Jacek Szwedo

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• Ph.D., DSc. habil.
• Professor at University of Gdańsk

The planthopper genus Montrouzierana Signoret, 1861 (Hemiptera: Fulgoromorpha: Tropiduchidae) and its only known endemic species to New Caledonia, M. oxycephala (Montrouzier, 1861), are redescribed. Illustrations of diagnostic characters including male and female genitalia are provided. The systematic position of the genus is briefly discussed. Mor...

The new genus and species Lutevanaphis permiana gen.n. et sp.n., represents the oldest Aphidomorpha, new superfamily Lutevanaphidoidea superfam.n. and new family Lutevanaphididae. This taxon is described from the Middle Permian of the Lodève Basin, southern France. The presence of very small Aphidomorpha in the Middle Permian contradicts the hypoth...

Following recent advances in the morphological interpretations of the tegmen basal cell margins in the Paraneoptera, a standardized and homology-driven groundplan terminology for tegmina types, structures and vein patterns in Hemiptera Fulgoromorpha, including fossils, is proposed. Each term is listed with a morphological definition, compared and l...

Paleogene arthropod biotas have proved important for tracing the faunal turnover and intercontinental faunal interchange driven by climatic warming and geodynamic events [1-5]. Despite the large number of Paleogene fossil arthropods in Europe and North America [5-8], little is known about the typical Asian (Laurasia-originated) arthropod biota. Her...

With the exception of a fly and a mite from the Triassic of Italy, all Mesozoic amber arthropods are from the Cretaceous. Late Jurassic Lebanese amber from Aintourine revealed a completely preserved adult coccid male (wing length 0.8 mm), Jankotejacoccus libanogloria gen. et sp. n., the earliest record of a plant sucking scale insect. Associated pl...

The new species Amecephala micra sp. nov. is described and illustrated on the basis of a well-preserved male psyllid (Liadopsyllidae) in a piece of Cretaceous Myanmar amber. This second species of the genus Amecephala Drohojowska, Szwedo, Müller et Burckhardt, 2020 exhibits a combination of features that have not been previously recognised among Li...

A new genus and species of Aleurodicinae whiteflies from the Eocene Baltic amber are described. Eogroehnia carsteni Drohojowska and Szwedo, nov. gen., nov. sp., is the third representative of Aleurodicinae from Baltic amber. Morphological features of this new whitefly are discussed in the context of features of other contemporaneous aleurodicine wh...

Planthoppers (Hemiptera: Fulgoromorpha) are a species‐rich and globally distributed insect clade with high economic, ecological and evolutionary importance. However, the relationships among planthopper lineages and families remain unclear. Previous efforts based on inconsistent morphological traits, a few genes or limited sampling often resulted in...

An elytra of a beetle was discovered from sediments of the Mandla Lobe Deccan Intertrappean beds of Umariya in Madhya Pradesh, India dating from the latest Maastrichtian (Late Cretaceous) to earliest Danian (early Paleocene). The elytra was examined with light microscopy and micro-CT and revealed a close resemblance to that of the Curculionoidea. T...

A new genus and species of fossil whiteflies (Aleyrodidae) and a psyllid (Psyllidae) were discovered in the Hindon Maar Complex, a Fossil-Lagerstätte within the Dunedin Volcanic Group in Otago, South Island of New Zealand. These represent the first fossil records of these sternorrhynchan families (Hemiptera) from New Zealand. Miotetraleurodes novae...

The whiteflies (Hemiptera: Aleyrodidae) are small sternorrhynchan insects, which have the potential to cause significant economic damage to agricultural crops. There is a paucity of knowledge regarding the diversity, disparity, and evolutionary history of these insects, with classification based on the immatures, called puparia. The fossil record o...

This paper describes a new genus and species of Achilidae (Hemiptera: Fulgoromorpha) Achiplecton stilleri gen. et sp. nov from the newly established tribe Achiplectini trib. nov. This tribe belongs to one of three Achilidae subfamilies, Myconinae, and is found solely in the West Cape of Southern Africa. The whole region is thought to be one the of...

Here we report a new species, Cretodorus lijuanae sp. nov., of extinct Mimarachnidae from mid-Cretaceous amber of northern Myanmar. C. lijuanae sp. nov. was described based on one adult and three nymphal fossils. The new adult specimen exhibits the diagnostic features of Cretodorus and possesses a unique number and arrangement of sensory pits in th...

Of the early Eocene amber deposits known across the world, Belgian amber has been mostly absent from the relevant literature. We reinvestigated amber held in the palaeobotanical collection of the Royal Belgian Institute of Natural Sciences, Brussels, which derived from three localities in Belgium that originated from two geographical areas (Leval-T...

The morphological data available for Jurassic Coleorrhyncha taxa are scarce and scattered across different sources, often vaguely discussing wing and body part morphology. Therefore, it is essential to discover, describe and apply new characters at both micro and macro scales. Progonocimicidae specimens are frequently reported from various Jurassic...

The aim of this study is to analyse the extensive collection of the zoological drawings by Samuel Niedenthal that has been preserved in Dresden. They were executed using various techniques and depict a variety of animal species. This article draws attention especially to studies of the local fauna. This legacy of Pomeranian zoology is unique and im...

This study presents a comprehensive and illustrated catalogue of the forewing venation of the Paleo- and Mesozoic Hemiptera planthoppers. It compiles fragmented information dispersed across various publications, ensuring a uniform interpretation of tegmina venation for fossil taxa. Taxa are presented by family and each species is documented with it...

Sinofulgoridium suni Zhang, Szwedo and Zhang, n. gen. n. sp., is established based on a complete tegmen from the Middle Triassic Tongchuan Formation in Shaanxi, China, and is assigned to the family Surijokocixiidae of Fulgoromorpha, Hemiptera. A key of all the known species of Surijokocixiidae is provided for the first time.

Extant cicada (Hemiptera: Cicadoidea) includes widely distributed Cicadidae and relictual Tettigarctidae, with fossils ascribed to these two groups based on several distinct, minimally varying morphological differences that define their extant counterparts. However, directly assigning Mesozoic fossils to modern taxa may overlook the role of unique...

A new monospecific fossil genus of ricaniid planthoppers (Hemiptera: Fulgoromorpha: Ricaniidae), Worako gen. nov., is described with Worako yulei sp. nov. The fossil is reported from the Late Eocene Insect Bed of the Bembridge Marls, Isle of Wight, UK. It is the second record of Ricaniidae from these strata and third fossil record of Ricaniidae fro...

The second species of the Aleyrodinae whitefly genus Snotra Szwedo et Drohojowska, 2016 is described based on an inclusion in the Eocene Baltic amber. It is second species of the genus, adding new data to knowledge on disparity and taxonomic diversity of whiteflies in the Eocene Baltic amber. The diversity of the group and its fossil record in the...

A new fossil ceratopogonid genus and species from Lower Cretaceous Lebanese amber, Baskintoconops maaloufi Pielowska-Ceranowska gen. et sp. nov., is described and illustrated. The studied material originates from a newly discovered amber site in the Lebanese village Baskinta at a locality dubbed Qanat Bakish. The described genus is typified by its...

Confirmation of occurrence of Issus muscaeformis (von Schrank, 1781) (Hemiptera: Issidae) in Pomerania. Issus muscaeformis (von Schrank, 1781) is rarely collected in Poland mostly in southern parts of the country. In the Gdańsk Pomerania it was recorded only once by Matsumura in 1906 from Żukowo village near Gdańsk. Here we confirm the occurrence...

Prompted by a rapid development in the field of palaeoentomology, fossil terrestrial arthropods and amber research, Palaeoentomology was established in late 2018 through a collaborative effort between the International Palaeoentomological Society (IPS) and Magnolia Press (Azar et al., 2018). The inception of this scientific journal was met with ent...

Chemical defense in insects is becoming an increasingly interesting topic and has the potential for providing unexplored compounds with unknown properties for drug and repellent discovery, so the secretions of different species of insects are being studied and new ways of collecting these secretions are being sought. Silica gel and activated carbon...

A synthesis of planthopper suprafamilial and familial classification is provided and discussed, solving some nomenclatural issues for the group, and adopting an enriched presentation with a clear distinction between the taxon name and its concept.

The second genus and species of the planthopper family Katlasidae, Dumpyawnus hpungwanus gen. et sp. nov., is described from mid-Cretaceous Kachin (Burmese) amber. It can be definitely attributed to Katlasidae mainly based on its tegminal structure (e.g., costal area absent, clavus closed, tegmen widened at membrane, multiplied forked of ScP+RA, MP...

A new genus and species of fossil whitefly, Praealeurolobus indicus gen. et sp. nov., is described and illustrated based on puparia found on leaves of a malvaceous plant from the Pliocene Rajdanda Formation in Eastern India. It is the first record of whitefly puparia on leaves from the Neogene deposits of Asia and the second taxon formally named an...

A new genus Amphignokachinia Brysz & Szwedo, gen. n. and species Amphignokachinia subversa Brysz & Szwedo,sp. n. of Achilidae (Hemiptera) planthoppers of the tribe Amphignomini are described. It is the second genus and first fossil rep-resentative of the tribe Amphignomini coming from the Cenomanian amber in Kachin State, Myanmar. Diagnosis of the...

A new Middle Triassic (Ladinian) planthopper of the family Surijokocixiidae (Surijokocixioidea, Fulgoromorpha) from the Tongchuan Formation in Shaanxi, NW China is established as Sinosurijikocixius tongchuanensis gen. et sp. nov. All the known taxa assigned to Surijokocixiidae are reviewed and compared. The palaeogeographic distributional pattern o...

The basal classification of planthoppers (Hemiptera Fulgoromorpha) is revised. The wrinkled ambient vein (appendix) on the fore wing is given as an apomorphic character for the Fulgoromorpha for which the Pennsylvanian, Moscovian genus Aviorrhyncha (310 Mya), Aviorrhynchidae, is the oldest representative to date. All other planthopper taxa belong t...

A new genus and species of planthopper – Yobuenahuaboshka rectangialatus gen. et sp. nov., preserved as inclusion in late Lower Miocene, Burdigalian amber from Hispaniola (Dominican Republic) is described and illustrated. Classification and fossil records of the family Nogodinidae, and their issues, are briefly overviewed. The morphological charact...

Trichomyia (Septemtrichomyia) grimaldii sp. nov. and Trichomyia (Trichomyia) fudalai sp. nov. from the Lower Miocene Dominican amber are characterised, described, illustrated and their taxonomic position discussed. These fossils constitute the first record of the Trichomyiinae and increase the knowledge on the biodiversity of the biological inclusi...

The previous issue of Palaeoentomology brought the first set of papers honoring David A. Grimaldi on the occasion of his 65th birthday. With the current one, it is continued, in recognition of his impact on the fields of amber studies, palaeontology, palaeo- and neoentomology, and evolutionary biology. After the success of Jurassic Park (both the M...

Libanonemopalpus grimaldii gen. et sp. nov. is characterised, illustrated, and described from the lower Barremian amber of Bqaatouta (Lebanon). It represents the earliest record, the first and only Bruchomyiinae psychodid with functional piercing mouthparts. The discovery of this Cretaceous psychodid fly improves our knowledge of the palaeobiodiver...

Based on new fossil materials, a new species Toxorhina ( Ceratocheilus ) christelius sp. nov. has been described herein with complete documentation of drawings and photographs. Features such as wide spine on the gonocoxite differentiating the new species of Toxorhina were discussed. Finding new interesting fossil materials also allowed for providin...

Laiyangella zhangi gen. et sp. nov., the first planthopper from the Lower Cretaceous Laiyang formation, Shandong Province of China, is described. Although exhibiting a cixioid venation that would seem to indicate its placement with or near the Cixiidae, the well-preserved impression of the genitalia of this female specimen is not of the orthopteroi...

Nymphs of extinct sternorrhynchan hemipterans are extremely rare, although very important for understanding of evolutionary traits of these insects. A protopsyllidioid nymph, in mid-Cretaceous amber from Kachin, Myanmar, placed in the family Postopsyllidiidae, is the first nymph of this family to be found in the fossil. Postopsyllidiidae previously...

Mimarachnidae is a Mesozoic planthopper family, with high palaeodiverisity at generic and species levels during Cretaceous. Herein, geometric morphometric analysis of mimarachinid forewings was executed. Our results of principal component and cluster analyses show that Dachibangus formosus and D. hui are morphologically distinct from the type speci...

A new genus and species of Aleurodicinae whiteflies from the Eocene Baltic amber is described. Medocellodes blackmani Drohojowska et Szwedo gen. et sp. nov. is the second representative of Aleurodicinae from Baltic amber, presenting mixture of plesiomorphic characters, as retention of median ocellus, with apomorphic states, shared also with other c...

Fossils are critical for understanding the evolutionary diversification, turnover, and morphological disparification of extant lineages. While fossils cannot be sequenced, phenome-scale data may be generated using micro-computed tomography ( -CT), thus revealing hidden structures and internal anatomy, when preserved. Here, we adduce the male caste...

The tropiduchid monotypic genus Laberia Stål, 1866, endemic to Madagascar, is placed in a new tribe of its own Laberiini trib. nov. The tribe is placed in the subfamily Elicinae, and can be distinguished from other representatives of the subfamily by the unique combination of morphological characters of the head, tegmina, legs and genital structure...

The superfamily Cercopoidea is commonly named as “spittlebugs”, as its nymphs produce a spittle mass to protect themselves. Cosmoscartini (Cercopoidea: Cercopidae) is a large and brightly colored Old World tropical tribe, including 11 genera. A new genus Nangamostethos gen. nov. (type species: Nangamostethostibetense sp. nov.) of Cosmoscartini is d...

The Early Miocene (16–23 Mya) amber of Ethiopia constitutes a new source of fossil ants for Africa, where they are otherwise poorly documented. Here we report a diversified assemblage of six subfamilies and at least 19 genera that are still predominantly alive in the Afrotropics today. In this first account, a particular reference is made to the su...

A new planthopper genus Tenebricosus gen. nov. from mid-Cretaceous Kachin amber with its type species Tenebricosus coriaceus sp. nov. is described in the family Mimarachnidae. The new genus and species is the fourteenth genus and 22nd species of the extinct family Mimarachnidae, which is the most diverse planthopper family among Kachin amber inclus...

Nymphs of extinct sternorrhynchan hemipterans are extremely scarce, however very important for understanding of evolutionary trait of these insects. A protopsyllidioid nymph, placed in the family Postopsyllidiidae is the first found in the fossil record, based on an inclusion in mid-Cretaceous amber from Kachin, Burma. Postopsyllidiidae so far comp...

The first records of Ceratopogonidae, or biting midges, are from the earliest Cretaceous. The early Cretaceous representatives of the family are currently interpreted as haematophagous. †Minyohelea Borkent, 1995 is one of six extinct, bloodsucking ceratopogonid genera currently recorded (Pielowska-Ceranowska et al., 2021). So far, nine species of t...

Gedanochila museisucini gen. et sp. nov. is described, based on inclusions in the Eocene Baltic amber. A morphological phylogenetic analysis supports the placement of Gedanochila gen. nov. into the tribe Achilini. Definition, content and subdivisions of the tribe as well as position of extinct taxa placed within are briefly discussed.

A new extinct genus and species, Zhangpumazus sheni n. gen. n. sp., of the planthopper family Caliscelidae Amyot and Audinet-Serville, 1843, is described from mid-Miocene Zhangpu amber. This is the second fossil record of this family and the first formally described planthopper species from Zhangpu biota. This new genus and species is apparently cl...

Fossilized tree resin, or amber, commonly contains fossils of animals, plants and microorganisms. These inclusions have generally been interpreted as hollow moulds or mummified remains coated or filled with carbonaceous material. Here, we provide the first report of calcified and silicified insects in amber from the mid-Cretaceous Kachin (Burmese)...

Fossil new genus and species Afroscatopse haennii gen. et sp. nov. from Miocene Ethiopian amber are described and illustrated and placed in the tribe Colobostematini. The gigantic adults of this scavenger fly, largest within the family, with dull body, long antennae and broad wings evidently were adapted to live on bark of trees. It is suggested th...

The second genus and species of recently established planthopper family Inoderbidae, Ingensala xiai gen. et sp. nov., is described based on a well-preserved specimen from mid-Cretaceous Kachin (Burmese) amber, and it can be definitely attributed to Inoderbidae mainly based on its head structure, pronotum and mesonotum without median and lateral car...

This study presents the morphology of calcar in adult Delphacidae based on representatives of the genera Ugyops Guérin-Meneville, 1834, Notuchus Fennah, 1969 (Ugyopini), Asiraca Latreille, 1798 (Asiracini), Kelisia Fieber, 1866, (Kelisini), Stenocranus Fieber, 1866 (Stenocranini), Chloriona Fieber, 1866, Megadelphax Wagner, 1963, Muellerianella Wag...

A new genus and species of the tropiduchid planthopper of the tribe Tropiduchini is described as Bitara augusta gen. et sp. nov. from Madang Province, Papua New Guinea. It is the 18th species of Tropiduchini and the 9th genus known east of the Wallace line. It is another taxon with characteristic asymmetric genitalia within this tribe.

A new, extinct species of Allodia Winnertz is described from early Miocene amber of Ethiopia. Allodia paleoafricana sp. nov. is mostly characterized by the scutum with strong anteromarginal, dorsocentral, and lateral setae and the wing with the stem of the M-fork slightly shorter than the vein r–m and the base of the M4–CuA fork aligned with the ba...

A new species, Dorytocus jiaxiaoae Song, Szwedo & Bourgoin sp. nov., from the mid-Cretaceous Burmese amber is described and illustrated. This species represents the first record of the adult of the extinct family Dorytocidae. The supplementary description for the fourth-instar nymph of Dorytocus ornithorhynchus Emeljanov & Shcherbakov is provided....

Fossilised insects probably brought man’s attention since the prehistory, since first amber with an insect entombed in resin was found. Amber was collected and used by humans first in the Upper Paleolithic period, perhaps as long ago as 20,000 years (Beck et al., 2009; Burdukiewicz, 2009; Płonka & Kowalski, 2017). The written testimonies on amber i...

A new species of the fossil genus Minyohelea (Diptera: Ceratopogonidae) from the lower Barremian (Lower Cretaceous) Lebanese amber is studied. The new species Minyohelea nexuosa sp. nov. is described, illustrated, and its unusual male genitalia is compared with those of the other fossil representatives of this extinct genus.

Ghost lineages have always challenged the understanding of organism evolution. They participate in misinterpretations in phylogenetic, clade dating, biogeographic, and paleoecologic studies. They directly result from fossilization biases and organism biology. The Cylindrotomidae are a perfect example of an unexplained ghost lineage during the Mesoz...

Three new genera and species of Cenomanian Cixiidae from Kachin amber (Myanmar) are described: Acrotiara Bourgoin & Luo gen. nov., type species A. multigranulata Luo & Bourgoin sp. nov., Delphitiara Bourgoin & Luo gen. nov., type species D. tibiocoronata Luo & Bourgoin sp. nov., and Pentacarinus Bourgoin & Luo gen. nov., type species P. kachinensis...

A new family of planthoppers, Szeiiniidae fam. nov., is described on the basis of a well-preserved forewing from the Upper Triassic Yanchang Formation of Huanglong County, Shaanxi Province, China. As the only specimen of the new family, Szeiinia huanglongensis gen. et sp. nov. differs from other families within Fulgoroidea by a unique combination o...

During the Mid-Miocene Climatic Optimum [MMCO, ~14 to 17 million years (Ma) ago], global temperatures were similar to predicted temperatures for the coming century. Limited megathermal paleoclimatic and fossil data are known from this period, despite its potential as an analog for future climate conditions. Here, we report a rich middle Miocene rai...

Here we describe and illustrate a new genus and species of the extinct froghopper family Sinoalidae, Cretomultinervis burmensis gen. et sp. nov. from Burmese (Myanmar) amber. The new genus can be recognized from all sinoalids by the combination of hind tibia armed with five interlaced lateral spines, leaf-shaped tegmen with very broad postcostal ce...

A planthopper originally assigned to the genus Boreocixius in Surijokocixiidae (Surijokocixioidea, Fulgoromorpha) from the Ladinian (Middle Triassic) of Tongchuan in Shaanxi, northwestern China is described as Boreocixius tongchuanensis Zhang et al. sp. nov. As the third species ascribed to the genus, it differs from its congeners in having the muc...

Ghost lineages have always challenged the understanding of organism evolution. They participate in misinterpretations in phylogenetic, clade dating, biogeographic, and paleoecologic studies. They directly result from fossilization biases and organism biology. The Cylindrotomidae are a perfect example of an unexplained ghost lineage during the Mesoz...

Four new protopsyllidioid species, Paraprotopsyllidium spinosum gen. et sp. nov., Angustipsyllidium minutum gen. et sp. nov., Burmapsyllidium setosum gen. et sp. nov., and Maliawa akrawna gen et sp. nov. are described from the mid-Cretaceous Burmese amber, and assigned to a new family, Paraprotopsyllidiidae fam. nov., that we establish. These taxa...

The family name Bernaeidae was introduced by Zherikhin (1980) for two genera of whiteflies from Lebanese amber, but no diagnostic characters for the taxon were mentioned. Therefore, Shcherbakov (2000) regarded it as a nomen nudum and described Bernaeinae as a new subfamily in Aleyrodidae, listing and discussing Bernaea, though without explicitly ci...

A new planthopper family Katlasidae fam. nov. is described based on a well-preserved specimen from mid-Cretaceous Kachin amber (Burmese amber). It is characterized by vertex separated from frons with two parallel, transverse carinae; antenna with pedicel densely covered with sensory lentiform flattened plaque organs; pronotum with elevated disc, co...

Professor Ryszard Szadziewski’s adventure with entomology started in 1968 with his participation in the “Wielka Gra” (The Great Game) Polish TV 1 game show. With his mathematics and physics background and mind, he took a part in this contest as an amateur (it was one of the rules of the show). Even though he did not win this contest, he won entomol...

Amecephala pusilla gen. et sp. nov. is described and illustrated on the basis of a well-preserved female psyllid (Liadopsyllidae) in a piece of Cretaceous Myanmar amber. The new genus differs from other members of Liadopsyllidae in details of the antennae and forewings. For the first time, the presence of a circumanal ring is documented for Mesozoi...

Biting midges, nematoceran flies family Ceratopogonidae, is represented by more than 6200 valid species. This group is still poorly known and not easy in identification. Structural coloration of insect wings (WIP, or Wing Interference Pattern) is an example of an optical phenomenon called iridescence, which is based on the formation of pigment-free...

The Society of Vertebrate Paleontology (SVP) has recently circulated a letter, dated 21st April, 2020, to more than 300 palaeontological journals, signed by the President, Vice President and a former President of the society (Rayfield et al. 2020). In this letter, significant changes to the common practices in palaeontology are requested. In our pr...

The Society of Vertebrate Paleontology (SVP) has recently circulated a letter, dated 21st April, 2020, to more than 300 palaeontological journals, signed by the President, Vice President and a former President of the society (Rayfield et al. 2020). In this letter, significant changes to the common practices in palaeontology are requested. In our pr...