Francesco Boschin

Università degli Studi di Siena | UNISI · Department of Environment, Earth and Physical Sciences

Most of the Middle Palaeolithic evidence of Central Italy still lacks a reliable chrono-cultural framework mainly due to research history. In this context Grotta dei Santi, a wide cave located on Monte Argentario, on the southern coast of Tuscany, is particularly relevant as it contains a very well preserved sequence including several Mousterian la...

Data about exploitation of carnivore, rodent and lagomorph taxa from the Epigravettian of Grotta Paglicci are presented in this paper. Grotta Paglicci is characterized by a long term stratigraphy, whose Epigravettian part covers, quite continuously, a time span from about 20,000 years cal. BP to about 13,500 years cal. BP. During this time, among m...

This paper poses a question on the interpretation of caprine “kill‐off patterns” in some prehistoric sites of the Caput Adriae (northern Adriatic region, Mediterranean area). In particular, caprine kill‐off data from layers 2 (Late Neolithic‐Copper Age) and 2a (Middle Neolithic) of Grotta dell’Edera (Trieste Karst, north‐eastern Italy) are presente...

The identification of the earliest dogs is challenging because of the absence and/or mosaic pattern of morphological diagnostic features in the initial phases of the domestication process. Furthermore, the natural occurrence of some of these characters in Late Pleistocene wolf populations and the time it took from the onset of traits related to dom...

The zooarchaeology and taphonomy of large mammal remains from Grotta dell’Edera (Trieste Karst, northern Italy) are presented in this paper. A Mesolithic (Sauveterrian) Preboreal assemblage testifies to a first human occupation, followed by a second longer Sauveterrian frequentation from the Boreal until the beginning of the Atlantic. The last Meso...

In this paper, we apply a two-dimensional (2D) Geometric morphometric analysis to a sample of Epigravettian lithic artefacts with the aim of assessing the potential of such an approach to study Epigravettian lithic assemblages. The lithic sample comes from layer 9c2 (Evolved Epigravettian, Upper Palaeolithic, about 18,000-19,000 years ago) of Grott...

X-ray microCT offers the possibility of studying the internal structure of animal remains by detecting age-related changes in bone microstructure. In the present work, we analyse developmental patterns of the diaphyseal structure in canids. In particular, the first metacarpal of current and archaeological individuals of red fox (Vulpes vulpes) and...

The mobility of hunter-gatherer groups is crucial in understanding Palaeolithic settlement dynamics. The concept of mobility cannot be separated from the space in which it occurs, including landscape components, localization of critical resources and of other sites, and routes between them. Nevertheless, the landscape is not constant in time due to...

Residue analysis in the field of Palaeolithic studies represents a powerful source of information, as well as a challenging operation of deduction, based on the interpretation of what remains of organic and inorganic compounds that testify to the ancient use of lithic artefacts. The disposal of reliable methods for residue characterisation is thus...

Evidence of plant food processing is a significant indicator of the human ability to exploit environmental resources. The recovery of starch grains associated with use-wear on Palaeolithic grinding tools offers proof of a specific technology for making flour among Pleistocene hunter-gatherers. Here we present the analysis of five grindstones from t...

Modern humans have populated Europe for more than 45,000 years1,2. Our knowledge of the genetic relatedness and structure of ancient hunter-gatherers is however limited, owing to the scarceness and poor molecular preservation of human remains from that period³. Here we analyse 356 ancient hunter-gatherer genomes, including new genomic data for 116...

The mobility of hunter-gatherer groups is crucial in understanding Palaeolithic settlement dynamics. The concept of mobility cannot be separated from the space in which it occurs, including landscape components, localization of critical resources and of other sites, and routes between them. Nevertheless, the landscape is not constant in time due to...

In this paper we apply for the first time a 2D shape analysis to a sample of Epigravettian lithic artefacts with the aim of evaluating the potential of such approach. The lithic sample comes from layer 9c2 (Evolved Epigravettian, Upper Palaeolithic) of Grotta Paglicci (Apulia, southern Italy). After extracting the outline coordinates from high-reso...

The current paper analyses the faunal remains deriving from inhumations (1st century AD) and cremations (1st-3rd centuries AD, with the main period of occupation dating from the Claudian period to the middle 2nd century AD) excavated at Potzneusiedl (Burgenland, Austria). The inhumations are unique because of their early dating and the grave goods...

The human microbiome has recently become a valuable source of information about host life and health. To date little is known about how it may have evolved during key phases along our history, such as the Neolithic transition towards agriculture. Here, we shed light on the evolution experienced by the oral microbiome during this transition, compari...

Predicting the effects of future global changes on species requires a better understanding of the ecological niche dynamics in response to climate; the large climatic fluctuations of the last 50,000 years can be used as a natural experiment to that aim. Here we test whether the realized niche of horse, aurochs, red deer, and wild boar changed betwe...

Residue analysis in the study of Palaeolithic represents both a well-developed field of research and a discipline with a great deal of potential yet to be explored and understood. As proven by previous works (Sano et al. 2019; Dominici et al. 2022), the use of techniques based on Synchrotron Radiation (SR) in dealing with degraded traces of organic...

Bones and teeth are biological archives, but their structure and composition are subjected to alteration overtime due to biological and chemical degradation postmortem, influenced by burial environment and conditions. Nevertheless, organic fraction preservation is mandatory for several archeometric analyses and applications. The mutual protection b...

In the last few years several studies have reassessed the attraction and the role of small game in the subsistence economy of hunter-gatherers across Europe and the Mediterranean region since the Middle Paleolithic. The exploitation of small mammals intensified during the Pleistocene-Holocene transition, when some unusual faunal assemblages were re...

Neanderthal foot bone proportions and morphology are mostly indistinguishable from those of Homo sapiens, with the exception of several distinct Neanderthal features in the talus. The biomechanical implications of these distinct talar features remain contentious, fueling debate around the adaptive meaning of this distinctiveness. With the aim of cl...

THE DIET DURING THE PALAEOLITHIC: A CASE STUDY FROM GROTTA PAGLICCI (RIGNANO GARGANICO – FG) – It is usually assumed that the diet of Palaeolithic hunter-gatherer communities was based almost exclusively on the consumption of meat and of other animal resources (e.g. marrow). Recent studies carried out on dental calculus, as well as on organic resid...

The origin, development, and legacy of the enigmatic Etruscan civilization from the central region of the Italian peninsula known as Etruria have been debated for centuries. Here we report a genomic time transect of 82 individuals spanning almost two millennia (800 BCE to 1000 CE) across Etruria and southern Italy. During the Iron Age, we detect a...

After the last interglacial [Marine Isotope Stage (MIS) 5e] Europe was affected by several harsh climatic oscillations. In this context southern Italy acted, like the rest of peninsular Mediterranean Europe, as a 'glacial refugium', allowing the survival of various species, and was involved in the spread of 'cold taxa' (e.g. woolly mammoth and wool...

Background Recently, the study of mitochondrial variability in ancient humans has allowed the definition of population dynamics that characterised Europe in the Late Pleistocene and Early Holocene. Despite the abundance of sites and skeletal remains few data are available for Italy. Aim We reconstructed the mitochondrial genomes of three Upper Pal...

The analysis of bone-surface modifications (BSM), such as butchering marks, is necessary to better understand how the exploitation of animal resources by past hominins influenced their biological and cultural evolution. In this paper, we try to quantify to what extent the depth of the cut marks influences the shape of their cross sections. This is...

Fishing, hunting and gathering at the sauveterrian site Galgenbühel/Dos de la Forca (Salorno, South Tyrol) - The fauna recovered at the mesolithic site give a picture of the former environment and allows to reconstruct fishing, hunting and gathering strategies as well as the diet of the groups living in the Adige valley during the 9th-8th millenniu...

An understanding of how ecological niches can change through time is key to predicting the effect of future global change. Past climatic fluctuations provide a natural experiment to assess the extent to which species can change their niche. Here we use an extensive archaeological database to formally test whether the niche of four European ungulate...

Clastic successions in rock shelters commonly host important archaeological findings, especially of prehistoric and protostoric times. The understanding of depositional and post-depositional processes in these environments is crucial to understand the lifestyle settings of humans, as well as the reliability of archaeological data obtained during ex...

The Oscurusciuto rock shelter (southern Italy) is crucial for the understanding of Neanderthals’ subsistence and settlement strategies as it contains a ~ 6-m-thick reliable deposit made up of several Middle Palaeolithic levels. This paper focuses on level SU 14, a 60-cm-thick deposit of volcanic tephra containing traces of human occupation only in...

The role of small game in prehistoric hunter-gatherer economy is a highly debated topic. Despite the general assumption that this practice was uneconomic, several studies have underlined the relevance of the circumstance of capture – in terms of hunting strategies and technology – in the evaluation of the actual role of small mammals in human forag...

PurposeIn the past, a role of thyroid hormones in human evolution has been hypothesized. T3, the metabolically active form, derives from extrathyroidal conversion of T4 by deionidase 2 (D2) enzyme encoded by DIO2 gene. In thyroid-deficient patients, decreased levels of free T3 have been associated with the polymorphism rs225014 A/G in DIO2, which c...

The Molare Rock shelter (S. Giovanni a Piro, Salerno, Italy) is a key site to carry out high-resolution chronological studies in the broader context of Italian Mousterian peopling dynamics. The stratigraphic sequence is to be referred to MIS 5 and is characterized by the presence of a number of thin anthropic levels (often consisting of largely und...

Grotta dei Santi represents a very suitable investigation field for carrying out an inquiry into the Neandertal behaviour at a high-resolution time scale, in order to obtain data useful for the "functional" reading of the economic and settling strategies. This cave opens into the Monte Argentario Promontory (southern coast of Tuscany, central Italy...

Personal ornaments are commonly linked to the emergence of symbolic behavior. Although their presence in Africa dates back to the Middle Stone Age, evidence of ornament manufacturing in Eurasia are sporadically observed in Middle Palaeolithic contexts, and until now, large-scale diffusion has been well documented only since the Upper Palaeolithic....

Most of the Middle Palaeolithic evidence of Central Italy still lacks a reliable chrono-cultural framework mainly due to research history. In this context Grotta dei Santi, a wide cave located on Monte Argentario, on the southern coast of Tuscany, is particularly relevant as it contains a very well preserved sequence including several Mousterian la...

The Oscurusciuto Rockshelter (Ginosa, Southern Italy) is a perfect sample-site for the reconstruction of multiple aspects of the last Neanderthals life. Different settlement strategies are attested in the excavated portion of the stratigraphic sequence, dated between ~ 55 and 43 ka BP. As a first goal, the reconstruction of the site spatial organiz...

The arrival of Modern Humans (MHs) in Europe between 50 ka and 36 ka coincides with significant changes in human behaviour, regarding the production of tools, the exploitation of resources and the systematic use of ornaments and colouring substances. The emergence of the so-called modern behaviours is usually associated with MHs, although in these...

Defining the processes involved in the technical/cultural shifts from the Late Middle to the Early Upper Palaeolithic in Europe (~50-39 thousand years BP) is one of the most important tasks facing prehistoric studies. In this debate Italy plays a pivotal role, due to its geographical position between eastern and western Mediterranean Europe as well...

Evidence of human activities during the Middle to Upper Palaeolithic transition is well represented from rock_shelters, caves and open_air sites across Italy. Over the past decade, both the revision of taphonomic processes affecting archaeological faunal assemblages and new zooarchaeological studies have allowed archaeologists to better understand...

Defining the processes involved in the technical/cultural shifts from the Late Middle to the Early Upper Palaeolithic in Europe (~50-39 thousand years BP) is one of the most important tasks facing prehistoric studies. Apart from the technological diversity generally recognised as belonging to the latter part of the Middle Palaeolithic, some assembl...

The arrival of Modern Humans (MHs) in Europe between 50 ka and 39 ka coincides with significant changes in human behaviour, notably regarding the production of tools, the exploitation of resources and the systematic use of ornaments and colouring substances. The emergence of the so-called modern behaviour is usually associated with MHs, although cl...

Evidence of human activities during the Middle to Upper Palaeolithic transition is well represented from rock-shelters, caves and open-air sites across Italy. Over the past decade, both the revision of taphonomic processes affecting archaeological faunal assemblages and new zooarchaeological studies have allowed archaeologists to better understand...

Microscopic analysis of backed lithic pieces from the Uluzzian technocomplex (45–40 thousand yr ago) at Grotta del Cavallo (southern Italy) reveals their use as mechanically delivered projectile weapons, attributed to anatomically modern humans. Use-wear and residue analyses indicate that the lithics were hunting armatures hafted with complex adhes...

Grotta di Castelcivita (Campania, Southern Italy) is a cave-site containing a key archaeological sequence for the study of the Middle to Upper Palaeolithic transition in Southern Italy. A Late Mousterian occupation, whose modelled dates span from 47.6 to 41.1 ka cal BP, is followed by layers bearing evidence of the Uluzzian techno-complex, which un...

Understanding the reason(s) behind changes in human mobility strategies through space and time is a major challenge in palaeoanthropology. Most of the time this is due to the lack of suitable temporal sequences of human skeletal specimens during critical climatic or cultural shifts. Here, we present temporal variations in the Sr isotope composition...

Despite the widespread application of high-resolution quantitative methods in bone taphonomy, very few studies have focused on projectile impact marks. Therefore, in a previous work, we explored the potential of 3D microscopy in distinguishing bone hunting injuries from other taphonomic marks, developing a widely-applicable diagnostic framework bas...

The Middle Bronze Age site of Gorgo del Ciliegio is located in the Afra valley (Upper Tiber basin) in the proximity of strategic trans-Apennine passes connecting the Tyrrhenian regions of Central Italy to the Po plain and the Umbrian-Marches territories. At Gorgo del Ciliegio a Middle Bronze Age small settlement composed of a single house and vario...

The Mesolithic faunal record recovered from the Galgenbühel/Dos de la Forca rock-shelter at Salorno/Salurn (Bolzano/South Tyrol, Italy, Eastern Alps) allows the reconstruction of the landscape and local environmental conditions in the Adige valley bottom during the Early Holocene thanks to the analysis of the species ecology. The predominance of a...

Bone accumulation in Palaeolithic archaeological sites is often the result of activities carried out by hunter-gatherer groups. Cultural choices may have influenced prey representation in archaeological assemblages, distorting their palaeoecological meaning. We present a comparison between large mammal and small mammal assemblages from the Upper Pa...

The surface texture of a stone represents a sensitive parameter in evaluating its conservation state. In monuments and sculptures, in fact, external agents continuously alter the appearance of stones, determining peculiar weathering patterns and modifying properties such as retention of water and particles, interaction with light, color, and finish...

Stratigraphic Unit 13 of Oscurusciuto Rockshelter (Ginosa, Taranto, Southern Italy) is a short Mousterian palimpsest representing the first stable occupation of the site soon after the deposition of a thick layer of tephra (Mt. Epomeo Green Tuff - Ischia datable around 55 kya BP). Different activities were identified by integrating the study of lit...

The Uluzzian techno-complex is commonly considered to be a "transitional industry" mostly on the basis of some inferred characteristics such as a chiefly flake-based production, a small amount of Upper Palaeolithic-like tools and a combination of Middle and Upper Palaeolithic elements both in the toolkit and in the technical systems. Following its...

Wild horses thrived across Eurasia until the Last Glacial Maximum to collapse after the beginning of the Holocene. The interplay of climate change, species adaptability to different environments, and human domestication in horse history is still lacking coherent continental-scale analysis integrating different lines of evidence. We assembled tempor...

Despite the widespread application of high-resolution quantitative methods in bone taphonomy, very few studies have focused on projectile impact marks (PIMs). Therefore, in a previous work, we explored the potential of 3D microscopy in distinguishing bone hunting injuries from other taphonomic marks, developing a widely-applicable diagnostic framew...

The site of Ficoncella is located to the north of Rome (Central Italy). It is an alluvial context where animal bones and lithic remains were buried over a short space of time in a floodplain environment during the Lower Palaeolithic. The main faunal evidence is represented by parts of a straight-tusked elephant carcass, but remains belonging to oth...

La caccia alle marmotte sulle Prealpi italiane durante il Tardoglaciale: dati sperimentali e analisi morfometrica 3D di impatti di proiettile su resti faunistici Parole chiave: Tafonomia, Lesioni da impatto, Marmotta, Proiettili litici, Microscopia 3D, Epigravettiano Recente. Nonostante la sempre più vasta applicazione di metodi quantitativi ad alt...

Il sito pluristratificato all'aperto di Arco via Serafini (Trento), posto alla quota di 80 m s.l.m., è collocato sulla porzione distale del conoide colluviale del Rio Bordellino, in prossimità della pianura alluvionale del fiume Sarca. Le indagini archeologiche iniziate nel febbraio del 2013 in un'area già nota per il ritrovamento di importanti evi...

The site of Elvas-Kreuzwiese p. f. 574 is located near Brixen-Bressanone (BZ) on the NatzSchabs plateau at about 600 m a.s.l. Excavations carried out at the beginning of this century attested the presence of Late Bronze Age, Iron Age and Roman Age materials. Almost 10,000 animal remains from all chronological phases are presented in this paper. Fau...

The site of Elvas-Kreuzwiese p. f. 574 is located near Brixen-Bressanone (BZ) on the Natz-Schabs plateau at about 600 m a.s.l. Excavations carried out at the beginning of this century attested the presence of Late Bronze Age, Iron Age and Roman Age materials. Almost 10,000 animal remains from all chronological phases are presented in this paper. Fa...