Hendrik Klein

Hendrik Klein
Saurierwelt Paläontologisches Museum Neumarkt · Paleontology

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252
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Publications

Publications (252)
Article
From Middle-Late Triassic continental deposits of the Timezgadiouine Formation (Argana Basin, Morocco), 17 lithofacies are identified, with alluvial fan, fluvial, floodplain and lacustrine associations. Invertebrate traces include Archaeonassa fossulata, Arenicolites isp., cf. Arenituba, Beaconites isp., cf. Camborygma, Cruziana problematica, Diplo...
Article
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Zusammenfassung Wir stellen ein neues Fährtenvorkommen aus der Hauptdolomit-Formation der S-charl-Einheit am Piz S-chalambert Dadaint (Val d'Uina, Scuol) vor. Die Fundstelle wurde 2018 von Mario Riatsch (Sent/ Scuol) entdeckt und liegt am Westgrat auf 2840 m ü. M. Die wissenschaftliche Dokumentation war nur mit Helikopter und Seilen möglich. Die Fä...
Article
Tridactyl theropod and ornithischian dinosaur tracks and trackways from Imilchil and Isli formations (Middle–?Late Jurassic, Bajocian–?) of the central High Atlas region (Morocco) are described. The Imilchil Formationconsists of brackish marine-continental deposits, and the Isli Formation is a continental red-bed succession. Considering numerous ne...
Article
Two newly discovered dinosaur track locations near the town of Bijie in Guizhou Province, southwestern China are named the Yuanbaoqing and Yejiatian tracksites. They represent the Lower Jurassic Maanshan Member of the Zijiujing Formation and Upper Jurassic Xiashaximiao Formation, respectively. The former Yuanbaoqing tracksite reveals a minimum of t...
Article
The Jurassic and Cretaceous tetrapod track record in China has incrementally expanded due to the discovery of small sites in widely dispersed geographical locations, which often lack body fossils. Many of these are theropod dominated tracksites where, despite morphological variation, identification of ichnotaxa is difficult at low taxonomic levels...
Article
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The Jiaguan Formation and the underlying Feitianshan Formation (Lower Cretaceous) in Sichuan Province yield multiple saurischian (theropod–sauropod) dominated ichnofaunas. To date, a moderate diversity of six theropod ichnogenera has been reported, but none of these have been identified at the ichnospecies level. Thus, many morphotypes have common...
Article
Fossil footprints provide the only evidence of Triassic-aged dinosaurs in Australia. While historically several Triassic dinosaur tracksites are reported, only one specimen exists for direct study. Evidence for the Rhondda colliery (Ipswich) trackmaker, Upper Triassic Blackstone Formation as a very large-bodied Triassic predatory dinosaur is not su...
Article
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The years 2020 and 2021 were marked worldwide by the corona pandemic with restrictions on fieldwork at home and abroad, with limited personal communication and restricted access to fossil and rock collections. Still, remarkable progress has been made by the international team of our working group as shown below by a number of publications, the part...
Article
The Jurassic-Cretaceous Tuchengzi Formation has yielded at least 16 known theropod-dominated tracksites from northeast China, and in this paper, a new locality, the Lishi site in Hebei Province, is documented. A tuff layer positioned stratigraphically 90 m below the trackway layer yielded a UPb date of 142.2 ± 0.9 Ma, placing the site in the Berria...
Article
The smallest non-avian theropods have often attracted attention of palaeobiological and ichnological research. Here, we describe a diminutive tridactyl track (10.2 mm in length) from the Lower Jurassic Ma’anshan Member of the Ziliujing Formation, Wuli site, Sichuan Province, China. The Wuli specimen may be the smallest theropod track in the world....
Article
An assemblage of eubrontid-type theropod tracks, representing terrestrial progression and turtle track swim tracks representing subaqueous progression characterises a newly reported site in the Middle Jurassic Zhiluo Formation in Shaanxi Province China. While eubrontid tracks are common in the Jurassic of China, turtle swim tracks have only compara...
Article
Full-text available
Rich tetrapod ichnofaunas, known for more than a decade, from the Huangyangquan Reservoir (Wuerhe District, Karamay City, Xinjiang) have been an abundant source of some of the largest Lower Cretaceous track collections from China. They originate from inland lacustrine clastic exposures of the 581–877 m thick Tugulu Group, variously divided into fou...
Article
Complete trackways of the Late Triassic archosaur ichnogenus Brachychirotherium are extremely rare and thus far mostly known from North America, though isolated imprints are abundant. Complete step cycles have been found on the upper and lower bedding surfaces in laminated mudstones and finegrained sandstones of the lowermost part of the Machraa Ab...
Chapter
Abstract. Tetrapod footprints from the marginal-marine facies of the Muschelkalk (Anisian-Ladinian) give rare insight into terrestrial vertebrate faunas of the Central European Basin, which was influenced by marine transgression during this time interval. Ichno-assemblages from both shallow-marine and fluvial deposits reflect vertebrate communities...
Article
A single well-preserved theropod trackway identified as Eubrontes was discovered in the Lower Cretaceous (Berriasian-Valanginian) Cangxi Formation on the northern margin of the Sichuan Basin This is the first report of tracks from this formation. The occurrence is consistent with reports of theropod dominated ichnofaunas from broadly coeval deposit...
Article
A newly discovered saurischian dominated tracksite in the Lower Cretaceous Jiaguan Formation in southeastern Sichuan province reveals 13 sauropod trackway segments representing ichnogenus Brontopodus and three theropod trackways. This is a typical Type 1 Jiaguan Formation deposit dominated by tetrapod tracks with no significant tetrapod body fossil...
Article
Zusammenfassung Zwei Chirotherien-Fußeindrücke aus dem Fränkischen Chirotheriensand-stein von Gambach in Unterfranken erweisen sich als die einzigen Belege für die so zahlreich beschriebenen Tetrapodenfährten aus dem Gambacher Profil im Oberen Buntsandstein. Sie werden hier als Chirotheriidae cf. Chirotherium barthii Kaup 1835 bestimmt, die man bas...
Article
A total of 31 Lower Cretaceous tetrapod tracksites have been reported from four areas in Gansu Province, China: Yanguoshia, Guanshan, Zhongpu and Honggu. Here we report on the previously undescribed track types found at seven of these sites, five of which are sauropod dominated, with one site yielding ornithopod tracks and another one yielding ther...
Article
Sauropod tracks, particularly those attributable to the ichnogenus Brontopodus are becoming increasingly well known in the Jurassic and Cretaceous of China. Globally there are indications that wide gauge forms, attributable to titanosauriform sauropods increase in the Cretaceous. However, differentiating wide and narrow gauge, broadly categorized a...
Article
Full-text available
The Zhaojue (ZJ) tracksites represent multiple steeply-inclined, track-bearing exposures in the clastic Feitianshan Formation of Sichuan Province that have been sequentially, excavated, mapped, expanded and destroyed, by quarrying, erosion or collapse, for almost three decades. The quarried area extends for more almost 1.0 km from north to south an...
Article
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Triassic tetrapod footprints have been studied since the 1830s and are now known from all of the World’s continents. An ichnotaxonomic revision of all Triassic tetrapod ichnogenera recognizes 34 valid ichnogenera: Anshunpes, Apatopus, Atreipus, Banisterobates, Batrachichnus, Batrachopus, Brachychirotherium, Brasilichnium, Capitosauroides, Characich...
Article
Full-text available
We report the first occurrence of the track type “Chirotherium” lulli from western North America. Three specimens were discovered at the “Feather Ridge Tracksite” in the Owl Rock Member of the Chinle Formation within Bears Ears National Monument of San Juan County, Utah. The ichnospecies occurs in association with Grallator isp., Brachychirotherium...
Article
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As part of a 1930s public works project, two specimens revealing a total of ~20 tracks, preserved as natural casts, were collected and built into the interior walls of what was then the Fruita Museum in western Colorado. Based on our knowledge of the regional tetrapod ichnofaunas we are able to designate these as a "Triassic specimen" and a "Jurass...
Article
Multiple sauropod-track-bearing horizons occur in the Middle Jurassic Shaqiaomu Formation, in the Chaya Group of Eastern Tibet. These add to the report of a previously described sauropod trackway, which is a famous tourist attraction and became part of a folkloric legend. From an ichnological point of view, multiple track-bearing levels are consist...
Article
Dinosaur and other tetrapod tracks are already known from multiple (~25) sites in the Early Cretaceous Hekou Group of Gansu Province. In addition to large sites (e.g. Yanguoxia) which have yielded more than 1500 tracks, small sites are important for verifying the distribution of morphotypes (ichnotaxa) over a wider region. Documentation of five new...
Article
A Late Jurassic ichnofauna from the Suining Formation at the Yejipo site, Huangba Village, Sichuan Province has been locally known for its fossil footprints since the 1980s as the name ‘Yejipo’, meaning feral chicken/pheasant indicates. The best-preserved trackway represents a large theropod (mean footprint length 55.0 cm) assigned to Megalosauripu...
Article
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Poorly preserved tracks have limited ichnotaxonomic or biotaxon utility, but may reveal useful information about the paleoenvironment, behavior and track taphonomy. Eight mostly parallel to sub parallel trackway segments (T1–T8) were registered on a truncation surface in the Lower Cretaceous Luohe Formation of Shaanxi Province. These attest to the...
Article
Full-text available
The Solvey Quarry of Bernburg is one of the most important ichnosites from the Muschelkalk of the Germanic Basin. Extensive surfaces with long chirotheriid trackways have been discovered and assigned to Chirotherium and Isochirotherium. Some undescribed step cycles from this site are analysed here and assigned to Synaptichnium isp. These footprints...
Article
Subaqueously-registered trackways of marine tetrapods are only rarely preserved; some of the best examples so far come from the Shizishan Member (Member II) of the Guanling Formation (Middle Triassic, Anisian-Ladinian) of Yunnan Province in southwestern China. The unit is well-known for the Luoping Biota including abundant skeletons of marine repti...
Article
The Xixipo dinosaur tracksite in the Chuanjie Formation of Yunnan Province is one of the 14 Chinese sites yielding sauropod tracks from between the Triassic–Jurassic and Jurassic–Cretaceous boundaries, but is only one of the two that represent the Middle Jurassic. Although it is a small site, it adds incrementally to the overall track record of the...
Article
Large dinosaur tracks were recently reported from locations in the Pterosaur-Yadan National Geological Park situated about 100 km south of Hami in Xinjiang Province, China. The park comprises a substantial area in a much larger arid region comprising and extensive spectrum of Cretaceous, siliciclastic, Tugulu Group, lithofacies representing proxima...
Article
Nonmarine biostratigraphic/biochronologic schemes have been created for all or parts of the late Carboniferous–Middle Triassic using palynomorphs, megafossil plants, conchostracans, blattoid insects, tetrapod footprints and tetrapod body fossils, and these provide varied temporal resolution. Cross correlation of the nonmarine biochronologies to the...
Article
A dinosaur track assemblage from a remote, high altitude (∼4,500–5,000 m) site on the is the first to be reported from this area. The location, here named the Yangguang tracksite, represents the Lower Cretaceous part of the Kezilesu Group consisting of feldspathic sandstones and dated as Hauterivian–Barremian on the basis of the palynoflora. As the...
Article
It has long been known that Lower Jurassic theropod track assemblages are dominated by theropod tracks including Eubrontes and Grallator, known from North America since the mid 19th century. It was only comparatively recently that it was recognised that these ichnogenera where also abundant in other regions including China, making them important co...
Article
An archosaur trackway consisting of 10 successive pes imprints has been recovered from the Late Triassic Irohalene Member of the Timezgadiouine Formation (Argana Basin, Morocco). Footprints are tetradactyl-pentadactyl, and show an elongated ‘heel’, probably enclosing the trace of digit V. The trackway lacks a manus impression, which could be due to...
Article
Our recent comprehensive review of the Permian-Early Triassic tetrapod tracksites from South Africa includes a revision of the ichnotaxonomy and the incorporation of a large quantity of new material. The paper also discusses, in light of the revised ichnotaxonomy and palaeontology of several sites, trackmaker attribution and the biostratigraphy of...
Article
The newly discovered Maoshajing tracksite in the Lower Jurassic Longtoushan Group of Guizhou Province, China is dominated by well-preserved medium-sized theropod tracks of the Grallator-Eubrontes plexus type, and associated with a few sauropod tracks. This saurischian dominated ichnofauna is typical of the Lower Jurassic biochron of China and elsew...
Article
Full-text available
From the beginning of 2018 to the autumn of 2019, a team of 18 authors worked very hard to publish Schneider et al. (2019) “Late Paleozoic–early Mesozoic continental biostratigraphy —Links to the Standard Global Chronostratigraphic Scale”. This follows some important publications on nonmarine Permian biostratigraphy and biochronology in the volume...
Article
Full-text available
From the beginning of 2018 to the autumn of 2019, a team of 18 authors worked very hard to publish Schneider et al. (2019) “Late Paleozoic–early Mesozoic continental biostratigraphy — Links to the Standard Global Chronostratigraphic Scale”. This follows some important publications on nonmarine Permian biostratigraphy and biochronology in the volume...
Conference Paper
Full-text available
We report on a new dinosaur tracksite in the Hauptdolomit Formation of the S-charl unit at Piz S-Chalambert Dadaint (Val d'Uina, Scuol). The site was discovered in 2018 by a local hunter on the western ridge at 2840 m a.m.s.l. and could only be accessed by us using ropes and helicopters. The footprints occur in the uppermost part of the Hauptdolomi...
Article
Slabs with tetrapod tracks from the Madigou Village quarry in the Tuchengzi Formation have famously been used as building stones at the imperial summer resort locality, also referred to in the ichnological literature as the “Chengde Summer Palace.” The Jurassic-Cretaceous boundary Tuchengzi Formation has elsewhere in the region yielded differently...
Article
A new dinosaur tracksite is reported from continental red beds of the Jurassic (Late Bathonian-?Callovian) Isli Formation along the northern flank of the Aït Ali ou Ikkou Syncline of the Imilchil area, Central High Atlas, Morocco. The succession was deposited in a fluvio-lacustrine environment, and contains at least fourteen track-bearing levels. T...
Conference Paper
Full-text available
Tracks documenting subaqueous activity of terrestrial or semi-aquatic tetrapods are common in the Mesozoic record. Numerous examples of swimming dinosaurs or other tetrapods, in particular turtles, phytosaurs and crocodylomorphs are known. Marine forms, however, rarely left their imprints on the bottom of shallow water while buoyant, drifting or ac...
Article
The Cretaceous Hengshan Formation at the Baxian tracksite, central Zhejiang has yielded more than 20 sauropod tracks with a considerable size range (pes length ranging from 22.7–83.7 cm). The assemblage consists of a more complete trackway of a large animal, a less complete trackway of a large animal with a similar orientation and, a small manus-pe...
Conference Paper
Full-text available
Trackways of Late Triassic Brachychirotherium (archosaur) are known from few specimens, mostly from North America. Complete step cycles have been found on the upper and lower bedding surface in laminated mudstone and fine sandstone of the upper Oued Oum Er Rbiaa Formation (Late Triassic). This is the first trackway of the ichnogenus in North Africa...
Conference Paper
A new dinosaur tracksite was discovered in continental red beds of the Isli Formation (late Bathonian-?Callovian) along the northern flank of the Aït Ali ou Ikkou Syncline in the Imilchil area, Central High Atlas, Morocco. The area has an extraordinary scientific potential, as the layers, which have been deposited in a fluvio-lacustrine environment...
Conference Paper
Trackways of Late Triassic Brachychirotherium (archosaur) are known from few specimens, mostly from North America. Complete step cycles have been found on the upper and lower bedding surface in laminated mudstone and fine sandstone of the upper Oued Oum Er Rbiaa Formation (Late Triassic). This is the first trackway of the ichnogenus in North Africa...
Conference Paper
Trackways of Late Triassic Brachychirotherium (archosaur) are known from few specimens, mostly from North America. Complete step cycles have been found on the upper and lower bedding surface in laminated mudstone and fine sandstone of the upper Oued Oum Er Rbiaa Formation (Late Triassic). This is the first trackway of the ichnogenus in North Africa...
Conference Paper
Middle to Late Triassic redbeds of the Timezgadiouine Formation (T4-T5, Anisian-Carnian) in the Irohalene area (Argana Basin, Western High Atlas, Morocco) consist of cyclical clastic sedimentary rocks, which are mainly composed of conglomerates, sandstones, siltstones, claystones and mudstones, that were deposited in an alluvial plain to mudflat en...
Book
Full-text available
Conference abstract volume and field trip guide for a 3-day field trip focused on continental ichnology of Germany
Article
The Middle Triassic (Anisian) Guanling Formation in Yunnan and Guizhou provinces, southwestern China, is well-known for its marine-reptile fauna with numerous well-preserved skeletons. Tetrapod footprints from this unit have been reported of both terrestrial and marine forms documenting a marginal marine environment in the eastern Tethys region. In...
Article
A sauropod track assemblage from a surface in the Upper Jurassic fluvial Tianchihe Formation at the Guanxi site, Shanxi Province, China, is re-documented and analysed in detail, together with a recently discovered new locality situated nearby. In contrast to earlier reports, the former includes four trackways plus isolated tracks (41 tracks in tota...
Article
Complete trackways of the Late Triassic archosaur ichnogenus Brachychirotherium are extremely rare and thus far mostly known from North America, though isolated imprints are abundant. Complete step cycles have been found on the upper and lower bedding surfaces in laminated mudstones and finegrained sandstones of the lowermost part of the Machraa Ab...
Article
Complete trackways of the Late Triassic archosaur ichnogenus Brachychirotherium are extremely rare and thus far mostly known from North America, though isolated imprints are abundant. Complete step cycles have been found on the upper and lower bedding surfaces in laminated mudstones and finegrained sandstones of the lowermost part of the Machraa Ab...
Article
Full-text available
Complete trackways of the Late Triassic archosaur ichnogenus Brachychirotherium are extremely rare and thus far mostly known from North America, though isolated imprints are abundant. Complete step cycles have been found on the upper and lower bedding surfaces in laminated mudstones and fine-grained sandstones of the lowermost part of the Machraa A...
Article
Complete trackways of the Late Triassic archosaur ichnogenus Brachychirotherium are extremely rare and thus far mostly known from North America, though isolated imprints are abundant. Complete step cycles have been found on the upper and lower bedding surfaces in laminated mudstones and fine-grained sandstones of the lowermost part of the Machraa A...
Conference Paper
Full-text available
The Carboniferous, Permian, and Triassic chronostratigraphic scales based on marine rocks and fossils are reasonably well defined and of global utility, but the situation is much different for non-marine deposits of this time interval. Due to the Carboniferous–Permian glaciation, the Hercynian-Appalachian-Ancestral Rocky Mountain orogenies, and oth...
Conference Paper
Presentation type: oral communication We report on a series of enigmatic traces from the Early Jurassic of Poland that co-occur with tridactyl dinosaur tracks on the same surface. Apart from theropods, ornithischian as well as tracks of sauropods are known from coeval deposits. Swim traces of theropods and an isolated footprint of a mammal have als...
Article
An Early Cretaceous dinosaur ichnofauna is reported from the Taoqihe Formation (Aptian–Albian) in Heilongjiang Province, China. The assemblage, which comprises >70 tracks in 5 trackways, includes parallel sauropod trackways, a toe-trace dominated tridactyl ornithopod trackway and elongate tridactyl theropod tracks, the latter tentatively be attribu...
Article
Full-text available
The Val Gardena Formation of the Dolomites region in northern Italy preserves the most significant assemblage of Late Permian tetrapod footprints in the world. More than 120 years of collecting resulted in about 900 publicly accessible specimens from the study area. This huge amount of data is comprehensively revised in the light of recent advances...