Laishram Kosygin

Laishram Kosygin
  • Doctor of Philosophy
  • Scientist at Zoological Survey of India

About

114
Publications
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469
Citations
Current institution
Zoological Survey of India
Current position
  • Scientist

Publications

Publications (114)
Article
Glyptothorax palakkadiensis, a new sisorid catfish, is described from the Chalakudy drainage in Kerala, Western Ghats, India. It is distinguished from its congeners in the Indian subcontinent in possessing the following combination of characteristics: the presence of numerous oblique branches of the laterosensory canal that are visible as cream-yel...
Article
Ompok malabaricus (Valenciennes 1840) was originally described from the Malabar, India. Earlier, this species was only found in the Indian peninsula, specifically in Maharashtra, Goa, Karnataka, Kerala along the Western Ghats Mountain range and Rajasthan. However, in this study, O. malabaricus is reported for the first time from West Bengal, and it...
Article
Ompok malabaricus was originally described from the Malabar. The fish was believed only to be found in the Indian peninsula, specifically in Maharashtra, Goa, Karnataka, and Kerala along the Western ghats mountain range. However, in this study, O.malabaricus is reported for the first time from Rajasthan, and its distribution is extended further nor...
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Glyptothorax chakpiensis is described from the Chakpi River, Chindwin drainage of Manipur, India. The new species is distinguished from its congeners in the Chindwin River in having a lanceolate-shaped thoracic adhesive apparatus, with a median depression almost enclosed posteriorly by skin ridges; a short nasal barbel not reaching the anterior mar...
Article
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Glyptothorax zeiladensis, a new catfish, is described from the Barak-Surma-Meghna River drainage in Manipur, India. It differs from its congeners by a combination of characteristics: densely tuberculate skin on the body; an elliptical thoracic adhesive apparatus, almost enclosed posteriorly with skin ridges extending over the entire apparatus inclu...
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Osteobrama peninsularis has been reported in southern Indian drainages, particularly in the Krishna River drainage. This study provides new findings of O. peninsularis in two distant locations: the Kangsabati River (Suvarnarekha River drainage) in West Bengal and Wyra Lake (Godavari River drainage) in Telangana. This marks the first record of O. pe...
Article
A new species of Garra (Labeoninae) is described from the Zubza River, a headwater of the Brahmaputra River basin, in Kohima district of Nagaland, North-eastern India. The new species is distinguished by the following combination of characters: a black spot located on the principal ray and first three branched rays of the dorsal (upper) lobe distal...
Article
Glyptothorax pongoensis, sp. n. is described from the Yongmon River in Nagaland, India. The new species can be distinguished from its congeners in the Indian subcontinent by the following combination of characters: an ovate-shaped thoracic adhesive apparatus with skin ridges present over the entire apparatus, including the subulate-shaped median de...
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An attempt has been made to prepare a systematic checklist of the fish fauna of Mujnai river, West Bengal. A total of 41 species of fish, which belong to 34 genera, 17 families, and 8 orders, have been recorded. The order Cypriniformes has the highest diversity with 23 species, followed by Siluriformes with 9 species. Among the families, the highes...
Article
Garra triangularis Shangningam, Rath and Kosygin,2021 was originally described from the Sakartod river at Khanvel, Dadra & Nagar Haveli, Western ghats, India. Later, it was reported from Odisha, Eastern ghats. It is reported here for the first time from Rajasthan.
Article
Glyptothorax heokheei, a new species of sisorid catfish is described from the Brahmaputra River drainage in India. It can be distinguished from all congeners, except G. dikrongensis, G. indicus, G. rugimentum and G. obliquimaculatus, in having the unculiferous striae of thoracic adhesive apparatus extending anteriorly onto gular region. Glyptothora...
Article
Glyptothorax primusplicae, sp. n., is described from the Pamtujang River, a tributary of the Chakpi River, Chindwin basin in Manipur, India. It is diagnosable by the following combination of characteristics: the presence of plicae on the ventral surface of the pectoral fin spine and on the first pelvic-fin ray; the dorsal fin located nearer to th...
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Glyptothorax siangensis, new species, is described from the Siang River in Arunachal Pradesh, India. The new species can be distinguished from its congeners from the Ganga-Brahmaputra drainage and its neighbouring drainages by the following combination of characters: a thoracic adhesive apparatus with skin ridges present over almost the entire appa...
Article
Glyptothorax siangensis, new species, is described from the Siang River in Arunachal Pradesh, India. The new species can be distinguished from its congeners from the Ganga-Brahmaputra drainage and its neighbouring drainages by the following combination of characters: a thoracic adhesive apparatus with skin ridges present over almost the entire appa...
Article
Parambassis lala was originally described from the Gangetic provinces in India. It is reported here for the first time from Maharashtra, India, and its distribution is extended to the Northern part of the Western ghats.
Article
Glyptothorax viridis, new species, is described from the Dujang, a hill stream tributary of the Chakpi River, Chindwin drainage in Manipur, India. It is distinguished from its congeners by the following combination of characters: presence of plicae on paired fins; thoracic adhesive apparatus with a deep, cone-shaped median depression opening caudal...
Article
The present study appraises the discrepancies patterns of morphometry for the first time on the three species of Garra. Garra arunachalensis, G. birostris, and G. quadratirostris were collected from various locations of the Siang River in Arunachal Pradesh, India. The results of the morphological analysis revealed that G. arunachalensis, G. birostr...
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The state of Meghalaya, in northeast India, has many caves in the Jaintia, Khasi and Garo Hills districts. As per the available reports, over 1000 caves have been explored or partially explored to yield over 450 kilometres of surveyed cave passage in the state. The true cave-adapted species or troglobionts mostly occupy the cave's deep zone, while...
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Olyra parviocula was originally described from the Kameng river, Brahmaputra river drainage in Arunachal Pradesh, India and was considered endemic in the state. It is reported for the first time from West Bengal, and its distribution is extended from Arunachal Pradesh to West Bengal.
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The occurrence of Garra triangularis Shangningam et al. (2021) is reported for the first time from Odisha, Eastern ghats. This fish has been reported earlier only from the type locality Sakartod River at Khanvel, Dadra & Nagar Haveli, Western ghats. The meristic counts, body measurements and descriptions are presented and the specimens have been co...
Article
Pethia chakpiensis sp. nov. is described from the Chakpi River of the Chindwin-Irrawaddy drainage in Manipur, India. This new species is characterized in having an incomplete lateral line with 4–7 pored scales, 21–24 scales in lateral series; 15 preanal scales; intercalated scale row originates above and after the 5th scale of lateral line scale ro...
Article
Pethia chakpiensis sp. nov. is described from the Chakpi River of the Chindwin-Irrawaddy drainage in Manipur, India. This new species is characterized in having an incomplete lateral line with 4–7 pored scales, 21–24 scales in lateral series; 15 preanal scales; intercalated scale row originates above and after the 5th scale of lateral line scale ro...
Article
A new catfish of the genus Glyptothorax is described from the Chakpi River, Chindwin drainage of Manipur, India. Glyptothorax waikhomi differs from its congeners in having a mottled skin, an oblanceolate leaf-shaped thoracic adhesive apparatus without a median depression, a serrated dorsal-fin and a short nasal barbel, not extending to anterior mar...
Article
A new species of sisorid catfish of the genus Exostoma is described from the Siang River in Arunachal Pradesh, northeastern India. The new species, Exostoma dhritiae, can be distinguished from congeners by the condition of the posterior extremity of the adipose-fin base, the degree of tuberculation on the dorsal surface of the head, and the shape o...
Article
During the taxonomic study of genus Mystus, a single deformed specimen collected from fish market in West Bengal, India was examined. The morphological features of abnormal specimens were compared with the normal specimen from the same location. The deformed specimen has its upper jaw shorter than the lower jaw; the front part of the head is abrupt...
Article
A new species, Aborichthys bajpaii, is described from the Siang River, Brahmaputra River drainage, in Arunachal Pradesh, India. It differs from all known congeners by the following combination of characters: a vent closer to the caudal fin base than to the snout tip, body with 17–22 fused bars along the mid-dorsal line, which are bifurcated into 28...
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Ichthyofaunal diversity of the Yu River in the Sagaing region of Myanmar is reported in this paper. A total of 49 species of fishes belonging to 38 genera and 19 families are recorded from the river. The region’s fish fauna includes two endangered, two vulnerable, one near threatened, twenty-nine least concern and three under data deficient categor...
Article
Glyptothorax yuensis, new species, is described from the Yu River, Sagaing division, Myanmar. It is characteristic in having a shallow adipose fin acutely incised at the posterior extremity of its base with an elongated pointed tip, adipose-fin base length 7.6-10.0 % SL; short nasal barbel, not extending to anterior margin of orbit; thoracic adhesi...
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The occurrence of Garra jaldhakaensis is recorded first in Uttarakhand, Northern India. The fish has been reported earlier only from the type locality Jaldhaka River near Jhalong, Kalimpong district, West Bengal, Brahmaputra River Drainage, India. The meristic counts, body measurements and descriptions are presented. The specimens have been compare...
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Freshwater crabs are considered as a valuable source of protein food in rural livelihood in northeast India. Although there is only little taxonomical work has been done for the freshwater crabs in the said area. The species Acanthopotamon fungosum (Alcock, 1909) has been reported for the first time from Zeilad Wild Life Sanctuary, Manipur on the b...
Preprint
A new species, Garra triangularis, is described from the Sakartod River at Khanvel, Dadra & Nagar Haveli, Western Ghats, India. The new species belongs to the proboscis species group and differs from the hitherto described congeners in having the following combination of characters: a triangular shaped labellum consisting of eight or nine transvers...
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Psilorhynchus magnaoculus sp. nov. (Teleostei: Psilorhynchidae) is described from the Yu River, Sagaing region, Myanmar. The new species is distinctive in having a squarish snout, a large eye, contributing 42.6−43.6 per cent of head length, almost equal to the length of snout and a notch on the ethmoid region of the snout.
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Garra jaldhakaensis sp. nov. (Teleostei: Cyprinidae) is described from the Jaldhaka River, Kalimpong district, West Bengal, India. The new species is distinguished from all its congeners in the Indian sub-continent in having a combination of the following characters: a prominent thick unilobed proboscis, protruding downward above the transverse gro...
Article
Sperata aorella (Blyth) is reported first from the Dadra & Nagar Haveli of Western ghats. The species is characteristic in having long supra occipital process, distinctly narrower anteriorly and rounded towards posterior, branched pectoral fin rays 9, branched anal fin rays 9 and gill rakers 20.
Article
Glyptothorax rupiri, a new sisorid catfish, is described from the Brahmaputra River basin in Arunachal Pradesh, northeast India. It differs from its congeners in the Indian subcontinent by the following combination of characters: the presence of plicae on the ventral surface of the pectoral spine and first pelvic-fin ray; a posteriorly serrated dor...
Book
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North-East Zone of India constitutes 5.2 per cent of total geographical area endowed with rich biodiversity. This zone depicts the faunistic and floristic resemblance of Indo-Malayan and Indo-Chinese bio-geographical regions. Zoological Survey of India made an attempt to compile all the available information on faunal diversity based on its surveys...
Chapter
North-East Zone of India constitutes 5.2 per cent of total geographical area endowed with rich biodiversity. This zone depicts the faunistic and floristic resemblance of Indo-Malayan and Indo-Chinese bio-geographical regions. Zoological Survey of India made an attempt to compile all the available information on faunal diversity based on its surveys...
Chapter
Full-text available
An updated systematic checklist of freshwater fishes of two biotic provinces of the northeast biogeographic zone of India has been provided. A total of 436 species under 117 genera, 42 families and 15 orders are documented here based on the present survey and available literature. Out of which 167 species are endemic to Northeast Biogeographic zone...
Article
Psilorhynchus brachyrhynchus Conway and Britz (2010) is reported for the first from India. The species is characteristic in having a short snout (43-48% HL), the scale-less ventral surface between paired fins, and features of dorsal and caudal-fin pigmentation. A total of 14 species of fishes belonging to 12 genera under 7 families and 4 orders are...
Article
A new species of glyptosternine catfish of the genus Exostoma is described from the Chakpi River, a tributary of the Chindwin-Irrawaddy drainage in northeastern India. The new species can be distinguished from congeners of the Chindwin-Irrawaddy drainage except E. chaudhurii and E. vinciguerrae in having an adipose fin confluent (vs. separate) with...
Article
This paper presents a first report of Glyptothorax igniculus Ng & Kullander (2013) from India. A checklist of the fish fauna of Chakpi River is presented based on the present collection and those reported by the earlier workers. A total of 90 species of fishes belonging to 53 genera and 21 families are recorded from the river. Based on the IUCN sta...
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A novel sisorid catfish, Pseudecheneis nagalandensis sp. nov., collected from the Tizu River, Chindwin Basin, Nagaland, northeast India is described. This new species differs from its congeners due to the presence of a long ray of pelvic fin that extends up to anal-fin origin, slender caudal peduncle and the thoracic adhesive apparatus with 13−14 t...
Article
Glyptothorax giudikyensis, a new species, is described from the Barak-Meghna-Surma River drainage in Manipur, north-east India. It is distinguished from its congeners except, G. scrobiculus in having a furrow running along the entire length of the ventral surface of the pectoral spine. However, the new species differs from G. scrobiculus in having...
Article
Glyptothorax kailashi, new species, is described from the Kaladan River drainage in Mizoram, northeast India. The new species can be distinguished from its congeners by the following combination of characters: a deep caudal peduncle; nasal barbel reaching the anterior margin of the eye or slightly beyond, maxillary barbel longer than head length; a...
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A new sisorid catfish, Glyptothorax distichus, is described from the Barak-Meghna-Surma drainage in Mizoram, North-east India. It is distinguished from its congeners in having a combination of the following characters: a chevron-shaped thoracic adhesive apparatus, with poorly developed median depression that is widely opened posteriorly by skin rid...
Article
The fish fauna of Kodayar river has been presented here based on an extensive survey and earlier reports. A systematic enumeration of fish fauna comprised 79 species representing eight orders, 23 families and 47 genera. The Kodayar river constitutes about 5.67 % to the endemic fish diversity of Western Ghats of India. Highest species diversity was...
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Pethia arunachalensis, a novel cyprinid species, is described from the Nao-dhing River, Brahmaputra Basin, Arunachal Pradesh, India. The new species is diagnosed in having an incomplete lateral line, 6-7 lateral-lined pored scales, absence of barbels and a distinct black caudal spot. A key to the species of the genus Pethia established in the Ganga...
Article
A new species of the genus Osteobrama is described from the Mahanadi River, Tikarpada, Angul District, Odisha state, India. Osteobrama tikarpadaensis, new species, differs from its congeners in having two pairs of minute barbels; iii–iv unbranched dorsal-fin rays with 25–33 serrae on the last unbranched ray; 15–16 branched pectoral-fin rays, and 25...
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Lepidocephalichthys arunachalensis is redescribed based on the specimens collected from the Brahmaputra drainage of Arunachal Pradesh, India. It can be distinguished from congeners in having the following combination of characters: a rounded or truncated caudal fin; body with 12-14 large, dark spots on the lateral sides; dorsal-fin origin conspicuo...
Article
Two new species of Aborichthys are described from the Pange River of the Brahmaputra River basin, in Arunachal Pradesh, northeastern India. Aborichthys kailashi, new species, is characterized in having a prominent dorsal and ventral adipose crest, 28-36 black bars on the body, an incomplete lateral line interrupted before pelvic-fin origin and vent...
Article
A new species of the genus Garra is described from the Ranga River of the Brahmaputra basin, Arunachal Pradesh, northeastern India. Garra magnacavus, new species, differs from its congeners in having an incipient proboscis with a transverse lobe, 15−19 rounded large pits on the snout, weakly developed non-fleshy central callous pad, and an elongate...
Article
Glyptothorax gopii, a new sisorid catfish, is described from the Kaladan River drainage in Mizoram, northeast India. It is distinguished from its congeners in the Indian subcontinent by the combination of the following characters: an axe-shaped anterior nuchal plate element extensively in contact with the posterior nuchal plate, plicae present on t...
Article
Garra simbalbaraensis, new species, is described from the Simbalbara River, Himachal Pradesh, India. The new species is distinguished from its congeners in having a combination of following characters: a prominent short, thick, unilobed and rounded proboscis, distinctly projecting downwards above the transverse groove; a black spot at the upper ang...
Article
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The Asian date mussel Arcuatula senhousia (Benson in Cantor, 1842) collected from Mandapam (Palk Bay), southeast coast of India is reported for the first time. The mussel were observed in the intertidal regions to depths of 2 m. The occurrence of the non-indigenous species, A. senhousia belonging to Mytilidae family is reported with its associate f...
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The species diversity of freshwater fishes from the transboundary river, Jaldhaka is still unknown to the scientific communities. We generated 40 DNA barcode sequences of 16 morphologically identified freshwater fishes and compared genetically with the database sequences. Ten species (Acanthocobitis botia, Barilius bendelisis, Crossocheilus latius,...
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Northeast India with two biodiversity hotspots is recognized as a biodiversity-rich region. However, several extant animals including chiropterans are currently at jeopardy due to habitat loss, electrocution mortality, and other anthropogenic threats. This study examines the efficacy of mitochondrial Cytochrome b (mtCytb) sequences for species-leve...
Article
A new nemacheiline loach, Aborichthys iphipaniensis sp. nov. is described from the Iphipani River in Arunachal Pradesh, Northeastern India. It can be distinguished from congeners in having the following combination of characters: vent closer to snout tip than caudal fin base, long caudal peduncle, shallow body, short pre-dorsal, small eye, lateral...
Article
Paracanthocobitis botia Hamilton, 1822 is reported for the first time from Balmiki Tiger Reserve, Bihar India. The species is distinguished from other members of the genus Paracanthocobitis in having a complete lateral line, axillary pelvic lobe absent and body colouration comprised of small dorsal saddles and 8–10 dark blotches. Distribution and m...
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Channa amari, a new ornamental species is described from Bhalka forest, West Bengal, India. It differs from congeners in having the following combination characters: absence of pelvic fins; 37 dorsal fin rays; 12-13 pectoral fin rays with 4-5 black bands; 23-25 anal fin rays; 10-11 caudal fin rays; 44-45 lateral line pored scales; 40-41 total verte...
Chapter
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A comprehensive list of the fish fauna of the Namdapha National Park and Tiger Reserve is provided here based on the present and earlier surveys conducted by the Zoological Survey of India. It includes a total of 50 species of fish belonging to 32 genera, 11 families and 4 orders. The fish fauna of the Namdapha National Park and Tiger Reserve is a...
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The greater hog badger, Arctonyx collaris is a venerable and widely distributed terrestrial carnivore mammal in Indo-Burma biodiversity hotspot. The population of A. collaris has largely declined in its range distribution due to the habitat loss and bushmeat crisis. The present study identified the amorphous burnt body parts of a mammal as A. colla...
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Nomenclature for the nameless is one of the successful events after the invention of the DNA barcoding technique in biodiversity research. The ornamental fish species are frequently hunted from the east and northeast India and trafficked with various elusive names and high demanding values. Such illegitimate trading stimulated the severe threats on...
Article
Psilorhynchus bichomensis, a new species, is described from the Bichom River in Arunachal Pradesh, northeast India. The new species is characterised by the following characters: body elongate with 46 scales along lateral line, unbranched pectoral-fin rays 9-10, predorsal scales 13-14, circumpeduncular scales 8 and caudal fin peppered with transvers...
Article
Psilorhynchus bichomensis, a new species, is described from the Bichom River in Arunachal Pradesh, northeast India. The new species is characterised by the following characters: body elongate with 46 scales along lateral line, unbranched pectoral-fin rays 9-10, predorsal scales 13-14, circumpeduncular scales 8 and caudal fin peppered with transvers...
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An updated assessment of the diversity and distribution of fishes in the Indian Himalaya revealed about 316 fish species which belong to 118 genera and 38 families under 13 orders. The assessed fish diversity includes 97 endemic fishes and 13 introduced species some of which are established in the Himalayan waters. The fish diversity known from the...
Article
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A new species, Olyra parviocula, is described from the Kameng River, Brahmaputra River drainage in Arunachal Pradesh, northeastern India. The new species differs from congeners in having small eye diameter 5–8% HL; short adipose fin, not confluent with caudal-fin, its base length 9–12% SL; dorsal-fin branched rays 6; anal-fin rays viii–xi, 8–10 and...
Chapter
The present study aims to update the freshwater fish fauna of Bilaspur district, Chhattisgarh. Through literature review and field surveys a total of 39 fish species belonging to 22 genera under 8 families are documented from the district.
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The present study aims to document the freshwater fish fauna of Guru Ghasidas National Park, Koriya district, Chhattisgarh. Through literature review and field surveys a total of 58 fish species belonging to 35 genera under 12 families are documented from the region in the present account.
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The present study aims to document the freshwater fish fauna of Barnawapara Wildlife Sanctuary, Raipur for the first time. The study provides an account of 14 fishes belonging to 11 genera under 5 families from the Sanctuary.
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A new cyprinid fish, Parapsilorhynchus swaini sp. nov., is described based on specimens collected from a stream nearHarisankar, Mahanadi River basin of Odisha, India. It differs from other species of the genus in having following combinationof characters: elongated and slender body (depth at dorsal fin origin 16.9-18.7% SL); narrow and slender head...
Article
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A new species Garra chindwinensis i.e., of cyprinid fish is described from the Chindwin basin in Manipur, northeast India. The details on their diagnosis, description, colour, morphometric date and distribution are also provided.
Article
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A new cyprinid fish, Parapsilorhynchus odishaensis, is described from Odisha, India. It is distinguished from its congeners by a combination of the following characters: poorly developed callous pad behind lower lip, which is not sharply delimited posteriorly; many prominent horny tubercles on the snout; deep head (depth at occiput 57.2–75.0% HL);...
Chapter
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abtract Indian freshwater fish diversity is very rich with as many as 1027 species, comprising of primary, secondary and alien freshwater fishes. Among them, primary freshwater fishes include 858 species belonging to 167 genera under 40 families and 12 orders. Further, 137 species of secondary freshwater fishes that frequently enter and thrive in f...
Article
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Garra tamangi, a new species of labeonine cyprinid fish is described from the Dikrong River, Brahmaputra River drainage in Arunachal Pradesh, northeastern India. Garra tamangi is distinguished from all its congeners in the Ganga-Brahmaputra River drainage in having roughly a triangular proboscis, trilobed with two small lobes that are more or less...
Article
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Glyptothorax senapatiensis, new species, is described from the Chindwin River drainage of Manipur, India. It is distinguished from its congeners by a combination of characters: deep body (depth at dorsal-fin origin 21.0-26.8 % SL, at anus 17.1-18.2 % SL); deep caudal peduncle (8.4-9.3 % SL); narrow head (width 72-82 % HL); short dorsal- fin spine (...
Article
A systematic checklist of freshwater fishes of Ganjam district of Odisha, India was provided. A total number of 97 species of fishes under 58 genera, 31 families and 9 orders has been recorded. Highest species diversity was observed in the Cyprinidae (36%) followed by Bagridae (9.2%). The fish fauna includes 2 Endangered (EN), 1 vulnerable (VU), 7...
Article
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Schistura paucireticulata, new species, is described from the Tuirial River in Mizoram, India. It is distinguished from its congeners by a combination of characters: 8-9 brown bars on the body, bars in front of the dorsal fin divided into 2-3 small bars forming a reticulations; basicaudal bar black, dissociated; dorsal fin spotted, with 4 simple an...
Article
Psilorhynchus rowleyi Hora and Misra 1941 known from the Chindwin–Irrawaddy drainage, formerly treated as a subspecies of the Ganga–Brahmaputra drainage species, Psilorhynchus homaloptera Hora and Mukerji 1935, is redescribed here based on newly collected materials and two syntypes in the Zoological Survey of India. It differs from the latter in ha...
Article
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A systematic, updated checklist of freshwater fishes of the Similipal Biosphere Reserve is provided with notes on status, distribution and habitat characteristics. It includes 66 species of fishes belonging to 42 genera, 19 families and 6 orders. A total of 24 species are reported here for the time from the reserve. As per IUCN status the fish faun...
Article
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A new freshwater fish Aborichthys waikhomi is described here from the streams of Noa-Dihing river, Brahmaputra basin in Arunachal Pradesh, India. It is distinguished from all other known species of Aborichthys in having truncate caudal fin with irregular black blotches and 12-16 oblique black bands on body, which are narrower than interspaces in ca...
Chapter
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This paper presented a checklist of the fish fauna of Tlawng river, Mizoram, based on present collection and those reported by the earlier workers. In total, sixty-four species of fishes representing thirty-four genera and fifteen families have been recorded from the river. Garra manipurensis Vishwanath & Sarojnalini and Tor barakae Arunkumar & Bas...
Article
An attempt was made to study the physico-chemical and biological characteristics of Awang Khujairok river, Manipur. Well mark seasonal variations were recorded in respect of physico-chemical characteristics. the collection included 11 species of fishes (belonging to 9 genera and 6 families), 3 species of snails (belonging to 3 genera and 3 families...

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