Fernando E Novas

Fernando E Novas
Museo Argentino de Ciencias Naturales "Bernardino Rivadavia" · Laboratorio de Anatomía Comparada y Evolución de los Reptiles Fósiles

Doctor in Natural Sciences

About

228
Publications
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Introduction
Specific topics on which we are working are: 1) dinosaur origins based on current information from Argentina and Brazil; 2) phylogenetic relationships of particular theropod taxa (Chilesaurus and Megaraptoridae); 3) origin of birds and their flight, based on comparative studies on the anatomy of the Patagonian unenlagiids with other avialans, and observations on function and behaviour of some living, flightless birds (Ratitae).

Publications

Publications (228)
Article
Skorpiovenator bustingorryi is a derived abelisaurid theropod represented by a fairly complete skeleton from the Late Cretaceous sedimentary beds of north-western Patagonia. Although some features were described in the original paper, mainly related to the skull, the appendicular anatomy remains undescribed. The aim of the present contribution is t...
Article
Few latest Cretaceous fossil floras are known for South America, and in particular for the Austral-Magallanes Basin. Recent studies carried out in the Chorrillo Formation (Maastrichtian) revealed a diverse array of fossil elements, including continental invertebrates, vertebrates, palynomorphs, fossil woods and leaf impressions. In this work, we de...
Article
The Greater Rhea (Rhea americana, Rheidae) is a flightless paleognath with a wide geographical distribution in South America. The morphology of its shoulder girdle and wings are different from those of flying birds and some characteristics are similar to basal birds and paravian theropods. We present a detailed osteological, myological, and functio...
Article
Full-text available
Megaraptora is a theropod clade known from former Gondwana landmasses and Asia. Most members of the clade are known from the Early to Late Cretaceous (Barremian–Santonian), with Maastrichtian megaraptorans known only from isolated and poorly informative remains. The aim of the present contribution is to describe a partial skeleton of a megaraptorid...
Article
Dicraeosaurid sauropods are iconically characterized by the presence of elongate hemispinous processes in presacral vertebrae. These hemispinous processes can show an extreme degree of elongation, such as in the Argentinean forms Amargasaurus cazaui, Pilmatueia faundezi and Bajadasaurus pronuspinax. These hyperelongated hemispinous processes have b...
Article
The deposits of the Chorrillo Formation (Maastrichtian) were accumulated during a ‘continental window’ that occurred during the Late Cretaceous in the Austral-Magallanes foreland basin, southern Patagonia, Argentina. The aim of the present contribution is to describe the depositional conditions as well as new vertebrate and plant fossils from this...
Preprint
Full-text available
Megaraptorans are a theropod clade distributed in former Gondwana landmasses and Asia. Most members of the clade are known from early Cretaceous to Turonian times whereas Maastrichtian megaraptorans are known just from isolated and poorly informative remains. The aim of present contribution is to describe a partial skeleton of a megaraptorid coming...
Article
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The fossil record of Early Cretaceous birds in South America has been restricted to members of Enantiornithes from the Crato Formation (Aptian) of Brazil. Here we describe a new genus and species of bird discovered at Pedra Branca Mine, Nova Olinda County, Ceará State, Brazil, which adds to the avian fossil record from the Crato Formation. The spec...
Article
The record of sauropods in the Upper Cretaceous of Patagonia is rich. However, there are still several blanks on this record. Here it is described a new sauropod assemblage coming from the Salitral Ojo de Agua Area, Río Negro province, northern Patagonia, Argentina (Allen Formation; Maastrichtian). Remains of indeterminate saltasaurines, aeolosauri...
Article
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Lewisuchus admixtus is an early dinosauriform described by Alfred Romer in 1972 on the basis of a single, incomplete skeleton, collected in lower Upper Triassic rocks of the renowned Chañares Formation, at the Los Chañares type‐locality, La Rioja Province, north‐western Argentina. Recent field explorations to the type‐locality resulted in the disco...
Article
Aerosteon riocoloradensis represents one of the most complete megaraptorans yet discovered. This theropod comes from Anacleto Formation (Campanian) of Mendoza Province, Argentina. The aims of this contribution are: to present a detailed, bone by bone description of this specimen with figures of each bone; provide comparisons to other closely relate...
Article
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We describe the basal mesoeucrocodylian Burkesuchus mallingrandensis nov. gen. et sp., from th Upper Jurassic (Tithonian) Toqui Formation of southern Chile. The new taxon constitutes one of the few records of non-pelagic Jurassic crocodyliforms for the entire South American continent. Burkesuchus was found on the same levels that yielded titanosaur...
Article
The fossil record of gondwanatherian mammaliaforms from Patagonia is represented by several species known on the basis of isolated teeth, with the single exception of a partial dentary with two molariforms of the Paleocene species Sudamerica ameghinoi. The aim of the present contribution is to describe both a fragmentary dentary (with the base of t...
Article
Full-text available
In 1976 John Ostrom published an enlightening paper about the anatomical transformations in the shoulder girdle and forelimb elements along the origin of birds. Most of his ideas were based on comparing Archaeopteryx lithographica with the extant New World vulture Cathartes aura. Ostrom offered innovative ideas about range of movements and function...
Article
The fossil record of ornithischians in South America is sparse, and they are clearly underrepresented when compared with sauropod dinosaurs. However, recent discoveries indicate that ornithischians were more diversified than thought. The aim of the present contribution is to describe isolated remains belonging to ankylosaurs, and ornithopods, inclu...
Article
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Unenlagia comahuensis was originally described as a phylogenetic link between nonavian dinosaurs and birds. Later it was interpreted by some authors as belonging to the deinonychosaurian clade Dromaeosauridae, and more recently as phylogenetically closer to birds than to dromaeosaurids. The only known specimen is represented by an incomplete skelet...
Article
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Triassic beds from Argentina and Brazil provide the most relevant fossil record of early dinosauriforms in terms of numerical abundance and taxonomic diversity. This record currently represents the best source to understand the origin and early evolutionary radiation of dinosaurs. In the present paper we offer an updated review focused on the avail...
Article
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Pterosaurs were the first vertebrates to evolve powered flight¹ and comprised one of the main evolutionary radiations in terrestrial ecosystems of the Mesozoic era (approximately 252–66 million years ago), but their origin has remained an unresolved enigma in palaeontology since the nineteenth century2–4. These flying reptiles have been hypothesize...
Article
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Gondwanatheria is an enigmatic mammaliaform clade distributed in the Cretaceous and Paleogene of South America, Africa, Madagascar, India and Antarctica. The Mesozoic record in South America is restricted to the Latest Cretaceous of Río Negro and Chubut provinces, Argentina and Magallanes Region of southern Chile. The aim of the present contributio...
Article
Abelisaurid theropods are well known in Cretaceous beds along South America, particularly Patagonia. However, the record of latest Cretaceous abelisauroids is still far from satisfactory. Until recently, few taxa were described from Maastrichtian beds: Carnotaurus sastrei and Quilmesaurus curriei, but also perhaps Abelisaurus comahuensis. In this c...
Article
Full-text available
Discussions about the origin of flight almost unanimously assume that early birds positioned (and moved) their wings in the same basic manner as living flying birds, with reconstructed wings extended with the airfoil surface parallel to the ground and forelimbs moving in a dorsoventral arc. Such reconstructions of wing posture and movements for ext...
Article
Carnotaurus sastrei is perhaps the most iconic representative of the Abelisauridae family. It is known by a fairly complete specimen, recovered from Upper Cretaceous beds of southern Argentina. Here we present a re-study of the skull of Carnotaurus including detailed osteological descriptions, CT scans analyses and comparisons with abelisaurid taxa...
Chapter
Discussions about the origin of flight almost unanimously assume that early birds positioned (and moved) their wings in the same basic manner as living flying birds, with reconstructed wings extended with the airfoil surface parallel to the ground and forelimbs moving in a dorsoventral arc. Such reconstructions of wing posture and movements for ext...
Article
Abelisaurids were one of the most successful theropod dinosaurs during Cretaceous times. They are featured by numerous derived skull traits, such as heavily ornamented bones, short and tall snout, and a strongly thickened cranial roof. Furthermore, nasals are distinctive on having two distinct nasal patterns: strongly transversely convex and heavil...
Article
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The Chañares Formation exhibits one of the most important archosauriform records of early Carnian ecosystems. Here we present new data on the palaeohistology of Chañares archosauriforms and provide new insights into their paleobiology, as well as possible phylogenetically informative traits. Bone microstructure of Lagerpeton chanarensis and Tropido...
Article
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Mosasaurs were a cosmopolitan group of marine squamate reptiles that lived during the Late Cretaceous period. Tylosaurinae mosasaurs were characterized for having an edentulous rostrum anterior to the premaxillary teeth. External morphology of the snout of the tylosaurine Taniwhasaurus antarcticus from the Upper Cretaceous beds at James Ross Island...
Article
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Aoniraptor libertatem is a mid‐sized megaraptoran that comes from the Late Cretaceous (Turonian) Huincul Formation at Río Negro province, Patagonia, Argentina. In this study, we conducted a detailed analysis of pneumaticity of the sacrum and tail of Aoniraptor . This shows a complex structure within these vertebrae, being composed by small divertic...
Article
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The fossil record of basal paravians in Gondwana is still poorly known, being limited to the Cretaceous unenlagiids from South America and the problematic Rahonavis from Madagascar. Here we report on a new paravian from the Cenomanian-Turonian (Late Cretaceous) of Río Negro province, NW Patagonia, Argentina. The new taxon exhibits a derived bird-li...
Article
Full-text available
The first fossil remains of vertebrates, invertebrates, plants and palynomorphs of the Chorrillo Formation (Austral Basin), about 30km to the SW of the town of El Calafate (Province of Santa Cruz), are described. Fossils include the elasmarian (basal Iguanodontia) Isasicursor santacrucensis gen. et sp. nov., the large titanosaur Nullotitan glaciari...
Data
The present 3D Dataset contains the 3D models analyzed in the following publication: Paulina-Carabajal, A., Ezcurra, M., Novas, F., 2019. New information on the braincase and endocranial morphology of the Late Triassic neotheropod Zupaysaurus rougieri using Computed Tomography data. Journal of Vertebrate Paleontology. https://doi.org/10.1080/027246...
Conference Paper
Full-text available
En aves voladoras actuales el músculo supracoracoides recorre hacia arriba la cara anteromedial del coracoides, rodea al proceso acrocoracoidal y atraviesa el foramen trióseo para finalmente insertarse sobre la cara dorsal del húmero. Su función es clave en la elevación del ala y la rotación longitudinal del húmero. John Ostrom describió la condici...
Article
Zupaysaurus rougieri is an early neotheropod from the middle Norian Los Colorados Formation of northwestern Argentina represented by an almost complete skull and several postcranial bones. Most of its braincase morphology has remained obscured by other skull bones and sediment. Additional mechanical preparation and X-ray computed tomography on the...
Conference Paper
Full-text available
La presente contribución se centra en la histología ósea de dos ornitópodos antárticos de la Formación Snow Hill Island (Maastrichtiano-Campaniano), Trinisaura santamartaensis y Morrosaurus antarcticus. El objetivo es establecer el estadio ontogenético de ambos taxones, así como determinar si su crecimiento presenta diferencias significativas con f...
Conference Paper
Full-text available
Paleontological exploration has a long history in Argentina. The rich and diverse Argentinian fossil record has proven to be of upmost importance, profusely contributing to the resolution of important evolutionary interrogations. The technological advances, in particular those regarding three-dimensional tomographic imaging, opened a new field of r...
Article
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Talenkauen santacrucensis represents one of the most complete South American ornithopods yet discovered. This dinosaur comes from the Mata Amarilla Formation (Turonian) of Santa Cruz Province, Argentina. The aim of this contribution is to present a detailed description of Talenkauen santacrucensis. Features of the cervical series of Talenkauen, whi...
Article
Phylogenetic relationships of megaraptorid theropods are under intense debate. Some authors interpret them as archaic allosauroids that survived up to Late Cretaceous, whereas others consider megaraptorids as basal tyrannosauroids. The recently described Patagonian taxon Murusraptor barrosaensis offers novel information on skull, axial and hind lim...
Article
Pipimorpha is a clade of tongueless anurans with a wide fossil record. Furthermore, the oldest South American fossils come from the Late Cretaceous (Cenomanian) of Patagonia, Argentina. The aim of the present contribution is to describe a new genus and species of Pipimorpha from the Crato Formation (Aptian, Early Cretaceous), Araripe Basin, Brazil....
Article
Full-text available
Recent years witnessed the discovery of a great diversity of early birds as well as closely related non-avian theropods, which modified previous conceptions about the origin of birds and their flight. We here present a review of the taxonomic composition and main anatomical characteristics of those theropod families closely related with early birds...
Article
Full-text available
Many dinosaur skeletons show evidence of behavior, including feeding, predation, nesting, and parental care. The resting posture of the forelimbs has been studied in some theropod species, in relation to the acquisition of flight in advanced maniraptoran theropods. Chilesaurus diegosuarezi is a bizarre tetanuran recently described from the Toqui Fo...
Article
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The fossil record of Late Cretaceous–Paleogene modern birds in the Southern Hemisphere includes the Maastrichtian Neogaeornis wetzeli from Chile, Polarornis gregorii and Vegavis iaai from Antarctica, and Australornis lovei from the Paleogene of New Zealand. The recent finding of a new and nearly complete Vegavis skeleton constitutes the most inform...
Article
Neuquensuchus universitas is an enigmatic crocodyliform known by a single, incomplete postcranial skeleton collected in the Upper Cretaceous (Santonian) Bajo de la Carpa Formation, Neuquén province, Patagonia, Argentina. Here we describe a second specimen of N. universitas that adds important features related to the neck and limb anatomy. The new s...
Article
Tail anatomy of unenlagiids is nearly unknown. The aim of the present contribution is to describe in detail the tail of the unenlagiid Buitreraptor gonzalezorum, and to compare it with Archaeopteryx and other paravians. A detailed specimen overview yielded a large number of similarities between Archaeopteryx and Buitreraptor, including long postzyg...
Article
Full-text available
Vegavis iaai is a neornithine bird coming from the Late Cretaceous Sandwich Bluff Member of the López de Bertodano Formation (Maastrichtian), Antarctic Peninsula. Vegavis constitutes the only unquestionable Cretaceous neornithine bird, and is known by the holotype and specimen MACN-PV 19.748. The goal of this paper is to present a detailed osteohis...
Article
Full-text available
Lewisuchus admixtus is a basal dinosauriform coming from Late Triassic outcrops of NW Argentina. Although this taxon was recently anatomically restudied, histological data is still wanting. The microstructure of the long bones (femur and two tibie) reveals a relatively fast rate of growth, comparable with that seen in other basal Dinosauriformes an...
Article
Full-text available
Neuquenraptor argentinus was described as the first undoubted deinonychosaurian theropod from Gondwana. The only known specimen is represented by a fragmentary skeleton, including a nearly complete foot, coming from Late Cretaceous beds of Neuquén Province, Patagonia, Argentina. Neuquenraptor was later considered as belonging to the Southern Hemisp...
Article
Buitreraptor gonzalezorum is a theropod dinosaur belonging to the clade Unenlagiidae. It is known by several specimens coming from the Upper Cretaceous of Northwestern Patagonia, Argentina. In spite of its completeness, only a preliminary description of its postcranial anatomy is available. The aim of the present contribution is to make a detailed...
Article
We describe isolated shark teeth collected from levels of the Calafate Formation at the SE coast of the Argentino Lake, Calafate city, Santa Cruz province, Argentina (Atlantic Ocean), and from the Algarrobo coast at the Valparaíso Region in central Chile (Pacific Ocean). The teeth belong to a new species of the echinorhiniform genus Echinorhinus. E...
Article
A new genus and species of panchelid turtle (Mendozachelys wichmanni gen. et sp. nov.) from the upper Campanian–lower Maastrichtian Loncoche Formation (southern Mendoza Province, Argentina) is described here. This species is represented by only one complete and articulated large specimen (carapace length estimated in 950 mm) that was recovered from...
Article
Full-text available
http://zoobank.org/urn:lsid:zoobank.org:pub:69F156B4-1A84-4C98-86C3-4FE4626024AC Citation for this article: Agnolín, F. L., S. Bogan, F. Brissón Egli, F. E. Novas, M. P. Isasi, C. Marsicano, A. Zavattieri, and A. Mancuso. 2016. A new lungfish (Dipnoi) from the Late Triassic of South America. Journal of Vertebrate Paleontology. DOI: 10.1080/02724634...
Article
Full-text available
From complex songs to simple honks, birds produce sounds using a unique vocal organ called the syrinx. Located close to the heart at the tracheobronchial junction, vocal folds or membranes attached to modified mineralized rings vibrate to produce sound. Syringeal components were not thought to commonly enter the fossil record, and the few reported...
Article
Full-text available
Novas, F.E., Aranciaga Rolando, A.M. and Agnolín, F.L. 2016. Phylogenetic relationships of the Cretaceous Gondwanan theropods Megaraptor and Australovenator: the evidence afforded by their manual anatomy. Memoirs of Museum Victoria 74: 49–61. General comparisons of the manual elements of megaraptorid theropods are conducted with the aim to enlarge...