José Manuel Gasca

University of Salamanca · Área de Estratigrafía

This work characterizes for the first time the 500–700 m-thick uppermost Cretaceous continental sedimentary succession of the Allueva Fm recorded in the northeastern margin of the Iberian basin, in the Montalbán subbasin (Maestrazgo domain, Spain). The middle-upper Campanian age of this unit constrained here by new paleomagnetic and biostratigraphi...

An enlarged theropod manual ungual (CSC1-4) from the Weald facies of Spain is described. The claw was found in the fossil locality of Caña Seca 1, Teruel province, within the El Castellar Formation of early Barremian (Early Cretaceous) in age. CSC1-4 is morphologically closer to megalosauroids than to any other theropod clade bearing enlarged manu...

Surface collecting is the first and sometimes the only type of sampling carried out in many fossiliferous localities, including vertebrate microfossil assemblages. Nevertheless, it is rare to test how representative these surface-collected samples are of the palaeobiocoenosis. A first approach to the palaeontological analysis of Los Menires, a Barr...

Se ha realizado la caracterización bioestratinómica del yacimiento “Pepe”, situado próximo a Ladruñán (provincia de Teruel) en rocas de la parte superior de la Formación Mirambel, de edad Barremiense. El yacimiento analizado se sitúa en facies continentales que corresponden con un sistema aluvial-lacustre dominado por lutitas de llanura aluvial afe...

A partir de la descripción de dientes aislados de crocodilomorfos del yacimiento Peñuquillo, se han definido 7 morfotipos, atribuibles a tres gru pos, Goniopholididae, Bernissartiidae y Atoposauridae. Esta asociación es común en muchos yacimientos mesozoicos de la península ibérica, pero hasta la fecha no se habían registrado conjuntamente en es...

A paleohistological comparison has been made between the dorsal ribs of two iguanodontid ornithopod specimens from the Azud Aliaga (Aliaga) and Pepe (Ladruñán) sites, both from the Barremian (Lower Cretaceous) of the Maestrazgo Basin (Teruel, Spain). Both specimens share a similar histo-logical development in early stages of ontogeny, while in more...

Los dinosaurios ornitópodos, y en particular los iguanodontios, fueron componentes esenciales de los ecosistemas continentales barremienses (Cretácico Inferior) en el sur de Europa si consideramos la frecuencia con la que aparecen registrados en asociaciones fósiles de esa edad. La Formación Mirambel, en la subcuenca de Morella (cuenca del Maestraz...

Se ha realizado el estudio histológico de dos costillas de un ornitópodo estiracosterno iguanodóntido del yacimiento Pepe de la Formación Mirambel (Barremiense, Cretácico Inferior), ubicado en el anticlinal de Ladruñán (subcuenca de Morella, Cuenca del Maestrazgo) en el noreste de la provincia de Teruel. Los elementos postcraneales recuperados (un...

Decapod crustaceans are rare in Early Cretaceous invertebrate assemblages from the Iberian Peninsula; however, current knowledge does allow differences in taxonomic composition between different basins and subbasins to be recognised. In order to understand the distribution of Barremian–Aptian decapod crustaceans of Iberia better, seven new localiti...

The Upper Cretaceous (middle Campanian–earliest Maastrichtian) Allueva Fm. is a fossil-rich continental unit deposited in the early stages of the Montalbán and Aliaga intramountain sedimentary basins formation (localities of Bea, Allueva and Segura de Baños, Iberian Range, Teruel province, Spain). The Allueva Fm., deposited in a middle-distal alluv...

Several remains of iguanodontid ornithopods have been reported from two fossil sites (Camino de La Algecira and Pepe) from the Mirambel Formation (Barremian, Lower Cretaceous). The sites are located in the Ladruñán anticline (Maestrazgo basin, Teruel province, Spain). The sampled bones are a neural spine of a dorsal vertebra, four fragments of vert...

New material of an iguanodontid styracosternan ornithopod from the Lower Cretaceous of the Iberian Peninsula is described. The cranial and postcranial skeletal remains are from the Barranco del Hocino-1 site. These fossils (Iguanodontidae sensu Xu et al., 2018) represent the first medium-sized styracosternan from the Upper Sequence of the Blesa For...

Dinosaur tracks have been identified in several Upper Jurassic-Lower Cretaceous sedimentary units across the Maestrazgo Basin in eastern Iberia, which are preserved in a variety of transitional and continental paleoenvironments. Here, we described the lower Barremian San Benón tracksite with the first occurrence of clear dinosaur tracks in the El C...

The reconstruction of the latest Jurassic–Early Cretaceous evolution of the Torrelapaja Subbasin (Cameros Basin, Spain) resulted in the characterization of three synrift sequences (SS-1, SS-2 and SS-3) bounded by major unconformities. Three major NW-SE normal faults combined with smaller scale faults of variable direction (around NE-SW) controlled...

Throughout the evolutionary history of Avialae, several members of this clade have evolved into giant forms, in different time periods and ecological contexts. In Europe, the first birds that show this condition, the Gargantuaviidae, occur during the Late Cretaceous (late Campanian–early Maastrichtian), but it is during the Paleogene when more grou...

The South-Pyrenean Basin (northeastern Spain) has yielded a rich and diverse record of Upper Cretaceous (uppermost Campanian−uppermost Maastrichtian) vertebrate fossils, including the remains of some of the last European dinosaurs prior to the Cretaceous-Paleogene (K-Pg) extinction event. In this work, we update and characterize the vertebrate foss...

Sequence-stratigraphic, biostratigraphic and strontium-isotopic data have made it possible to update what is known of the stratigraphy and sedimentary evolution of the Kimmeridgian–Berriasian synrift successions recorded in four subbasins (Aguilón, Oliete, Morella, Galve) of the central Iberian Rift System (NE Spain). The studied successions are ar...

Ornithopods are traditionally a poorly known clade of dinosaurs within the Upper Cretaceous fossil record of Argentina. In the last few years, the number of discoveries has increased significantly in the out-crops near Rincón de los Sauces village (Neuquén province), where remains have appeared in two different formations (Plottier and Bajo de la C...

Detailed logging and analysis of the facies architecture of the upper Tithonian to middle Berriasian Aguilar del Alfambra Formation (Galve sub-basin, north-east Spain) have made it possible to characterize a wide variety of clastic, mixed clastic–carbonate and carbonate facies, which were deposited in coastal mudflats to shallow subtidal areas of a...

A review of the onset of the synrift sedimentation and synsedimentary extensional tectonics of the Oliete sub-basin (north- western Maestrazgo basin, East Spain) is presented here based on new data acquired after extensive sedimentological, struc- tural and palaeontological analysis of the Barremian Blesa Fm. The lower boundary of the Blesa Fm is a...

Introduction: Barranco del Hocino-1 is a new fossil site located near Estercuel, Teruel province, Spain. The fossil site is located geologically within the Oliete sub-basin, in the Blesa Formation (Barremian in age). Barranco del Hocino-1 shows a diverse assemblage of tetrapod vertebrates similar to other sites in the Blesa Formation. Materials and...

Aragosaurus-IUCA (University of Zaragoza) is a research group in vertebrate paleontology, originally focused in two lines of work: Mesozoic vertebrates from the Iberian Peninsula and Patagonia, and Pleistocene vertebrates from Spain (especially the small vertebrates of the Atapuerca locality). Lately, the group has developed a notable labor prospec...

We describe a new species of an ornithischian dinosaur, Isaberrysaura mollensis gen. et sp. nov. The specimen, consisting in an almost complete skull and incomplete postcranium was collected from the marine-deltaic deposits of the Los Molles Formation (Toarcian-Bajocian), being the first reported dinosaur for this unit, one of the oldest from Neuqu...

Full text here: http://sepaleontologia.es/wp-content/uploads/2017/02/5-Moreno-Azanza-et-al-ok.pdf. Here we describe the new fossil site of El Horcajo in the vicinity of Trevijano (La Rioja, Spain). It is located in the palustrine facies of the Enciso Group (Cameros Basin). This new locality has provided dozens of eggshell fragments, together with...

Up to now, the ichnological vertebrate record from the Barremian Mirambel Formation (NE Spain) has remained completely unknown despite the fact that osteological findings have been reported in recent years. Here we provide an overview of 11 new dinosaur tracksites found during a fieldwork campaign in the year 2011. The majority of these tracksites...

Ornithischian dinosaurs were an important component of the Mesozoic fauna. Their bones and tracks are commonly found in Jurassic and Cretaceous outcrops, and more rarely in the Upper Triassic. They are particularly ubiquitous in certain temporal ranges and geographical areas, such as the Early Cretaceous of Europe and the Late Cretaceous of North A...

Las facies continentales Weald de Inglaterra y de España presentan algunas de las asociaciones de dinosaurios barremianos más diversas de todo el Cretácico Inferior de Europa. En la Cuenca Ibérica Mesozoica (en el este de España) aflora la Formación Mirambel, una unidad compuesta por sedimentos aluviales y lacustres que ha proporcionado un importan...

Abnormalities in the histo- and ultrastructure of the amniote eggshell are often related to diverse factors, such as ambient stress during egg formation, pathologies altering the physiology of the egg-laying females, or evolutionarily selected modifications of the eggshell structure that vary the physical properties of the egg, for example increasi...

Raw EBSD data of PS-TEC,215. CPR and CRC Channel 5 files containing the acquired EBSD data, after the combination of the 27 acquired maps. (ZIP)

Here we present material from the new vertebrate fossil locality of Barranco del Hocino 1, located within the municipality of Estercuel (Teruel, Spain). Outcrops are located in the southwestern area of the Oliete Subbasin, within the Cretaceous Maestrazgo Basin. Geologically, this fossil locality is placed in the middle part of the Blesa Formation,...

Here we present material from the new vertebrate fossil locality of Barranco del Hocino 1, loca- ted within the municipality of Estercuel (Teruel, Spain). One campaign of eldwork on August 2015 has enabled us to recover roughly one hundred bone remains and to identify a moderately diverse assemblage of vertebrates. The fossil association is dominat...

El Barremiense inferior es un intervalo del Cretácico Inferior con escaso registro de dinosaurios en Europa, salvo excepciones como el Weald en Inglaterra y la Cordillera Ibérica en España. En este intervalo se ha identificado la mayor diversidad del registro fósil de vertebrados en general y dinosaurios en particular dentro de los yacimientos del...

Abstract Crocodiloid eggshell is considered to be one of the most conservative among amniotes. This contrasts with the high body diversity observed within the crocodylomorph lineage, which extebds from the Triassic to the present. This incongruence raises a fundamental question in palaeology: is the crocodylomorph eggshell structure that conservati...

Crocodyloid eggshell is considered to be the most conservative amongst amniotes. Known since the Late Jurassic, all fossil eggshells attributed to crocodylomorphs have been included in the oofamily Krokolithidae. This contrasts with the high diversity observed within the crocodylomorph lineage during the Cretaceous. This incongruence raises a funda...

Triassic vertebrate tracks are known from the beginning of the 19th century and have a worldwide distribution. Several Triassic track ichnoassemblages and ichnotaxa have a restricted stratigraphic range and are useful in biochronology and biostratigraphy. The record of Triassic tracks in the Iberian Peninsula has gone almost unnoticed although more...

Triassic vertebrate tracks are known from the beginning of the 19th century and have a worldwide distribution. Several Triassic track ichnoassemblages and ichnotaxa have a restricted stratigraphic range and are useful in biochronology and biostratigraphy. The record of Triassic tracks in the Iberian Peninsula has gone almost unnoticed although more...

Triassic vertebrate tracks are known from the beginning of the 19th century and have a worldwide distribution. Several Triassic track ichnoassemblages and ichnotaxa have a restricted stratigraphic range and are useful in biochronology and biostratigraphy. The record of Triassic tracks in the Iberian Peninsula has gone almost unnoticed although more...

Delapparentia turolensis Ruiz-Omeñaca, 2011 is the only iguanodont taxon erected in the Barremian of Spain. It is described on the basis of a partial postcranial skeleton discovered in the 1950s near the village of Galve (Teruel Province), within the Camarillas Formation. Recently, new remains from the same individual have been recovered, and these...

In this work we describe fossils of iguanodontian ornithopod dinosaurs from five new fossil localities of the Mirambel Formation (Barremian, Lower Cretaceous) in the municipalities of Castellote and Tronchón (Teruel, Spain). The studied fossils include an anterior fragment of a left maxilla bearing functional teeth, remains of three caudal vertebra...

In this work we describe fossils of iguanodontian ornithopod dinosaurs from five new fossil localities of the Mirambel Formation (Barremian, Lower Cretaceous) in the municipalities of Castellote and Tronchón (Teruel, Spain). The studied fossils include an anterior fragment of a left maxilla bearing functional teeth, remains of three caudal vertebra...

The fossil record of dinosaurs from the Mirambel Formation (Barremian, Teruel province, Spain) is relatively scarce, being cited the presence of iguan-odontian ornithopods and basal tetanuran theropods until now. In this paper, we document the presence of sauropods in this unit for the first time. They are fragmentary postcranial remains found in d...

The fossil record of dinosaurs from the Mirambel Formation (Barremian, Teruel province, Spain) is relatively scarce, being cited the presence of iguan-odontian ornithopods and basal tetanuran theropods until now. In this paper, we document the presence of sauropods in this unit for the first time. They are fragmentary postcranial remains found in d...

Trigonoolithus amoae oogen. et oosp. nov. is described on the basis of abundant eggshell fragments from the La Cantalera site in the Late Cretaceous (early Barremian) Blesa Formation, Teruel Province, northern Spain. The surface ornamentation, the most diagnostic feature of the new oogenus, consists of closely spaced sub-triangular or rounded protu...

Iguanodont fossils

Spalacotheriids, basal trechnotheres with an acutely angled, reversed-triangle pattern of major molar cusps, are relatively rare pan-Laurasian mammals that ranged through much of the Cretaceous but are generally not well represented in the fossil record. Herein we describe newly collected specimens from the Molino Alto 1 locality, Teruel Province,...

Altered iguanodontian dinosaur teeth have been studied to analyse the unusual replacement of biogenic apatite and to deduce how the alterations affect the preservation of these fossils. The fossils were recovered from alluvial lacustrine facies (Barremian, Early Cretaceous) in Teruel (Spain). Mandibular teeth and sediments were studied in hand samp...

We describe an isolated distal femur of a large-sized theropod dinosaur from the Mirambel Formation, Teruel province, Spain. It comes from the Ladruñán-3 fossil locality, which represents a sandstone channel deposit that is early Barremian (Early Cretaceous) in age. This femur can be identified as belonging to an undetermined stem carcharodontosaur...

Th e Spalacotheriidae symmetrodonts are Cretaceous mammals having molars with two roots and crowns with three principal cusps on upper and lower molars. Th e cusps are arranged in a nearly symmetrical triangle pattern in the trigon/trigonids and lacking talon/talonids, being the last a character shared with modern mammals. Symmetrodontans appear to...