all co...

Ectromopsis merkli sp. n. (Coleoptera: Tenebrionidae: Helopini) is described from Central Anatolia (coast of Eğirdir Lake). The species is the most similar to E. tantilla Ménétriés, 1848 from the Caspian depression (European Russia, West Kazakhstan) and differs in the body shape, structure of genae, the shape of pronotum and elytral punctation. Bri...

The description of an extinct species of Metaclisa Jacquelin du Val, 1861 (Tenebrionidae) is presented. This genus and the tribe Metaclisini are recorded as fossils for the first time, from Eocene Baltic amber. The new species Metaclisa ottoi sp. nov. belongs to the subgenus Trichometaclisa subgen. nov. and differs from all other Metaclisini in pos...

Additional diagnostic characters of the little-known alleculine species Cornucistela serrata Campbell, 1980 are discussed. Variability of ocular index, a character widely used in the systematics of Alleculinae, is demonstrated. The distribution of C. serrata is amended; the species is known from the central part of Saudi Arabia and not from western...

Based on a single well-preserved specimen from Eocene Baltic amber, a new comb-clawed beetle Isomira lobanovi sp. n. (Alleculinae: Alleculini: Gonoderina) is described and illustrated. The new species belongs to the subgenus Mucheimira Novák, 2016, with extant species distributed in South China, Himalaya and the south of Arabian Peninsula. Isomira...

Species of the genus Helopelius Reitter, 1922 (type species: Stenomax aeneipennis Allard, 1876) were known from Rhodes and North Africa. We found that two species described from Rhodes belong to the genus Stenohelops Reitter, 1922, and Helopelius can be interpreted as a subgenus within latter, guided by the article 23.2 of ICZN. The following synon...

Reitter described two genus-rank taxa within the tenebrionid tribe Helopini (Tenebrionidae) in the same work: Euryhelops Reitter, 1902 (type species Helops tiro Reitter, 1902) as a subgenus of the genus Helops Fabricius, 1775 and Euryhelops Reitter, 1902 as a separate genus (type species Helops championi Reitter, 1891 = Helops subaeneus Reitter, 18...

A new tenebrionid species Ceratanisus diyarbakiricus sp. n. (Pimeliinae: Ceratanisini) is described from southeastern Anatolia. This is the most eastern representative of the mucoreus species-group. The species is similar to C. mucoreus (Waltl, 1838) from the Aegean region and C. purcharti Nabozhenko et al., 2016 from Taurus mountains. New distribu...

Full catalogue of the tribe Helopini (Coleoptera, Tenebrionidae) of the Palaearctic is published

Full catalogue (with synonymy, distribution of species) of the tribe Blaptini (Coleoptera: Tenebrionidae) of Palaearctic is published

The genus Scaurus Fabricius, 1775 in Turkey is briefly reviewed. A new synonymy is proposed: Scaurus macricollis Allard, 1882 = Scaurus puncticollis var. syriacus Reitter, 1914, syn. n. Scaurus julioferreri sp. n. is described from Eastern Turkey (Kars Province); it differs from all Caucasian, Near and Middle East Scaurus by granulated (without pun...

A new species of darkling beetles of the tribe Helopini is described from Southern Israel: Catomus chikatunovi n. sp. A new species differs from all Near Eastern Catomus Allard, 1876 in the presence of long sparse erected hairs on lateral margins of pronotum and elytra, the latter being otherwise bare dorsally. The new species exhibits arenicolous...

Two new species of the tribe Helopini (Coleoptera: Tenebrionidae) are described from Turkey: Nalassus (Nalassus) becvari sp. nov. (Elazığ Province) and Hedyphanes (Hedyphanes) kmenti sp. nov. (Artvin Province). The fi rst species is characterized by strongly thickened antennomeres 2-8 and diff ers from all Turkish Nalassus s. str. Mulsant, 1854 in...

Based on a single well-preserved specimen from Eocene Baltic amber, a new tenebrionid beetle Neomida groehni Nabozhenko et Bukejs sp. nov. (Diaperinae: Diaperini) is described and illustrated using phase-contrast X-ray microtomography. This oldest representative of the genus differs from all known extant and extinct congeners by the dorsally very w...

A brief review of the eight species of the genus Odocnemis Allard, 1876 from the Near East (Cyprus, Syria, Lebanon, Israel, Jordan, Egypt) is given. All species (excluding probably O. intrusicollis from Cyprus) belong to the praelonga species-group. Odocnemis evae sp. n. is described from Lebanon. The new species is similar to O. praelonga Baudi di...

The oldest tenebrionid beetle Palaeobasanus neli gen. et sp. n. of the tribe Scaphidemini (subfamily Diaperinae) is described from the Paleocene of Menat (Thanetian) in France. The new extinct genus shows a distinct complex of characters of the tribe Scaphidemini, such as modified procoxal cavities, modified prosternal "cowling" and mesocoxal cavit...

The fossil record of Tenebrionidae (excluding the Quartenary) is presented. In total, 122 fossil species, clearly belonging to the family, are known; some beetles were determined only to genus; 78 genera are listed in the fossil record, including 29 extinct genera. The great diversity of tenebrionids occurs in the Lower Cretaceous Lagerstätte of Ch...

The karyotype and cytogenetic features of darkling beetle Accanthopus velik-ensis were analysed using conventional and differential staining. The diploid number was determined as 2n = 20 and the presence of Xy p sex determination system was observed with DAPI and silver staining as well as conventional staining. Although a single nucleolar material...

Four new species of tenebrionid beetles of the tribe Helopini are described and fi gured: Nalassus (s. str.) kadleci sp. n., N. (s. str.) kataevi sp. n., Eustenomacidius (Caucasohelops) glebi sp. n. (all the three species are from Loristan Province) and Odocnemis (s. str.) denisi sp. n. from Zanjan and Gilan provinces. The new species of Nalassus M...

The tenebrionid species Blaps scabiosa (tribe Blaptini) is redescribed and illustrated. This species was twice described under the same name by Baudi di Selve in 1874 from Tashkent (now Uzbekistan) and by Faust in 1875 from Tash (Iran, Semnan Province). In 1876 Faust synonymized his species with Baudi’s species. Nabozhenko interpreted these taxa as...

The taxonomic history, composition, morphology, distribution, and bionomics of the genus Dila Fischer von Waldheim, 1844 are discussed. The following new generic synonymy is established: Dila Fischer von Waldheim, 1844 = Caenoblaps König, 1906, syn.n. As a result, four species are transferred from Caenoblaps to the genus Dila : Dila difformis (Köni...

New localities and data on distribution of darkling beetles of the tribe Helopini are presented. In total, 46 species from 12 genera (Adelphinus Fairmaire et Coquerel, 1866, Catomus Allard, 1876, Euboeus Boieldieu, 1865 (Pelorinus Vauloger, 1900), Helops Fabricius, 1775, Raiboscelis Allard, 1876, Entomogonus Solier, 1848, Hedyphanes Fischer von Wal...

Aim. In this work, we aimed to study the distribution and habitat of rare Caucasian tenebrionid beetle Hedyphanes nycterinoides, as well as to identify reasons for the population re‐ duction and to develop a basis for its protection. Methods . Materials from the largest collection of the Zoological Institute RAS (St Petersburg) along with the autho...

A list of six Jordanian species of the darkling beetles of the tribe Helopini is given. Entomogonus (Delonurops) amri Nabozhenko et Tichý, sp. n. is described from Jordan and Syria. Two new synonyms are proposed: Catomus (s. str.) acutipennis (Reiche et Saulcy, 1857) = Catomus laenoides Reitter, 1922, syn. n.; Catomus (s. str.) hesperides (Reiche,...

A second species of the previously monotypic genus Cornucistela (C. anichtchenkoi sp. nov.) is described, illustrated and compared with C. serrata Campbell, 1980 (type species). It is a first record of the genus in Iran (South Khorasan Province).

The new monotypic subgenus Stenolassussubgen. nov. of the genus Stenohelops Reitter, 1922 is described for Stenohelops klebsi (Nabozhenko, Perkovsky et Chernei, 2016) comb. nov. (Coleoptera: Tenebrionidae), which was previously described within the genus Nalassus Mulsant, 1856. This species is transferred to Stenohelops Reitter, 1922 based on the s...

A new tenebrionid genus Socotraphanes gen. nov. and two new species S. krali sp. nov. and S. dementor sp. nov. (Tenebrionidae: Helopini) are described from Socotra Island (Yemen). The genus differs from all Old World Helopini by hidden frontoclypeo-labral membrane, which originates under the inner side (not from the anterior edge) of frontoclypeus....

Based on well preserved specimen, Gonialaena groehni gen. et sp. nov. (Coleoptera: Tenebrionidae) is described from Eocene Baltic amber and assigned to Gonialaenini trib. nov. within the subfamily Lagriinae due to the combination of the following characters: labrum elongate; antennal sensilla simple; procoxal cavities closed at least externally; na...

The aim of the present study is to reveal the consequences of the blood ark (Anadara kagoshimensis) invasion of Taman Bay (the Sea of Azov), one of the biggest boreal brackish coastal lagoon. Anadara kagoshimensis was found there for the first time in 2003. During our work from 2003 to 2013, the blood arks were found at eleven of 83 stations in the...

Three new species of comb-clawed beetles of the tribe Alleculini (Tenebrionidae: Alleculinae) are decribed. Gonodera baygushevae Nabozhenko et I. Chigray, sp. n. (Eocene Baltic amber) is similar to extant Gonodera rufoaenea Reitter, 1900 by very finely and sparsely punctated elytral intertriae, but differs in the small size of body (6.6 mm, while o...

A brief history of classification of the tenebrionid beetles of the tribe Helopini Latreille, 1802 (Coleoptera: Tenebrionidae) is discussed. Until now, the system of Helopini had been interpreted in accordance with the four main classifications used independently by specialists from different regions. Three subtribes within the tribe Helopini are p...

A brief review of the genus Cryphaeus Klug, 1833 of the Russian Far East is given. Two species occur in this region: C. amurensis (Heyden, 1884) in the Primorsky and Sakhalin regions and C. albopilosus Chûjô et Lee, 1993 in the Primorsky Region (new record for Russia). A new synonymy is proposed: Cryphaeus amurensis (Heyden, 1884) = Anthracias duel...

A brief review of the genus Cryphaeus Klug, 1833 of the Russian Far East is given. Two species occur in this region: C. amurensis (Heyden, 1884) in the Primorsky and Sakhalin regions and C. albopilosus Chûjô et Lee, 1993 in the Primorsky Region (new record for Russia). A new synonymy is proposed: Cryphaeus amurensis (Heyden, 1884) = Anthracias duel...

Two new species of darkling beetles of the genus Nalas-sus are described: N. (s. str.) farsistanus sp. n. (Iran: Fars Province) and N. (Helopondrus) striatissimus sp. n. (Iran: Mazandaran Province). New faunistic records of little known species of the tribe Helopini (gen-era Nalassus Mulsant, 1854, Catomus Allard, 1876, Hedyphanes Fischer von Waldh...

The Red Data Book of the Kabardino-Balkarian Republic is an official publication containing information on the characteristics, distribution, ecological, biological and phytocenotic features, limiting factors and measures for the protection of plant species, fungi and animals, approved by the Government of the Kabardino-Balkarian Republic "On the l...

The taxonomic review of the tenebrionid genus Calyptopsis Solier, 1835 (Tenebrionidae: Tentyriini) of Turkey and Western Transcaucasia is given. Four new species and one new subspecies are described: C. capnisiformis volcanica subsp. nov. (from calderas of extinct volcanoes in Karaman and Manisa provinces), C. fouquei sp. nov. (Artvin Province), C....

A taxonomic review of the tenebrionid-beetle genus Calyptopsis Solier, 1835 from Kazakhstan and Middle Asia is given. In total, five species are listed: C. clypeata Faust, 1875 (Turkmenistan, Iran), C. deplanata Faust, 1875 (Turkmenistan), C. incerta Faust, 1875 (Turkmenistan, Iran), C. punctiventris Baudi di Selve, 1874 (Kazakhstan, Uzbekistan, Ta...

The genus Erulipus Fairmaire, 1903 (previously in the subfamily Tenebrioninae, tribe Helopini), after the study of the type species (Erulipus fruhstorferi Fairmaire, 1903), is transferred to the genus Ainu Lewis, 1894 (subfamily Stenochiinae, the tribe Cnodalonini) as a subgenus. Consequently, two new combinations are established: Ainu (Erulipus) f...

A new tenebrionid species, Toxicum elviraesp. n. (Tenebrioninae: Toxicini), is described from Primorskii Territory of Russia. The new species belongs to the subgenus Mutiloxicumsubgen. n. and differs from the congeners in the absence of horns on the male head (only supra-orbital swellings and a small epistomal tubercle are present), in the erect pu...

A taxonomic revision of the tenebrionid-beetle genus Hedyphanes Fischer von Waldheim, 1820 from Kazakhstan, Middle Asia, Iran and Afghanistan is presented. The history of the classification of this genus is described; the descriptions, synonymy, bibliography, data on the distribution and ecology, and a key to 19 species are given. Four new species...

A new species Entomogonus (Delonurops) doguanatolicus Nabozhenko sp. n. (Coleoptera: Tenebrionidae) is described from Diyarbakır (Turkey).It differs from all members of the subgenus Delonurops Reitter, 1922 in the flattened lateral sides of pronotum. The newly described is similar to E. clavimanus Reitter, 1903, E. makovskyi Nabozhenko et Tichy, 20...

A new tenebrionid species, Toxicum elvirae sp. n. (Tenebrioninae: Toxicini), is described from the Primorsky Krai of Russia. This new species belongs to the subgenus Mutiloxicum subg. n. and differs from congeners in the absence of horns on the male head (only supra-orbital swellings and a small epistomal tubercle are present), the erect pubescence...

A redescription of the male of Scaurus araxinus Richter, 1945 is given and the female is described for the first time. The species occurs in Azerbaijan (Nakhichevan Republic), Armenia (Tigranashen and Yeghegnadzor) and Eastern Turkey (first record in Iğdır province). It had been regarded as conspecific with S. syriacus Reitter, 1914 and S. punctico...

A taxonomic review of the genus Prosodes Eschscholtz, 1829 of Iran is given. Two new species from the subgenus Meropersina Reitter, 1909 are described, both from West Azerbaijan Province: P. kasatkini sp. n. and P. shokhini sp. n. Three new synonymies are proposed: Prosodes (Dilopersina) mithras Reitter, 1904 = Prosodes (Dilopersina) chorassanica G...

A review of the following Eastern Palaearctic (China, Russian Far East, Korean Peninsula, Japan and Taiwan) genera of the tribe Helopini (Coleoptera: Tenebrionidae) is given: subtribe Helopina: Nipponohelops Masumoto, Ando et Akita, 2006, Apterotarpela Kaszab, 1954, Catomus Allard, 1876, Stenohelops Reitter, 1922; subtribe Cylindrinotina: Nalassus...

The taxonomic history and position of the genera Euboeus and Probaticus within the tribe Helopini are discussed. A new synonymy, based on morphological characters is established: Euboeus Boieldieu, 1865 = Probaticus (s. str.) Seidlitz, 1896, syn. n. As a result, 67 new combinations (from Probaticus to Euboeus) are estabished for all species formerl...

The third stage larva of Nalassus sareptanus (Allard, 1876) (tribe Helopini), type species of the subgenus Helopondrus Reitter, 1922, is described and figured for the first time. The species has “cylindrinotoid” type of the aedeagus and female genital tubes but belongs to the “nalassoid” group of genera based on the structure of larval labrum, whic...

A review of tenebrionid beetles of the genus Anatolica Eschscholtz, 1831 of the Don River sands is given. In total, six species and one subspecies were registered: Anatolica lata (Steven, 1829), A. angustata (Steven, 1829), A. subquadrata (Tauscher, 1812), A. abbreviata Gebler, 1832, A. eremita (Steven, 1829), A. gibbosa gibbosa (Steven, 1829), A....