Juan A. Zaragoza

• Bachelor of Science
• Honorific Collaborator at University of Alicante

Globochthonius satapliaensis (Schawaller & Dashdamirov, 1988) (n. comb.) original-ly described as Chthonius (Chthonius) satapliaensis from a single female collected in the Sataplia II Cave, Imereti Region (Georgia) is redescribed and depicted based on specimens of both sexes from caves of Sataphlia-Tskaltubo karst massif and Zemo Imereti Plateau. I...

Faunistic sampling in the western Caucasus (Russian side) has yielded two new species belonging to the subgenus Neobisium (Neobisium) Chamberlin, 1930 which are described and illustrated here: Neobisium (N.) catherineae n. sp. and Neobisium (N.) kamenskyi n. sp. The species Neobisium (N.) speleophilum Krumpál, 1986 is redescribed and illustrated ba...

Faunistic sampling in the western Caucasus (Russian side) has yielded two new species belonging to the subgenus Neobisium (Neobisium) Chamberlin, 1930 which are described and illustrated here: Neobisium (N.) catherineae n. sp. and Neobisium (N.) kamenskyi n. sp. The species Neobisium (N.) speleophilum Krumpál, 1986 is redescribed and illustrated ba...

The ecological functioning of dryland ecosystems is closely related to the spatial pattern of the vegetation, which is typically structured in patches. Ground arthropods mediate key soil functions and ecological processes, yet little is known about the influence of dryland vegetation pattern on their abundance and diversity. Here, we investigate ho...

A review of the pseudoscorpion fauna from the Limatambo and Yanaoca districts, Cusco department, Peru, is presented. The species Parachernes (Parachernes) peruanus Beier, 1955 and Stenolpiodes gracillimus Beier, 1959, are reported for the first time for the Cusco department and a new species of Neochelanops, N. unsaac, is described.

A review of the pseudoscorpion fauna from the Limatambo and Yanaoca districts, Cusco department, Peru, is presented. The species Parachernes (Parachernes) peruanus Beier, 1955 and Stenolpiodes gracillimus Beier, 1959, are reported for the first time for the Cusco department and a new species of Neochelanops, N. unsaac, is described.

A case of phoresy by a pseudoscorpion on a longhorn beetle is reported from Italy. It represents the first record involving a pseudoscorpion and the genus Rosalia Audinet-Serville, 1833. Primer registro de foresia en el género Rosalia (Coleoptera: Cerambycidae) a cargo de un pseudoescorpión (Arachnida: Pseudoscorpiones) Resumen: Se registra un caso...

A new genus and species of epigean pseudoscorpions belonging to the subfamily Neobisiinae Chamberlin, 1930, is described from western Georgia and its diagnostic characters are illustrated. Cornuroncus n. gen.resembles the genus Roncus L. Koch, 1873 in several characters but differs by having a short dorso-distal spine on tarsus IV, and the presence...

Pseudoscorpions inhabiting mature Quercus pyrenaica Willd. trees have been surveyed using hollow emergence traps and window traps in Salamanca, Spain. Three new species are described: Neobisium (N.) hispanicum Zaragoza & Hernández- Corral, n. sp., Rhacochelifer gaeli Zaragoza & Hernández-Corral, n. sp. and Rhacochelifer nonidezi Zaragoza & Hernán-...

A new troglobitic species of the genus Antillobisium is described from Sancti Spiritus province, Cuba. Antillobisium toma-si n. sp. is the third known species of the genus and is well characterized by its large size and extremely slender appendages. Biogeographical and ecological considerations on the genus are given. Resumen Se describe una nueva...

A new troglobitic species of the genus Antillobisium is described from Sancti Spiritus province, Cuba. Antillobisium tomasi n. sp. is the third known species of the genus and is well characterized by its large size and extremely slender appendages. Biogeographical and ecological considerations on the genus are given.

Five new species of the recently created genus Occidenchthonius Zaragoza, 2017 are described from caves of Portugal: Occidenchthonius alandroalensis sp. nov., O. algharbicus sp. nov., O. duecensis sp. nov., O. goncalvesi sp. nov. and O. vachoni sp. nov. The species Occidenchthonius cardosoi (Zaragoza, 2012) and Chthonius ischnocheles (Hermann, 1804...

Se presenta una asociación forética entre Pselaphochernes scorpioides (Hermann, 1804) y Achyrolimonia decemmaculata (Loew, 1873), siendo ésta la primera cita de foresia de un pseudoescorpión sobre un limónido en la Península Ibérica. Asimismo, la cita de P. scorpioides es novedosa para Cataluña (provincia de Barcelona). New record of a phoretic a...

First world phoretic record by Pseudoscorpiones on Diptera, Rhinophoridae Abstract: New phoretic record between pseudoscorpions and dipterans from Órgiva, Granada province, Spain. It represents the first world record for a phoretic pseudoscorpion, Pselaphochernes lacertosus, on family Rhinophoridae. A table compiles the records of phoretic associat...

Several phoretic records by pseudoscorpions on Hymenoptera are documented, all them observed in Órgiva, Granada province, Spain. Phoresy by pseudoscorpions on Evaniidae is the first record for Europe and other four records on Ichneumonidae are also mentioned. In all cases, the phoront was assigned to Pselaphochernes lacertosus (L. Koch, 1873).

El presente listado muestra la representación del orden Pseudoscorpiones en la península ibérica e islas Baleares actualizado a fecha 31 de julio de 2017. Los táxones se presentan siguiendo una estructura jerárquica y por orden alfabético dentro de cada categoría. Cuando en una especie hayan sido descritas subespecies, se indicará aquella (o aquell...

Se reportan cuatro especies de pseudoescorpiones de los bosques de manglar del sur del Golfo de Morrosquillo, Caribe Colombiano. Se reportan por primera vez para Colombia y se amplía la distribución conocida de las especies Americhernes reductus, Epactiochernes insularum, Parachernes (Parachernes) aff. setosus y Serianus gratus. Se presenta una cla...

During a spider inventory project in the Ciudad Real and Toledo provinces, Castilla-La Mancha region, Spain, some pseudoscorpion samples were also collected. Four pseudoscorpion species have been identified: Atemnus politus (Simon, 1878), Pselaphochernes lacertosus (L. Koch, 1873), Olpium pallipes pallipes (Lucas, 1849) and Withius hispanus (L. Koc...

The pseudoscorpions Allochernes deceuninckorum Henderickx & Vets, 2003 and Paraliochthonius singularis (Menozzi, 1924) are recorded from Portugal for the first time. The myrmecophilous A. deceuninckorum was found among layered stones and rubble south of Lisbon in the Arrábida Mountains. The intertidal P. singularis was found in coastal habitat unde...

Resumen: Se ofrecen los datos obtenidos en el III Aracnoblitz-GIA, desarrollado durante las XV Jornadas GIA en marzo de 2015. Se han analizado todos los grupos de Arachnida (Scorpiones, Pseudoscorpiones, Opiliones, Acari y Araneae), de modo que el estudio ofrece una primera imagen aracnológica conjunta de algunas zonas del Parque Natural de Cabo de...

A taxonomic revision of the subgenus Chthonius (Ephippiochthonius) in the Iberian Peninsula, Balearic Islands and Macaronesia is presented. New and previously insufficiently studied characteristics are shown to be taxonomically useful, such as the chelal lyrifissures patterns, chaetotaxy and condylar complex. Three taxa previously treated as subgen...

Interesting information (new scientific records or rarities) on 37 species of hypogean invertebrates found in subterranean habitats from Sierra de Segura (Jaén) is presented. Ten of them are nematodes and 27 arthropods.

El orden pseudoescorpiones son artrópodos que pertenecen a la Clase Arachnida. Se caracterizan por poseer pedipalpos quelados, con glándulas venenosas en los extremos de los dedos de la quela. Otros caracteres importantes son el cuerpo dividido en dos tagmas, prosoma y opistosoma, unidos sin estrechamiento intermedio, la ausencia de telson o cola y...

Abstract: Intensive sampling over a period of a year has been carried out in Coves de Campanet, Mallorca, Balearic Islands, producing three pseudoscorpion species. Two of these are epigean species Chthonius (Chthonius) ischnocheles and Chthonius (Ephippiochthonius) tetrachelatus. The third is a previously undescribed species, Chthonius (Chthonius)...

Four new hypogean species are described from the Jaén province (southern Spain): Chthonius (Ephippiochthonius) espanyoli sp. nov., C. (E.) giennensis sp. nov., C. (E.) villacarrillo sp. nov. and Neobisium (Ommatoblothrus) perezruizi sp. nov. New records are given for the species Chthonius (E.) cazorlensis, C. (E.) perezi, C. (E.) tetrachelatus, Neo...

Three new hypogean species of the Iberian genus Roncocreagris Mahnert, 1974 are described from mainland Portugal: R. borgesi sp. nov. and R. gepesi sp. nov. from caves in the Sicó massif, and R. occidentalis sp. nov. from caves in the Montejunto and Cesaredas karst plateau. This brings to nine the number of known hypogean species of the mostly Iber...

The new pseudoscorpion genus Lusoblothrus of the family Syarinidae is described from a cave in the Algarve region, southern Portugal, to accommodate L. aenigmaticus sp. nov., whose morphological affinities within the Holarctic syarinid fauna are not clear and resembles the Gondwanan genera. This discovery emphasizes the relevance of the Algarve reg...

Abstract: The new species Chthonius (Ephippiochthonius) cardosoi is described from a hypogean location. This is the sixth species of the subgenus recorded from mainland Portugal. Keywords. Pseudoscorpiones, Chthoniidae, Chthonius, Ephippiochthonius, hypogean, Portugal. Taxonomy. Chthonius (Ephippiochthonius) cardosoi sp. nov.

El primer Encuentro Ibérico de Biología Subterránea, celebrado en Valencia el pasado julio de 2009, logró reunir un nutrido elenco de investigadores y profesionales en diversos aspectos de la Biología Subterránea que trabajan en España y Portugal. Entre los objetivos del citado encuentro se pretendía publicar los resúmenes de las ponencias que abor...

http://www.sea-entomologia.org/gia/jornadas/jornadas_gia_xii.pdf

The pseudoscorpion Neobisium (Blothrus) birsteini Lapschoff is recorded for the first time from the famous Krubera-Voronya cave, Abhazia. Short description of the unique specimen found is provided and morphological variability discussed.

Female of Chthonius (Ephippiochthonius) ventalloi cazorlensis is reported and described for the first time from type locality, Cueva Secreta del Sagreo. Differences between C. (E.) ventalloi subspecies are discussed and Chthonius (Ephippiochthonius) cazorlensis stat. nov. is elevated to specific rank.

The new genus Titanobochica is described for Titanobochica magna sp. nov. from caves of the Algarve karstic massif, in Portugal. The new genus is assigned to the family Bochicidae and its particular characteristics and geographical isolation suggest a relictual condition. A key to the genera of Bochicidae is provided. The composition of the cave-dw...

Arcanobisium gen. nov. is described for Arcanobisium comasi sp. nov., from a limestone cave in the Valencian Community, Castellón province, Spain. By its unique features, compared to other syarinids, it warrants the creation of a new subfamily, Arcanobisiinae, for its accommodation. Some characters of the Syarinidae are evaluated and the genera Chi...

Additional descriptive data are provided for Mesochelifer fradei Vachon, 1940, based on type material from Portugal and additional samples from Spain (provinces of Cádiz and Huelva) and Morocco. Mesochelifer insignis Callaini, 1986 is shown to be a junior subjective synonym of M. fradei, based on an examination of paratype specimens from Algeria.

Roncus elbulli sp. n. is described from Cap de Creus Nature Park (Catalonia, Spain). Th e new species is morphologically close to Roncus cadinensis Zaragoza, 2007 (Barcelona province), but diff ers by being smaller in size and having a more robust chela with a diff erent microsetae pattern. Th e new species seems to be restricted to coastal semiari...

Abstract: The subspecies of Roncocreagris galeonuda Beier have been examined and new characteristics for their separation are considered. The chelal microsetae pattern and the shape of coxa I are shown to provide good diagnostic characters for these taxa, which are here elevated to specific rank. Remarks on the genus Roncocreagris and its relations...

Roncus montsenyensis sp. nov. is described from Montseny Natural Park (Catalonia, Spain). The new species is geographically and morphologically close to Roncus cadinensis Zaragoza, 2007, but can be separated from it by palpal morphometrics, the chelal microsetae pattern and karyology. The diploid number was found to be 2n=16 in R. montsenyensis, wi...

Roncus montsenyensis sp. nov. is described from Montseny Natural Park (Catalonia, Spain). The new species is geographically and morphologically close to Roncus cadinensis Zaragoza, 2007, but can be separated from it by palpal morphometrics, the chelal microsetae pattern and karyology. The diploid number was found to be 2n=16 in R. montsenyensis, wi...

Mesochelifer fradei Vachon, 1940, is recorded for the first time from Spain, provinces of Cádiz and Huelva. Up to now unknown habitat for the species is described and its phoretic association with saproxylic cerambycid beetles is studied.

We present new data on cave pseudoscorpions from Gipuzkoa and neighbouring regions of the Basque Country and Northern Navarra (Spain). The discovery of new troglobiont species, new localities of collection and distribution area extensions of known species are pointed. A total of 30 taxa, 22 of them found in caves and 17 troblobites or strict cave-d...

Abstract: We report ecologic, faunistic and taxonomic results from a one-year, extensive sampling of pseudoscorpions at the Cadí-Moixeró Natural Park. A total of 86 specimens belonging to 8 different species were collected. We describe the new species Roncus cadinensis sp. nov.; Neobisium geronense Beier stat. nov.is elevated to species and Neobisi...

Abstract: All the available taxonomic literature on the arachnid order Pseudoscorpiones present in the Iberian Peninsula and the Balearic Islands up to the year 2005 has been examined, from which has been produced a list of 178 taxa in 38 genera. Fourteen families, are represented, of which the Neobisiidae and Chthoniidae are the most diverse and p...

We present the results of sampling cave terrestrial fauna in several Majorcan caves: Cova del Pirata, Cova des Pont, Cova de sa Piqueta and Cova des Xots, all of them from the Can Frasquet zone, in the municipality district of Manacor. The knowledge on the biological communities of these caves has been increased with new records for the island of M...

Pseudoscorpiones.

Mundochthonius gallaecicus sp. nov. Mundochthonius gallaecicus new species from Galicia, mainland Spain, is described. This is the first record of the genus Mundochthonius Chamberlin from Western Europe and suggests that the genus may be widespread across the entire continent.

Abstract: Roncus judsoni sp.n., a new hypogean Roncus from Catalonia, Spain, is described. Roncus drescoi Heurtault, 1986 is proposed to be a junior subjective synonym of Roncus duboscqi Vachon, 1937 and the importance of the group of microsetae close to trichobothria eb/esb as a specific taxonomic character is emphasized. Key words: Pseudoscorpion...

Larca lucentina sp. n. and Larca fortunata sp. n. are described from the south-eastern Spanish provinces of Alicante and Murcia. These cave-dwelling species are compared with the other species of the genus, particularly the European taxa. The hypothetical descent of cavernicolous European species from populations of a common ancestor similar to the...

Abstract: Two species of pseudoscorpions have been collected during a prospecting year in caves in the area of Machico town, Madeira island. The new taxon Paraliochthonius cavalensis n. sp. is described, which represents the first troglobitic pseudoscorpion for the Madeira archipelago. Relationship of the pseudoscorpion fauna from Madeira with the...

Abstract: Epigean species of the genus Acanthocreagris Mahnert are reported for the first time for continental Spain. Up to now only three species and three subespecies, all of them cave dwellers and relictic, were known from continental Spain, along with one epigean species from the Balearic Islands. Two new species are described: Acanthocreagris...

Abstract A new species of cave-dwelling pseudoscorpion of the genus Roncocreagris Mahnert is described from Asturias (northern Spain). This discovery represents an interesting link between the troglobitic species from Portugal and Galicia on the one hand and Roncocreagris aurouxi Zaragoza from Cantabria on the other hand. This shows that cave-dwell...

Abstract Two new cave-dwelling pseudoscorpion species of the genus Roncocreagris from Galicia, NW-Spain, are described.They constitute a geographical link with the other three troglobitic species of the genus from the Iberian Peninsula, two from Portugal and one from Cantabria (northern Spain). Comparisons between the five species are made. Key Wor...

A new species of the genus Minniza is described which represents the first record for the Iberian Peninsula of this genus. The Minniza algerica Beier, 1931, type is redescribed and its similarities with M. vermis E. Simon, 1881, are discussed. One specimen of M. graeca (L. Koch, 1873) from the greek island of Santorini and others from Maroc are stu...

En el presente trabajo se estudian algunas especies cavernícolas de pseudoscorpiones de Asturias, Cantabria y el País Vasco, en el Norte de España. Se ofrecen nuevas citas para Neobisium (Blothrus) jeanneli (Ellingsen) y N. (B). nonidezi (Bolivar); ésta última y N. (B.) vasconicum vasconicum (Nonidez) son redescritos sobre ejemplares recogidos en s...

Se cita la presencia de dos especies de pseudoscorpiones (Olpium pallipes e Hysterochelifer sp.) en los contenidos estomacales y heces de Acanthodactylus erythrurus y Psammodromus algirus en dunas de la provincia de Alicante. Se confirma la relativa abundancia de la primera de ellas sobre ramas y hojas de Thymelaea hirsuta, mediante mangueos mensua...

Main Systematic Groups.. In the group classitematic groups are known three suborder, of them nine family groups are kmown from Europe, all them with representant in the Iberian Peninsula.

La propuesta formulada más abajo para una nueva especie del subgénero Ephippiochthonius, viene a aumentar la lista de endemismos propios del Sureste Español. Nuevamente se pone en evidencia que, la búsqueda sistemática en esta Zona y tanto en biotopos hipogeos o epigeos, puede proporcionar un número aún considerable de nuevas especies. A new cave-d...

A new cave-dwelling specie of Chthonius (Pseudoscorpions, Chthoniidae) from Beniarbeig, Alicante (Spain), is described and figured: Chthonius (E.) mahnerti n. sp.

A new species of Pseudoscorpion belonging to the family Neobisiidae is described: Roncus (Parablothrus) setosus. Easily distinguishable of any other species of the genus by the extraordinary number of setae on the carapace and tergites, representing another case of «neochétotaxie majorante prosomatique» J. HEURTAULT 1980, this being confirmed by th...

El presente artículo es una recopilación de la bibliografía existente hasta la fecha en relación a los pseudoscorpiones de la península Ibérica, Baleares y Macaronesia. Se incluyen artículos con citas de localizaciones, descripciones de taxones endémicos, ecología, biodiversidad y catálogos faunísticos.