Khang Sinh Nguyen

• Researcher at The Institute Of Ecology and Biological Resources

The Himalayas represent a complex mountain system housing some of the world's richest floras along with a high level of endemism. Among them, Lysionotus (Gesneriaceae) stands out as a small genus (~34 species) that is unexpectedly distributed across southern to eastern Asia. Within this genus, the mountain forests of the south Pan-Himalaya region e...

A detailed description and illustrations are provided for the new species, Aschistanthera phucanhensis, from Phu Canh Nature Reserve in Hoa Binh province, Northwest Vietnam, are presented in this work. This marks the second known species within the genus. The novel species is differentiated from its related species, the type species of this genus,...

The paper presents new data on the orchid diversity of Vietnam obtained mainly during the fieldwork of 2023–2024, as well as a result of investigation of the herbarium and garden collections of Russia and Vietnam. It includes five species new to science, namely Cleisostoma truongquanii, Cymbidium bavuongii, Hemipilia uniflora, Micropera vietnamensi...

Begonia triquetra, a new species endemic to the limestone karst regions of Son La province in Northwestern Vietnam, is described and illustrated here. This new species resembles B. xenos in its suberect habit, subpeltate lamina, 4-tepaled staminate flowers, and 2-locular ovary. However, B. triquetra differs significantly from B. xenos in having tri...

This paper continues to publish new discovered from successive investigation of the species diversity of the Aspidistra genus in recent years. Six new species of Aspidistra (Asparagaceae) are described and illustrated: four from China, Aspidistra dentosa, A. dissecta, A. dongxingensis (SW Guangxi Zhuang Autonomous Region) and A. anlongensis (SW Gui...

Begonis laxiflora is described as a new species endemic to Vietnam from granite boulder habitats in the Dakrong district. It is most similar to B. abbreviata in its creeping habit, broadly ovate leaves, persistent bracts, and the number of tepals in both staminate and pistillate flowers. The new species is different in its inflorescence having inte...

Oreocharis hapii, a new species endemic to Quang Nam Province in central Vietnam, is described and illustrated here. This species bears a superficial resemblance to O. cotinifolia in its elliptic leaves, campanulate corolla tube, and the stamens protruding from the tube. However, the new species differs in its stoloniferous habit (vs. absent stolon...

Sonerila quangnamensis, a new species from Truong Son Mountain Range, Vietnam, is described and illustrated here. Like S. plagiocardia, S. quangnamensis has obliquely ovate leaves, which are unequal in size within each pair. However, S. quangnamensis is obviously distinct in several characters: the cross-section of its stems and petioles is terete...

Ophiorrhiza liuyanii, a new species from south-western China and northern Vietnam, is described and illustrated. The new species is characterised by the glabrous surfaces on almost all plant parts, congested inflorescences and broad-ovate to ovate bracts 9–22 × 4–11 mm. It morphologically differs from the closest species, O. baviensis, mainly in mo...

Mitreola sphaerocarpa (Loganiaceae), a species occurring in Peninsular Malaysia, is newly reported for the flora of Vietnam. The description, photographs, information on ecology, preliminary conservation status and taxonomic notes are also presented. In order to facilitate identification, we provide an identification key and checklist for the 4 kno...

The genus Xanthocyparis was described in 2002 for a new conifer species X. vietnamensis from northern Vietnam, which became well‐known for its rarity. Recently, natural diploid populations were found in a small area of northern Guangxi, China, whereas material from Vietnam plus newly discovered populations from southern Guangxi were determined to b...

This study establishes two new species, Aspidistra tenuistyla and A. luteo-rubra (Asparagaceae, Convallarioideae). The first one, Aspidistra tenuistyla, is described here as a new species for science. It was discovered in the lowland limestone area of Ba Be National Park (Bac Kan Province). A detailed illustrated description of this species, along...

Hiptage yangshuoensis K.Tan & K.S.Nguyen, a new species of Hiptage collected from a karst cliff close to the Lijiang River, Northeast of Guangxi Zhuang Autonomous Region, China, is described and illustrated based on molecular and morphological data. Hiptage yangshuoensis shares some morphological similarities with the H. multiflora F.N.Wei, but eas...

The paper presents new data on the orchid diversity of Vietnam obtained mainly in the course of the fieldwork in 2020–2023, as well as from the investigation of the collections stored at the Herbaria of Russia and Vietnam. It includes descriptions of four species (Appendicula spicata, Dendrobium khanhii, D. vietii, Listera anthropophora) and one va...

Primulina hoangmongii, a new species from Yen Bai Province of northern Vietnam, is described and illustrated. It is similar to P. albicalyx in its robust rhizome, rosette leaves, and yellow flowers. However, P. hoangmongii is clearly distinguished by its linear to narrowly-lanceolate bracts, 8–12 × 2–3 mm (vs. narrowly ovate to ovate, 18–25 × 9–14...

This paper continues the publication of Aspidistra species newly discovered in the flora of Vietnam. It includes an illustrated description of A. pentagona , found in the northern part of the country. Detailed analytical color illustrations, data on morphology and ecology, phenology, tentative relationships, distribution and expected conservation s...

39 Novelties in Aspidistra (Asparagaceae, Convallarioideae) with erect stems in the flora of Vietnam: A. quanii, a new species, and A. erecta, a new national record Abstract The paper reports two species of Aspidistra newly discovered in the flora of Vietnam. It includes a description of A. quanii, a species new to science found in the central part...

This paper continues the publication of Aspidistra species newly discovered in the flora of Vietnam. It includes an illustrated description of A. pentagona, found in northern parts of the country. Detailed analytical color illustrations, data on morphology and ecology, phenology, tentative relationships, distribution, and expected conservation stat...

A new monotypic genus, Quangnamia (Hamamelidaceae), and its species, Q. syncarpa, discovered in central Vietnam are described and illustrated. Quangnamia syncarpa is evergreen treelet or tree with non-stellate in-dumentum on vegetative and floral parts, glabrous twigs and leaves, axillary inflorescences in form of spadix-like many-flowered heads, h...

With the aim of inferring phylogenetic relationships among 86 species (including 45 species from Vietnam) mostly of the subfamily Convallarioideae (=Nolinoideae) (Asparagaceae sensu APG IV), we analyzed their chloroplast DNA sequences (rbcL and trnL-F) by both Bayesian inference (BI) and maximum likelihood (ML) methods. Our dataset included six of...

Understanding the driving forces of speciation is a central question in evolutionary biology. Cycas bifida (Dyer) K. D. Hill and Cycas micholitzii Dyer are two morphologically similar Cycas (Cycadaceae) species with deeply pectinate megasporophylls and dichotomously divided leaflets. Geographically, they are isolated by the Red River Fault Zone (RR...

A synopsis of the orchid genus Didymoplexiella is presented for Cambodia, Laos and Vietnam. Uncertainties in morphological delimitation of the species of Didymoplexiella are addressed in the context of the new data on variation of labellum structure; re-identification of a considerable number of specimens is performed in accordance with the current...

The seventh paper of the series on orchid diversity in Vietnam includes descriptions of four species new to science (Goodyera umbonata, Myrmechis brachyscapa, Rhomboda obcordata, and Vietorchis proboscidea) and five new national records (Calanthe speciosa, Habenaria coultousii, Myrmechis urceolata, Paphiopedilum barbigerum var. barbigerum, and Peri...

Meistera muriformis (Zingiberaceae: Alpinioideae), a new species from southern Vietnam is described and illustrated. The description is accompanied by additional notes on etymology, ecology and phenology, distribution, and preliminary IUCN assessment. Notes on differences from morphologically similar congeners are provided. The new species is diagn...

The paper presents an annotated checklist of the genus Raphiocarpus (Gesneriaceae) in the flora of Vietnam with a key for identification of all 13 species hitherto recorded in the country, including newly described species. The data for each species include the accepted name, synonyms, citations of relevant regional literature, typification, inform...

Two new species, Peliosanthes annamensis and P. virescens from Dak Lak Province in southern Vietnam, are described and illustrated. The former is morphologically similar to P. crassicoronata, while the latter to P. argenteostriata. We also describe and illustrate one new variety, P. nivea var. acutiloba from Khammouane Province, central Laos. Pelio...

Peliosanthes linearifolia, a new species of Asparagaceae, is described and illustrated. The species was discovered in 2022 in Quang Nam Province, southern Vietnam. Peliosanthes linearifolia is remarkable for its linear to narrowly lanceolate leaves that are less than 1.3 cm wide. Such leaves were previously known only in P. graminea. Based on the f...

Phytochemical study on the leaves of Amentotaxus yunnanensis led to the isolation of seventeen phenolic compounds including sixteen neolignans and lignans, and one flavone glycoside. Three among the isolates were previously unreported neolignans and named as amenyunnaosides A-C, respectively. Their structures were elucidated by extensive analyses o...

This paper presents new data on the orchid diversity of Vietnam obtained from field explorations made mainly in 2015–2022, as well as from the study of some historical collections housed at the Herbaria of Russia and Vietnam. It includes descriptions of two species new to science (Coelogyne phitamii and Cymbidium sangii), reports of one genus (Stig...

Ophiopogon muongnhensis, a new species from Dien Bien Province, NW Vietnam, is described and illustrated. In having slenderly petioled clavate leaves it is most similar to O. griffithii originally described from NE India, but clearly distinguishable mainly by its wider (1.8–2.9 vs. 0.3–1.9 cm) leaf blades, wider (3.5–7.5 vs. 2–2.8 mm), obliquely (v...

A new species of Hedyotis L. (Rubiaceae), Hedyotis konhanungensis B.H. Quang, T.A. Le, K.S. Nguyen & Neupane, is described and illustrated from the central highlands of Vietnam based on morphological and phylogenetic evidence. The new species belongs to the morphologically diverse tribe Spermacoceae (ca. 1000 species) of the family Rubiaceae, which...

Chase et al. (2009) attempted a subfamilial classification for the expanded asparagalean families Amaryllidaceae, Asparagaceae and Xanthorrhoeaceae sensu APG III (2009). In Asparagaceae they recognized seven subfamilies including Nolinoideae Burnett (1835: 985, 1095, 1106). The Nolinoideae circumscribed by them consisted of 16 genera including Noli...

Parahellenia trongduyii, a new species of Costaceae from Vietnam, is described and illustrated. The new species is similar to P. tonkinensis, but differs in unbranched or barely branched stems, narrowly triangular-lanceolate glabrous buds, bract apex with a soft mucro, yellow to whitish yellow labellum with longitudinal orange to red lines at cente...

Raphiocarpus taygiangensis , a new species of Gesneriaceae family discovered in Tay Giang District, Quang Nam Province, Central Vietnam, is here described and illustrated. The new species is diagnosed by the combination of its stem up to 2 m long, sericeous hairs on young stem, leaf petiole and adaxial mid-vein, sparsely and minutely serrate leaf m...

This paper continues the publication of new data obtained during field studies in remote and poorly studied regions of Laos in 2017-2022. It provides descriptions of one species and one variety new for science (Oberonia sinuata Aver. and Cleiso stoma fuerstenbergianum Kraenzl. var. minutiflorum Aver.) and reports five species newly recorded in the...

Ceriscoides glabra, a new species of the tribe Gardenieae (Rubiaceae), is described and illustrated. The species was discovered in 2017–2018 in Kon Chu Rang Nature Reserve (Gia Lai Province, southern Vietnam), in the vicinity of the locally well-known K50 waterfall, also known as Hang En waterfall. The new species differs from its congeners by the...

Leptomischus multiflorus, a new species of the tribe Argostemmateae (Rubiaceae), is described and illustrated. The species was discovered in 2022 in Quang Nam Province, southern Vietnam. The new species is characterized by rich indumentum on almost all plant parts, distylous flowers, and minutely puberulent ovary disk. Leptomischus multiflorus diff...

This paper presents new data on the orchid diversity of Vietnam obtained in 2020–2022. It comprises descriptions of two species new for science (Cleisostoma tatonii and Sarcoglyphis hiepii) and descriptions of four species new for the flora of Vietnam (Cleisostoma aspersum, Gastrochilus nanus, Micropera mannii and Trichoglottis ramosa). All species...

Millettia is a complex genus of the tribe Millettieae (Fabaceae). In this paper we provide a synopsis of the species of the genus from Vietnam, based on the examination of literature and specimens as well as field surveys. Millettia boniana and M. setigera are proposed as new synonyms of Derris tonkinensis and M. cubittii, respectively. Millettia d...

Hedyotis konhanungensis B.H.Quang, T.A.Le, K.S.Nguyen & Neupane (Rubiaceae, Spermacoceae) , a new species from Vietnam, is described and illustrated based on morphological and phylogenetic evidence. The new species is morphologically distinct from all southeastern Asian Hedyotis L. in its set of morphological traits such as the leaf type (shape and...

Tupistra annamensis, a new species from central Vietnam is described and illustrated. In the habitats it occurs as an epiphyte or a lithophyte. Having a pendulous scape, this species is similar to such species as T. fungilliformis and T. clarkei, but differs distinctly from them and all other members of this genus by its outer and inner floral brac...

Tupistra annamensis, a new species from central Vietnam is described and illustrated. In the habitats it occurs as an epiphyte or a lithophyte. Having a pendulous scape, this species is similar to such species as T. fungilliformis and T. clarkei, but differs distinctly from them and all other members of this genus by its outer and inner floral brac...

Argostemma tortilobum, a new species endemic to Vietnam, is described and illustrated. The species is morphologically most similar to A. sarmentosum, but differs in having erect stem without runners, densely pilose peduncle and twisted corolla lobes with ciliate margins.

Zanthoxylum avicennae (Lam.) DC. is a medicinal and aromatic plant, belong to the Zanthoxylum genus. The plant is found in some Asian contries, including Viet Nam. The various parts of this plant has been used to medicine and contain essential oils. In this study, the essential oil of Z. avicennae was isolated. Two essential oils from the leaves an...

Exploring the driving forces promoting speciation is a key question in evolutionary biology. Cycas bifida and C. micholitzii are two leaflets dichotomously divided Cycas species that have morphological similarities. Their distribution areas are separated by the RRFZ, a biogeographical barrier for Cycas species. The hypothesis for explaining their d...

Raphiocarpus taygiangensis , a new species of Gesneriaceae from Tay Giang District, Quang Nam Province, Central Vietnam, is here described and illustrated. The new species is diagnostic by the combination of its long stem, serrate leaf margin, purple spots, purple longitudinal lines, and glandular short hairs inside corolla. Morphological distincti...

This paper presents new data on the orchid diversity of Vietnam obtained in 2015–2021. It contains descriptions of three new species (Cleisostoma furcatum, Malleola micrantha, Schoenorchis hungii), one new variety (Chiloschista parishii var. minutiflora) and one new form (Cleisostoma furcatum forma albiflora). Additionally six species, Ascocentrops...

Lysionotus hagiangensis, a new species from northern Vietnam, is described and illustrated. It morphologically resembles L. chatungii and L. petelotii, but differs markedly by the character of leaf, inflorescence, corolla color, the shape of stamens, and the number of well developed staminodes. We provide detailed information, including morphologic...

Five new species of Urophyllum Wall. endemic to Cambodia and Vitnam are herein described and illustrated, and U. annamense (Pierre ex Pit.) Yooprasert, Culham & Utteridge, stat. nov. is raised to species status. The new species are unique due to the morphological combination of indumentum, secondary venation, stipule shape and inflorescence structu...

Peliosanthes luteoviridis is described and illustrated as a new species from southern Vietnam (Dong Nai Province, Cat Tien National Park). It is most similar to P. macrostegia in having stamens forming a hemispheric corona and a pistil somewhat abruptly narrowing to a conoid style, but differs mainly in dense inflorescence, terete peduncle, entirel...

We transfer Chirita crassifolia to Primulina as P. crassifolia and provide detailed amended descriptions for this species and for P. quanbaensis, both species found in the karst habitats in the Bat Dai Son mountains in Ha Giang province, northern Vietnam.

Previous studies recognized three major lineages of the family Costaceae: a South American clade, an Asian clade and a Costus clade. However, the genus Hellenia within the Asian clade has been shown to be non-monophyletic and its morphology has not been studied carefully. Therefore, the complete plastid genomes of Hellenia species were obtained and...

Alpinia is considered the largest genus of Zingiberaceae with ca. 250 species occurring in tropical and subtropical Asia, Australia, and Pacific Islands. The historical biogeography of Alpinia was conducted to explain where Alpinia originated and how it migrated to other regions. The phylogeny, divergence times and ancestral area reconstruction of...

A new species of Aspidistra, A. sonlaensis, is described and illustrated from Son La Province in northern Vietnam. The new species is morphologically close to A. micrantha, but differs in number of perigone lobes, anther position and pistil shape. The new species is accessed as Data Deficient (DD) according to the IUCN categories and criteria.

This paper continues the publication of Aspidistra species newly discovered in the flora of Vietnam. It includes illustrated diagnosis of two species new to science, A. liliiflora and A. sessilistigma found in northern and central parts of the country. Detailed analytical color illustrations, data on morphology, ecology, phenology, tentative relati...

A new species of Rubiaceae, Spiradiclis detianensis is described from a limestone karst area of southwestern China. This new species is morphologically similar to S. cordata and S. spathulata . All of them have rosetted habit and long peduncles, but it differs from the former by the cuneate leaf bases (vs. basally cordate) and much longer corolla t...

Two new species of Ophiopogon, O. dolichocaulis from Son La Province and O. trigonantherus from Cao Bang Province in northern Vietnam, are described and illustrated. The former is similar to O. chingii, while the latter is similar to O. rupestris and O. dracaenoides. This paper not only details their taxonomic features and relationships to similar...

This research provides a description and illustrations for a new species from the Phong Nha-Ke Bang National Park, Cynanchum quangbinhense. The distribution, habitat, phenology and conservation assessment for the species are also presented. The new species differs from closely related species, C. ovalifolium in many characters such as leaf shape, i...

The chemical composition and anti-inflammatory activity of the endemic Lysimachia baviensis were investigated for the first time in this study. A phytochemical fractionation of the methanol extract of L. baviensis resulted in the isolation of a new stilbene (bavienside A, 1) and two new chalcone glycosides (baviensides B and C, 2 and 3). Their stru...

Hemiboea chanii , a new species of Gesneriaceae from Ha Giang Province, northern Vietnam, is here described and illustrated. It has many branched stems, diamond-shaped involucre with two cirrose opposite apices, a pink corolla, red spotted inside, and a flowering time in January-February. Among congeners with an externally hairy corolla, this new s...

The paper provides additions to the known diversity of Begonia in Vietnam and Laos. One species new to science from Lao PDR (B. vivipara) and three new species from Vietnam (B. aspersa, B. gracilifolia, B. maculifolia) are described. Three species (B. rotundilimba, B. versicolor, B. wenshanensis) are reported as new records for the flora of Vietnam...

Peliosanthes longiracemosa, a new species from limestone areas of northern Vietnam, is described and illustrated. It is most similar to P. macrostegia in having a distally incurved staminal corona and a distinct pyramidal style, but clearly distinguishable chiefly by the broader leaf blades with more longitudinal veins and closer transverse veinlet...

Essential oils from roots, flowers, fruits, and leaves plus stems of Angelica sinensis cultivated in Vietnam were obtained by hydrodistillation and were analyzed using GC/MS-FID. In total, 36, 30, 33, and 27 compounds were found accounting for 98.7%, 98.8%, 99.2%, and 100% of the oils compositions, respectively. (Z)-Ligustilide (49.4%, 29.6%, 25.8%...

Alpinia laosensis was previously placed in synonymy under A. conchigera. However, a critical examination of herbarium specimens (including type material) and living plants of A. laosensis and A. conchigera reveals that they are two separate species. Alpinia laosensis can be distinguished from A. conchigera by the shape of its fruits, shape and size...

We describe and illustrate Ophiopogon robustus and Peliosanthes rubra from northern Laos and P. convallarioides from central Vietnam as species new to science. Ophiopogon robustus is close to O. platyphyllus from southern China, P. rubra is similar to P. densiflora from Laos and P. convallarioides is close to P. argenteostriata from Laos and Vietna...

Begonia catbensis , a new species in Begonia sect. Coelocentrum is described and illustrated. The new species was discovered in lowland limestone hills at Cat Ba National Park and can be easily distinguished from all its congeners by having dendritic hairs on the petiole, adaxial veins and stipules, fimbriate bracts and bracteoles, dense conical bu...

Hemiboea chanii , a new species of Gesneriaceae from Ha Giang Province, northern Vietnam, is here described and illustrated. It has many branched stems up to 90 cm, with 12–16 nodes, involucre diamond with two cirrose opposite apexs, pink corolla with red spotted inside, and flowering time in January–February. Among congeners with abaxially hairy c...

Leptomischus anisophyllus, a new species of Rubiaceae, is described and illustrated, and its conservation status is discussed. The species was discovered in 2017 during a botanical survey of Pu Mat National Park (Nghe An Province, Central Vietnam). The new species is remarkable by its anisophylly, a rare feature in Leptomischus. Besides, Leptomisch...

Aspidistra peltata, a new species morphologically allied to A. tonkinensis (= A. fungilliformis subsp. formosa; A. formosa) discovered in northern Vietnam, is described and illustrated. From the closely related A. tonkinensis, the new species differs in having an erect stem 4–14 cm tall, much smaller leaves, smaller flowers, filamentose stamens, a...

Mouretia oblanceolata, discovered in central Vietnam, is described as a new species of the tribe Argostemmateae in the family Rubiaceae. The new species is most similar to Mouretia tonkinensis and M. vietnamensis but differs from them by the following traits: erect to sub-erect habit, abaxially glabrous, obcordate stipules, oblanceolate leaves, 12–...

Rohdea dangii, a new species from limestone areas of Son La and Dien Bien Provinces in northwestern Vietnam, is described and illustrated. It is apparently most closely related to R. tonkinensis described from northern Vietnam, for these two species share an elongate aerial stem with no cataphylls (except for those at the base of the peduncle), nar...

Aspidistra magnifica, a new species from Mount Cà Đam, Quảng Ngãi Province, Central Vietnam, is described and illustrated. It is similar to A. maguanensis in having purple-black flowers, twelve perigone lobes and stamens, and a table-shaped pistil, but differs by its somewhat elliptic leaf blades, short flowering peduncles, much large broadly campa...

The paper describes a new species of Aspidistra discovered in Laos, namely A. nitens Aver., K. S. Nguyen et Tillich in addition to ten species (A. elegans, A. glandulosa, A. khangii, A. laongamensis, A. laotica, A. lurida, A. medusa, A. purpureomaculata, A. globulifera, A. subrotata) earlier reported in this country. The new species is most similar...

This study presents the morphological characteristics, colour illustration, biological information, ecology and distribution of Aspidistra papillata G.Z. Li, formerly considered as an endemic species to China, but recently discovered and recorded for the Flora of Vietnam, to identify new aspects of this species.

The present study reports newly recorded species of vascular plants for the flora of Vietnam found in the recently established Bat Xat Nature Reserve in Lao Cai province, close to the border with Yunnan province of China. Thirteen species belonging to eleven families are reported: Acanthaceae (Strobilanthes helicta), Actinidiaceae (Actinidia mellia...

This paper continues the publication of Aspidistra species newly discovered and reported in the flora of Vietnam. It includes illustrated diagnosis of one species new for science, A. angustata Aver., K.S. Nguyen & Tillich and notes on two species, A. chunxiuensis C.R. Lin & Yan Liu, and A. guangxiensis S.C. Tang & Yan Liu newly recorded in the coun...

Leptomischus hiepii , a new species of the tribe Argostemmateae from Son La province, northwestern Vietnam, is described and illustrated. Morphologically it allies to L. wallichii , L. erianthus and L. funingensis by sharing a similar habit, large stipules and similar corolla shape, but it differs by its anisophyllous leaves, 1-flowered inflorescen...

The paper presents new data on the orchid diversity in Vietnam obtained in 2016-2020. It contains descriptions of one genus (Apetalanthe) and two species new to science (Apetalanthe gracilis, Nervilia appressifolia), as well as one genus (Neottianthe) and four species (Calanthe tricarinata, Collabium yunnanense, Dendrobium praecinctum, Neottianthe...

his paper continues the publication of newly obtained results from a continuing taxonomic investigation of the genus Aspidistra in Laos and Vietnam. It includes illustrated diagnoses of two new species, A. melanaster Aver., K.S. Nguyen & Tillich, A. obliqua K.S. Nguyen & Aver., two new varieties, A. semiaperta Aver. & Tillich var. globulifera Aver....

The border area between south-eastern Yunnan, China and northern Vietnam is one of the regions with richest biological diversity including that of the fern genus Angiopteris (Marattiaceae). Based on the analysis of morphology and DNA sequences of multiple chloroplast regions ( atpB , rbcL , rps4-trnS spacer and trnL-F spacer), we revised Angiopteri...

We describe a new species, Peliosanthes cupuliformis, from central Laos with photographic illustrations and relevant taxonomic data. The species appears to be a local endemic confined to a karstic limestone area of Khammouane Province (Boualapha District). It is readily distinguishable from all other congeners by a suit of unique traits such as the...

Three groups of beetles inhabit cones of cycads (Cycadales) in the northern hemisphere and are believed to be involved in their pollination. The primitive weevil subtribe Allocorynina (Coleoptera: Belidae) is restricted to the New World cycad genera Dioon Lindl. and Zamia L. One group of weevils (Curculionidae), found only in Cycas L., appears to b...

The Bat Dai Son Mountain area belongs to Bat Dai Son Nature Reserve and additionally includes allied territories of Quan Ba District in Ha Giang Province of northern Vietnam. The study of the area was based on cooperation between the Institute of Ecology and Biological Resources (Vietnam Academy of Sciences and Technology) and the Komarov Botanical...

Tupistra nganii is described and illustrated as a new species from Ha Giang Province in northern Vietnam and Yunnan Province in southwestern China. The new species is most similar to T. muricata from Thailand and Laos, from which it differs by the greenish white or yellow perianth and longer exserted stamens. Additionally, Tupistra longispica is re...

HERBARIUM COLLECTIONS FROM BAT DAI SON MOUNTAIN AREA (NORTHERN VIETNAM, HA GIANG PROVINCE, QUAN BA DISTRICT) MADE IN 1995–2019 Averyanov L.V., K.S. Nguyen, T.H. Tran, A.L. Averyanova, H.T. Nguyen, T.V. Maisak. The document contains labels for 2569 collecting numbers made in Bat Dai Son Mountain area administratively belonging to Quan Ba District, H...

Two new neolignan sesquiterpenoids, chevalierinol A (1) and chevalierinol B (2), were isolated from the ethyl acetate extract of Magnolia chevalieri leaves, together with twelve known compounds (3–14). Their structures were elucidated by analysis of spectroscopic data including 1D, 2D and mass spectra and compared with the reported data. Compounds...

Ophiorrhiza hiepii and O. hainanensis are reported as a new species and a new record for Vietnam respectively. O. hiepii is morphologically similar to O. subrubescens but differs by its smaller habit, usually unbranched stems, fewer secondary veins, umbelliform and sub-congested cymes, shorter corolla tubes and shorter stigma lobes in short-styled...

Peliosanthes crassicoronata is described and illustrated as a new species from southern Vietnam. It somewhat resembles P. macrostegia in having drooping bowl-shaped purplish flowers, but is readily distinguishable by the larger leaves with more numerous longitudinal veins, internally thickly ribbed staminal corona, and pistil with a hexagonal ovary...

Based on morphological data, Paraboea villosa from limestone areas of northern Vietnam is described and illustrated as a new species. It is similar to Paraboea middletonii Triboun, P. rufescens (Franch.) B.L. Burtt and P. umbellata (Drake) B.L. Burtt, but differs in its almost ebracteate inflorescence, small white flowers, very short peduncles, abs...

This paper continues the publication of new data obtained during field studies in remote and poorly studied regions of Laos in 2015–2018. It provides descriptions of two new species (Bulbophyllum brachyscapum Aver. and Sunipia saccata Aver.), emended descriptions of two previously published but insuffi ciently known species (Cleisostoma verrucosum...

Spiradiclis hainanensis H.S. Lo and S. purpureocaerulea H.S. Lo (Rubiaceae) are reported as new additions to the flora of Vietnam. In Vietnam, the genus Spiradiclis has been represented by only two species, S. leptobotrya (Drake) Pit. and S. caespitosa Blume. During our field investigations in the Lao Cai and Ninh Binh Provinces of northern Vietnam...

Aspidistra babensis K.S. Nguyen, Aver. & Tillich, a new species of Asparagaceae from limestone mountains of Bac Kan Province in northern Vietnam, is described and illustrated. Morphologically, it is close to A. stricta, sharing a perigone that is upright and split almost to base, and with fleshy perigone lobes. Aspidistra babensis however differs f...

Two new species in Begonia sect. Coelocentrum, Begonia acutis, and B. remusatifolia, have been discovered in the karst regions of northern Vietnam, one of the richest habitats for Begonia. The differences of the two new species were compared with other peltate-leaved species in China and Vietnam, including B. cavaleriei, B. pulvinifera and B. wangi...

Begonia comprises at least 1850 species, most of which are local endemics. The greatest genus diversity is observed in tropical zone of America and Asia with particular rich area in Indochinese Peninsula. Meanwhile, the inventory of the genus in this region remains far from acceptable completing. Until recently, only 20 species were documented in L...