Francisco J. Goin

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• Dr.
• Senior Researcher at National University of La Plata

Desde hace más de una década se vienen realizando trabajos de prospección y rescate paleontológicos en canteras sujetas a la actividad minera en el área de Sierras Bayas, Partido de Olavarría, en el centro de la Provincia de Buenos Aires. Los mismos han permitido conocer nuevas secuencias sedimentarias tanto neógenas como cuaternarias, las que apor...

Carneiro et al. (2025) described a new genus and species of derorhynchid, Streptorhynchus tropicalis, from the Itaboraí Basin (Itaboraí II, early Eocene, Itaboraian SALMA sensu Bergqvist et al. 2024), southeastern Brazil. However, Streptorhynchus is preoccupied by a Permian brachiopod Streptorhynchus King, 1850. Therefore, we propose a new generic...

Derorhynchidae is a Paleogene clade of South American and Antarctic metatherians. Among the localities with representatives of this clade, the Itaboraí Basin stands out by the identification of three genera and at least five species. Herein, we present a new derorhynchid from this locality, Streptorhynchus tropicalis gen. et sp. nov., and discuss i...

The Itaboraí Basin, located in Itaboraí, Rio de Janeiro, Brazil, has preserved one of the most important early Eocene vertebrate assemblages of the Southern Hemisphere. Among its representatives, Metatheria comprises its most diverse group, with a minimum of 29 genera and 43 species. Considering this diversity, tiny and very small taxa are represen...

Carolocoutoia ferigoloi is one of the largest metatherians of the Itaboraí fauna (Early Eocene, Southeastern Brazil). It was originally identified from a single upper molar with a markedly bunodont crown, wrinkled enamel, and a robust protocone. Here, we describe a nearly complete right dentary of C. ferigoloi with the alveoli of the premolars and...

The fossil metatherian assemblage from La Venta (Middle Miocene, Colombia) is one of the most diverse in South America, and it is critical to understand the Neogene radiation of this group in this continent. La Venta contains the northernmost record of Thylacosmilidae Riggs, 1933 (Metatheria, Sparassodonta): Anachlysictis gracilis Goin, 1997, the f...

Se describe a Lutreolina tonnii sp. nov. (Mammalia, Metatheria, Marsupialia, Didelphimorphia, Didelphidae) procedente de niveles del Pleistoceno Temprano (o Temprano–Medio) de las barrancas costeras de los alrededores de Necochea, Provincia de Buenos Aires, Argentina. El holotipo, un maxilar derecho con los M1-3, se destaca por su gran tamaño compa...

Estelestes ensis Novaceck et al., 1991 is a curious Paleogene metatherian mammal recognized on the basis of a single specimen from Baja California (Mexico) in southern North America. It comes from early Eocene (Wasatchian age) levels of the Las Tetas de Cabra Formation at “Marsupial Hill” in the Lomas Las Tetas de Cabra site (also known as Punta Pr...

Durante mucho tiempo la principal fuente de información sobre mamíferos del Cretácico Superior de Gondwana estuvo dominada por los registros provenientes de la Patagonia argentina y Madagascar, en conjunto con registros más antiguos distribuidos en África continental y Australia. En el registro del Campaniano-Maastrichtiano predominan numerosas esp...

A noticeable aspect of metatherian evolution in South America is the influence of climatic oscillations in their macroevolutionary patterns, involving major radiations during warming phases, and extinctions and functional turnovers during cooling phases. Two aspects explain why southern metatherians were more successful than their Northern Hemisphe...

Morotodon aenigmaticus gen. et sp. nov. (Mammalia, Metatheria, ?Herpetotheriidae) from the early or early-middle Miocene of equatorial Africa (Moroto II locality, Moroto District, northeastern Uganda) is characterized by a short anterior cingulum, a buccal shelf, a well-developed hypoconulid in a central position, and a trigonid and talonid with si...

The terrestrial mammal diversity recovered from levels of Banco Negro Inferior (Lower Black Bank) at Punta Peligro and surrounding areas, in southeastern Chubut Province (Argentina) is detailed. Most of the metatherian taxa here described are new for this mammalian assemblage, which in turn is the base of the late Danian (early Paleocene, Peligran...

The territory of the Chubut province, Argentina, is one of the richest regions in vertebrate fossils from the Lower and Middle Cenozoic of South America. In many of its classic localities there are several successive fossil levels superimposed and associated with sediments of volcanic origin that allow a precise chronological control. During this (...

The Cretaceous was an important period in the evolution of vertebrates, both on land and in the oceans. Reconfiguration of landscapes by the Pangaean break-up and subsequent oceanic phase that opened new seaways enabled the establishment of new migration routes and also created wide, shallow epicontinental seas that are considered to boost diversit...

The fossil record from Cenozoic sediments provides a great deal of information that has direct bearing on the early assembling of modern Patagonian ecosystems. In this synthesis, we revise selected fossil marine and terrestrial records from the last 66 Ma with the aim of understanding major shifts of Patagonian biotas. From the Paleocene to the mid...

The Upper Cretaceous (Cenomanian) Mata Amarilla Formation in western Central Santa Cruz Province of Argentina has yielded fragmentary teeth of a large ?docodontan, an australosphenidan, a meridiolestidan (Amarillodon meridionalis gen. et sp. nov.), and a stem dryolestid (Treslagosodon shehuensis gen. et sp. nov.). These represent the first possible...

Hasta recientemente, la principal fuente de información sobre mamíferos mesozoicos sudamericanos provenía de la Patagonia argentina, con numerosas especies de mamíferos no-Theria, incluidos en los clados Gondwanatheria/Allotheria, Meridiolestida y Driolestidae. Sin embargo, hallazgos recientes en rocas del Cretácico Superior de la Formación Dorotea...

A new early Eocene, small-sized metatherian from the Itaboraí fauna is described. The new taxon is recognized on the basis of an incomplete dentary recovered from fissure fillings in the travertine limestones from the Itaboraí Basin, State of Rio de Janeiro, Brazil. The phylogenetic analysis placed the new genus and species as the sister taxon of D...

A new species of Philander (Marsupialia, Didelphidae) from the late Miocene of northwestern Argentina. A new species of Philander (Marsupialia, Didelphimorphia, Didelphidae), from levels of the Andalhuala Formation (late Miocene) at Tio Punco locality, Tucumán Province (Argentina) is described. Compared to the living species of the genus, the new s...

The record of extinct African metatherians (Mammalia, Theria) is scanty, restricted in time (Eocene–Miocene), and its taxonomy is still subject of debate. A review of all African metatherians, or alleged metatherians, known up to now, led us to the recognition of only three taxa referable to this group: (1) Kasserinotherium tunisiense (Peradectoide...

The Mesozoic plate tectonic and paleogeographic history of the final break up of West Gondwana had a profound effect on the distribution of terrestrial vertebrates in South America. As the supercontinent fragmented into a series of large landmasses (South America, Antarctica, Australia, New Zealand, the Indian subcontinent, and Madagascar), particu...

In the last decades, several discoveries have uncovered the complexity of mammalian evolution during the Mesozoic Era, including important Gondwanan lineages: the australosphenidans, gondwanatherians, and meridiolestidans (Dryolestoidea). Most often, their presence and diversity is documented by isolated teeth and jaws. Here, we describe a new meri...

Durante la mayor parte del Cenozoico, los grandes vertebrados carnívoros de América del Sur estuvieron repre-sentados por aves fororacoideas, cocodrilos, serpientes gigantes y mamíferos metaterios como los Borhyaenidae. Entre estos últimos, Arctodictis munizi Mercerat, 1891 representa al boriénido de mayor tamaño hasta ahora conocido. Un nuevo espé...

Modern grasses in the family Poaceae exhibit exceptional taxonomic and ecological diversity, with grassland-dominated habitats covering about 40% of Earth’s land surface. Grasslands today occur in a range of temperate to tropical ecosystems, spanning low to high altitudes and wet to arid climates. However, because of the scant fossil record of gras...

Materials from the localities of Araia d'Alcora in the Ribesalbes-Alcora Basin (Spain, early Miocene, Bio-zone C, MN4) have yielded an assemblage of erinaceids and metatherians, relatively rich for an Iberian site. The most common erinaceid is the gymnure Galerix symeonidisi Doukas, 1986, present in almost all of the studied sites. Other erinaceids...

Se describe a Magallanodon baikashkenke gen. et. sp. nov., un nuevo mamífero gondwanaterio del Cretácico tardío de la Región de Magallanes, en el sur de Chile (Valle del Río de Las Chinas, Estancia Cerro Guido, norte de Puerto Natales, Provincia de Última Esperanza). Las capas portadoras se ubican entre los niveles del Campaniano tardío-Maastrichti...

The diversity of the latest Paleocene-early Eocene metatherian fauna of Itaboraí locality, Brazil, is presented. Almost all major lineages (orders) of South American Cenozoic marsupials are already present in the Itaboraí fauna: Sparassodonta (Borhyaenoidea), Pediomyidae, Peradectoidea, “Didelphimorphia”, Paucituberculata, and Polydolopimorphia. Re...

We describe Magallanodon baikashkenke gen. et. sp. nov., a new gondwanatherian mammal from the Late Cretaceous of the Magallanes Region in southern Chile (Río de Las Chinas Valley, Estancia Cerro Guido, north of Puerto Natales city, Última Esperanza Province). The mammal-bearing layer is placed within the Late Campanian-Early Maastrichtian levels o...

A new Paleogene metatherian from locality IAA 1/90, Marambio (Seymour) Island in the Antarctic Peninsula is described. Pujatodon ektopos, gen. et sp. nov., is recognized on the basis of a tiny lower left molar recovered from early Eocene (late Ypresian) levels of the Cucullaea I Allomember, La Meseta Formation. The tooth is characterized by its sma...

With two recognized species, the extinct (late Eocene) Groeberia is periodically at the centre of systematic disputes. On the basis of its best-preserved specimen, we describe in detail the dental and functional morphology of the masticatory apparatus of Groeberia minoprioi. A review of its dental anatomy indicates that Groeberia has a tribosphenic...

Sparassodonts were the main mammalian predators during most of Cenozoic in South America. The lower Eocene Itaboraí Basin/Formation includes the second oldest fossil records of this group in South America: Patene and cf. Nemolestes. Patene is by far the most abundant sparassodont from this formation, with more than 30 specimens referable to a singl...

The sparassodonts were the main mammalian predators during most of Cenozoic in South America. The Itaboraí Basin/Formation (lower Eocene) includes two of the oldest fossil records assigned to this group: Patene Simpson, 1935 and cf. Nemolestes Ameghino, 1902. Patene is by far the most abundant sparassodont from this formation, with more than 30 spe...

The fossil record of terrestrial mammals in Antarctica is temporally and geographically constrained to the Eocene outcrops of La Meseta and Submeseta formations in Seymour (Marambio) Island in West Antarctica. The faunal assemblage indicates a clear South American imprint since all the groups have a close phylogenetic relationship with Cretaceous a...

La asociación de mamíferos del Mioceno medio de La Venta (Colombia), particularmente de aquellos aflorantes en el Desierto de La Tatacoa (Departamento del Huila), es una de las más diversas hasta ahora conocidas para el Neógeno de América del Sur. Constituye, además, una referencia obligada para el estudio de varios linajes de mamíferos vivientes (...

Sparassodonts were the main mammalian predators during most of Cenozoic in South America. The lower Eocene Itaboraí Basin/Formation includes the second oldest fossil records of this group in South America: Patene and cf. Nemolestes. Patene is by far the most abundant sparassodont from this formation, with more than 30 specimens referable to a singl...

Marsupials and their extinct relatives comprise the Subclass Metatheria of Mammalia. Several physiological and morphological features distinguish this group of mammals from all others, e.g., a reproductive cycle characterized by a short gestation period and a long lactation, and a generalized, adult dental formula that includes five upper and four...

The Pediomyiodea has been largely regarded as a Late Cretaceous North American metatherian lineage; however, some studies have suggested that a few South American taxa could be related to this family. Herein, Austropediomys marshalli gen. et sp. nov. from the Itaboraí Basin, Brazil (lower Eocene – Itaboraian SALMA), is described. Austropediomys gen...

In the last three decades, records of tribosphenidan mammalsfrom India, continental Africa, Madagascar and South Americahave challenged the notion of a strictly Laurasian distributionof the group during the Cretaceous. Here, we describe a lowerpremolar from the Late Cretaceous Adamantina Formation, SãoPaulo State, Brazil. It differs from all known...

Despite a global fossil record, Metatheria are now largely restricted to Australasia and South America. Most metatherian paleodiversity studies to date are limited to particular subclades, time intervals, and/or regions, and few consider uneven sampling. Here, we present a comprehensive new data set on metatherian fossil occurrences (Barremian to e...

The Quebrada de Los Colorados Formation (Los Cardones National Park, Salta Province, north-western Argentina), is an Eocene–Oligocene unit well represented in the Calchaqu�ı Valley area. Here we describe a new metatherian association recorded from the base of this formation, inferred as middle Eocene. Represented taxa are: Sparassodonta, family ind...

We compare the main histological features of the digestive tract of three selected New World didelphid marsupials (Mammalia, Metatheria, Didelphimorphia, Didelphidae): the White‐eared Opossum Didelphis albiventris Lund, 1840, the Short‐tailed Opossum Monodelphis dimidiata Wagner, 1847, and the Patagonian Opossum Lestodelphys halli Thomas, 1921. The...

Recientemente se dio a conocer un nuevo marsupial (aún innominado) procedente de niveles del Alomiembro Cucullaea I (Fm. La Meseta, Eoceno temprano) en la localidad fosilífera IAA 1/90, Isla Marambio (Seymour), al este de la Península Antártica (Reguero et al. 2016). El ejemplar (MLP 14-I-10-20) consiste en un molar inferior izquierdo aislado, comp...

We summarize the configuration of plates, geographical barriers, and possible dispersal events during the Late Cretaceous–Cenozoic between North and South America, the Caribbean, Antarctica, and Australia. The arrival of metatherians in South America was a Late Cretaceous event, and probably a Maastrichtian one. There are few doubts that the first...

The early Paleocene diversity of metatherians in Tiupampan faunas of South America and the pre-Tiupampan Paleocene polydolopimorphian Cocatherium speak in favor of an earliest Paleocene or Late Cretaceous dispersal of metatherians from North America. No Late Cretaceous metatherian or eutherian mammals have been recovered to date in South America, b...

South American living metatherians are relatively inconspicuous and comprise ca. 10 % of the region’s mammal species richness. Most of them are small-sized (<150 g), with long tails and grasping hands and feet, and resemble one another in their general appearance. Individuals are solitary, nocturnal or crepuscular, and most of them are arboreal or...

We describe a new mammal coming from site IAA 1/90, Marambio (Seymour) Island, Antarctic Peninsula. The specimen (MLP 14-I-10-20) is a tiny lower left molar partially worn (total length: 2.24 mm; trigonid length: 0,72 mm; trigonid width: 1.11 mm; talonid width: 1.45 mm), and represents a new taxon for the Antarctic Paleogene mammalian assemblage. I...

The Cretaceous was a special time for the Indian plate as it was separated from Gondwana landmasses and started its northward journey across the Tethys Sea towards the Equator. The northward movement of this plate implied shifting latitudes and climate belts, until it finally collided with Asia during the early Cenozoic. Geophysical data and plate...

The Cretaceous was a special time for the Indian plate as it was separated from Gondwana landmasses and started its northward journey across the Tethys Sea towards the Equator. The northward movement of this plate implied shifting latitudes and climate belts, until it finally collided with Asia during the early Cenozoic. Geophysical data and plate...

A diverse assemblage of extinct mammals of early–middle Eocene age (Ypresian–Lutetian boundary) come from the Patagonian localities of La Barda and Laguna Fría around Paso del Sapo in northwestern Chubut Province (Argentina). Metatherians are well represented, mostly by dental remains of ‘Didelphimorphia’, Paucituberculata, Sparassodonta, Microbiot...

With less than 3 g of estimated body mass, the early Eocene Minusculodelphis minimus Paula Couto (Mammalia, Metatheria, Jaskhadelphyidae) is one of the smallest mammals, living or extinct. It has alternatively been regarded as a didelphid or a derorhynchid Bameridelphian,^ or even as an eometatherian marsupial. Here, we describe a new species of Mi...

A new species of Sparassodonta (Mammalia, Metatheria), Lycopsis padillai, sp. nov., is described on the basis of a partial left maxilla with M1–M4 and fragments of lacrimal and jugal. The material comes from the early to early middle Miocene Castilletes Formation, La Guajira Peninsula, Colombia. This specimen represents the northernmost record of a...

With more than 100 species, living South American marsupials (Mammalia, Metatheria) give only a glimpse of the much higher taxonomic and ecological diversity acquired by metatherians throughout the Cenozoic Era. The term Metatheria designs a taxon within Mammalia that includes not only Marsupialia but also all therian mammals more related to Marsup...

Diversity, dietary, and body mass analyses suggest that the early Eocene represents the major radiation event in South America metatherian evolutionary history. During this period, representatives of all orders typical of the Paleogene reached their greatest diversity (i.e., “basal ameridelphians”; Polydolopimorphia Polydolopiformes, and Bonaparthe...

The Metatheria include not only marsupials but all therians more related to Marsupialia than to the Eutheria. Marsupialia is considered as a metatherian crown group including all extant marsupials, their common ancestor and all of their descendants. “Ameridelphia” is not a natural group. Australidelphia includes the Microbiotheria and all Australas...

In order to understand the timing of metatherian radiations, adaptations, and extinctions, it is important to review the variety of strongly interrelated contexts that defined them. (1) During the Cenozoic Era, global climates shifted from Greenhouse to Icehouse conditions; this major change was quite obvious by the Eocene–Oligocene boundary (ca. 3...

This book summarizes major aspects of the evolution of South American metatherians, including their epistemologic, phylogenetic, biogeographic, faunal, tectonic, paleoclimatic, and metabolic contexts. A brief overview of the evolution of each major South American lineage ("Ameridelphia", Sparassodonta, Didelphimorphia, Paucituberculata, Microbiothe...

Supplementary Data for: Catalina Suarez, AnalÍa M. Forasiepi, Francisco J. Goin & Carlos Jaramillo (2016) Insights into the Neotropics prior to the Great American Biotic Interchange: new evidence of mammalian predators from the Miocene of Northern Colombia, Journal of Vertebrate Paleontology, 36:1, DOI: 10.1080/02724634.2015.1029581

http://www.tandfonline.com/eprint/brK48uYJ53PsMdUaijm2/full

We describe a new metatherian of the genus Gaylordia Paula Couto, from the early Eocene Itaboraí Basin, Brazil. This new species (estimated body mass ~60-90.9 g) is based on lower and upper dentition elements and is characterized by a more plesiomorphic dental morphology with respect to G. macrocynodonta Paula Couto (with a larger body mass, estima...

The platyrrhine primates, or New World monkeys, are immigrant mammals whose fossil record comes from Tertiary and Quaternary sediments of South America and the Caribbean Greater Antilles. The time and place of platyrrhine origins are some of the most controversial issues in primate palaeontology, although an African Palaeogene ancestry has been pre...

In 2009 the Smithsonian Tropical Research Institute initiated a series of field trips to prospect deposits in the northernmost areas of the La Guajira Peninsula in northern Colombia. As a result of these efforts, hundreds of fossil vertebrates have been collected, including reptiles, mammals, fishes, and birds. Here, we report the first record of a...

The Early Eocene strata (Cucullaea I Allomember) of La Meseta Formation of Seymour Island, Antarctic Peninsula contain a diverse mammalian assemblage including gondwanatherians, marsupials, pilosans, litopterns and astrapotheres. This composition reflects a strong paleogeographical link between Paleogene faunas of Patagonia and West Antarctica. A s...

We describe a new metatherian mammal, Fieratherium sorex gen. et sp. nov., found in western Argentina (Quebrada Fiera locality, southern Mendoza Province), in Late Oligocene deposits (Agua de la Piedra Formation, Deseadan age). The only known specimen is a juvenile with fragments of both dentaries, the right maxilla and a fragment of the left prema...

An appraisal of Paleogene floral and land mammal faunal dynamics in South America suggests that both biotic elements responded at rate and extent generally comparable to that portrayed by the global climate pattern of the interval. A major difference in the South American record is the initial as well as subsequent much greater diversity of both Ne...

Late Cretaceous sedimentary rocks are only exposed around the northern part of West Antarctica, on the South Shetland Islands and James Ross Island Group, Weddell Sea. They were deposited in very different tectonic settings and environments. The South Shetland Island sequence represents a Cenozoic outer-arc (Birkenmajer 1995) or fore-arc (Elliot 19...

West Antarctica is an ensemble of blocks that have moved independently of each other and of cratonic East Antarctica. The onset of Terra Australis orogenesis was responsible for the termination of passive margin sedimentation along the greater part of the Pacific margin of Gondwana and began a long-lived process of accretion that added much of the...

One of the most intriguing paleobiogeographical phenomena related to the final stage of Gondwanan breakup is the close similarities and, in most cases, inferred sister-group relationships, of a number of terrestrial and marine/coastal vertebrate taxa recovered from Paleogene deposits of West Antarctica with those from other continents (South Americ...

Pocos días antes de las festas de Navidad y Año Nuevo del pasado año, la comunidad científca nacional e internacional recibió con consternación y dolor la noticia de que el 23 de diciembre había fallecido en la ciudad de La Plata, a los 87 años de edad, el Dr. Rosendo Pascual. Fue Investigador Superior del CONICET, Jefe de la División Paleontología...

Patagonia (Argentina) is a unique region where two orders of American marsupials (Paucituberculata and Didelphimorphia) and one order of Australasian marsupials (Microbiotheria) are currently found. Dromiciops gliroides (Microbiotheria) is one of four species of marsupials living in southern Argentina as well as southern Chile. Dromiciops (monito...

Pocos días antes de las festas de Navidad y Año Nuevo del pasado año, la comunidad científca nacional e internacional recibió con consternación y dolor la noticia de que el 23 de diciembre había fallecido en la ciudad de La Plata, a los 87 años de edad, el Dr. Rosendo Pascual. Fue Investigador Superior del CONICET, Jefe de la División Paleontología...

During the past two decades, geological explorations of the James Ross Basin, Weddell Sea, have revealed that this basin, located off the northeast tip of the Antarctic Peninsula (West Antarctica), contains one of the most important records of Late Cretaceous and Early Paleogene life in the Southern Hemisphere. The early explorer and scientist Otto...

En este trabajo se encara la revisión sistemática de dos órdenes de Metatheria (Mammalia) sudamericanos de la Edad Colhuehuapense (Mioceno temprano): Microbiotheria y Polydolopimorphia. En primer lugar, se comentan y discuten las homologías de las cúspides de los molares superiores e inferiores de los Argyrolagoidea (Polydolopimorphia). En segundo...

Late Cretaceous/Paleogene strata of the James Ross Basin, West Antarctica (63°S), yield the only association of marine and terrestrial vertebrates and plants known from anywhere in the continent. Fossil marine vertebrates from the extensive Cretaceous (Coniacian, Campanian and Maastrichtian) marine deposits of the James Ross Basin include marine re...

The Late Cretaceous terrestrial biota from Antarctica come from the marine sediments of the James Ross Basin and the western flank of the Antarctic Peninsula. A compilation of data for Cretaceous and Paleogene Antarctic floras from these areas provides different perspective on floristic and vegetation change when compared with those coeval floras f...

Despite the abundant fossil content of the Mata Amarilla Formation (Southern Patagonia, Santa Cruz Province, Argentina), its age has always generated a considerable number of questions and debates. The chronological data provided by invertebrates, dinosaurs, fish, turtles, plesiosaurs and fossil flora are contradictory. In this work, twenty U-Pb sp...

At least fi ve successive phases in South American mammalian evolution can be envisaged, the oldest one being largely hypothetical: (1) Early Gondwanan (?Late Triassic-Early Creta-ceous), mammals of Pangaean (triconodontids with amphilestid affi nities) and Gondwa-nan (autralosphenids) origin; (2) Late Gondwanan (Late Cretaceous), strong endemism i...

We describe two isolated molariforms recovered from early-middle Eocene (early Lutetian) levels of northwestern Patagonia, Argentina. Comparisons with major lineages of therian and non-therian mammals lead us to refer them to a new genus and species of Gondwanatheria (Allotheria). There is a single root supporting each tooth that is very short, wid...

Isolated teeth of a new species of the rare batomorph genus Hypolophodon were found in Paleocene beds of the Lefipán Formation (Maastrichtian-Paleocene) of southern Argentina. The new species H. patagoniensis differs from the type species H. sylvestris in the absence of an occlusal tranverse crest on the crown of unworn teeth and of a broad and rou...