Félix Pérez-Lorente

• PhD
• Retired at University of La Rioja

The Iouaridène Formation (Upper Jurassic, High Atlas, Morocco) is a relevant area for the study of dinosaur footprints, but up to now little is known about the palaeoenvironmental conditions at the time of its formation. This work consists of a sedimentological, geochemical, and mineralogical study of part of the sedimentary succession of the Iouar...

In the ensemble of theropod footprints from the Imilchil macrosite, a group of small size separated from the rest is distinguished. We had detected this grouping in other sites studied by our team in La Rioja and in the work of drawing up scatter plots and frequency histograms of the measurements, on a global scale, of the length and width of the t...

Three types of ichnites from a new site discovered in Imilchil were studied and described. The structures of pes and manus prints and tail traces of tetrapods provide data to separate: real ichnites (theropod footprints and crocodile tail drag marks); and sauropod’ subtraces. Most of the theropod footprints are deformed by collapse of their walls,...

This paper presents the study of 14 new sites with theropod footprints in Imilchil

This paper presents the study of 14 new sites with theropod footprints in Imilchil. The study focuses on ichnotaxonomic and extramorphological characteristics of the footprints, the analysis of their trackways, and the correlation with the descriptions of published tridactil footprints. The objective has been to provide a comparative view of therop...

The Sierra de las Cabras (Lat. 38° 28′ 53″N, Lon. 1° 24′ 52″W) is an exceptional ichnofossil site of late Miocene age, located in the Prebetic ranges near the town of Jumilla (Murcia Province, SE Spain). The site contains abundant vertebrate ichnofauna preserved in carbonate facies that were deposited in a semi-arid wetland system with shallow pond...

Upper Miocene sedimentary deposits of Sierra de las Cabras in the Prebetic Ranges (Jumilla, Región de Murcia, Spain) yield a exceptional fossil site of diverse vertebrate ichnofauna, including mammals and birds. This paper presents a detailed study of those ichnites of that site which correspond to mammals. The footprints are located in stratigraph...

Three types of ichnites from a new site discovered in Imilchil were studied and described. The structures of pes and manus prints and tail traces of tetrapods provide data to separate: real ichnites (theropod footprints and crocodile tail drag marks); and sauropod' sub traces. Most of the theropod footprints are deformed by the collapse of their wa...

En El Cantalar (Galve, Teruel) dentro de la Formación Villar del Arzobispo (Jurásico Superior-Cretácico Inferior), se conocían dos rastrilladas: una de un enorme cocodrilo y otra de un terópodo de andar semiplantígrado. El lavado y erosión del afloramiento ha limpiado la parte conocida y, además, ha puesto al descubierto varias pisadas que añadir a...

https://www.tandfonline.com/doi/full/10.1080/10420940.2020.1835661 Abstract During a study of the Talmest ichnological site (2011–2012), where theropod and sauropod tracks were discovered, other ichnites were found that are analyzed in the present work. There are two basic types attributed to two different activities produced by the same vertebrate...

In this work we present the description and analysis of a palaeoichnological site (7.9TAG) that only contains crocodile tracks. The site contains well preserved and abundant crocodilian manus and pes prints; there are no belly and/or tail traces. The crocodyle tracks have been mapped and the biomorphic and morphometric characters of the tracks and...

The present study evaluates a methodological workflow that could identify dinosaur tracks and trackways more comprehensively at outcrop scale. The approach described here is based both on 3D modelling by photogrammetry at different resolutions, and on suitably processed digital elevation models (DEMs). The ichnosite of Anza, Morocco, was chosen to...

The Anza tracksite, discovered in 2013, is one of several sites in the Western High Atlas of Morocco that contain well-preserved and diverse dinosaur and pterosaur trackways. The occurrence of rare tracks at Anza combined with the stratigraphic position, coastal location, and easy accessibility, have inspired the scientific community, non-governmen...

In this paper the study of dinosaur tracks of La Rioja, which began in 1970, is presented as an example of knowledge development, achievements, and events occurring around a paleontological resource having strong media impact. Fieldwork, research, and dissemination since the publication of the first scientific works have been accompanied by activit...

Two morphotypes (first and second) of sauropod ichnites in the Imilchil sites have been described for the first time. The second morphotype has its own characters not defined in any other site. If each ichnotype has to be defined by a certain number of characters that differentiate it from the others, the classification of the sauropod ichnites is...

El aumento de estudios y publicaciones pone de manifiesto el valor natural, cultural y científico de las huellas de dinosaurio de Marruecos. Más de 3.000 huellas inventariadas y un número desconocido en los yacimientos todavía inexplorados componen la riqueza patrimonial del país norteafricano, con más de 27 lugares con huellas ubicados mayoritaria...

Cretaceous pterosaurs tracksites are very rare worldwide. Until now,only one African Cretaceous site withtracks of (Agadirichnus elegans and Pteraichnus) was known. This makes the discovery of a new outcrop in the Upper Cretaceous of Anza (Morocco) the third manifestation of this type of footprint in Africa, extending the existence of such traces f...

In this paper a new Upper Cretaceous site with sauropod footprints is described. The outcrop, found in Chott Tigri, in the meridional area of the High Plateau of Morocco, between the Saharan Atlas and the Middle Atlas, is separate from other known sites with dinosaur footprints. We have found: three trackways of titanosauriform features and variabl...

Introduction This work is a detailed description of ankylosaurian natural casts found in Galve (Teruel, Spain), with the identification and classification of the major dynamic and static structures that depend on the phase of autopodial (hand) movement. Geological setting Specimens come from the Lower Cretaceous (Barremian) of the eastern sector o...

The discovery of the Larañe (La Rioja, Spain) ichnological site provides the first outcrop pterosaur footprints in the Tera Group of the Cameros Basin. The footprint have characteristics similar to some of those already known in the Oncala Group, suggesting that possibly pterosaurs, which lived in this area during the deposition of the Tithonian–Be...

Abstract The Anza site shows large ichnological surfaces indicating the coexistence in the same area of different vertebrate footprints (dinosaur and pterosaur) and of different types (tridactyl and tetradactyl, semiplantigrade and rounded without digit marks) and the footprint variability of long trackways. This area may become a world reference i...

The Anza site shows large ichnological surfaces indicating the coexistence in the same area of different vertebrate footprints (dinosaur and pterosaur) and of different types (tridactyl and tetradactyl, semiplantigrade and rounded without digit marks) and the footprint variability of long trackways. This area may become a world reference in ichnolo...

Full text here: http://sepaleontologia.es/wp-content/uploads/2017/02/5-Moreno-Azanza-et-al-ok.pdf. Here we describe the new fossil site of El Horcajo in the vicinity of Trevijano (La Rioja, Spain). It is located in the palustrine facies of the Enciso Group (Cameros Basin). This new locality has provided dozens of eggshell fragments, together with...

Sur l’estran de la petite commune d’Anza, situé au nord de la ville d’Agadir, se trouve un gisement d’empreintes de dinosaures. Le site a été découvert en 2013, grâce aux membres de l’association marocaine de l’orientation et de la recherche scientifique (AMORS) et des blogueurs. L’étude préliminaire réalisée (Masrour M. et Perez-Lorente F. 2015) m...

Galve (Teruel, Spain) is a town in the interior of a synclinal fold with Upper Jurassic marine limestones along its flanks, and, in its core, Upper Jurassic–Lower Cretaceous continental and shoreline sediments crop out. The core sediments cover an area about 8 km², and contain a concentration of sites with footprints, bones, and eggshells of dinosa...

Small shorebird-like footprints have been discovered in Cenicero (La Rioja, Spain). They are preserved in a sandstone block of the transition unit between the Na´jera and Haro Formations from the Ebro Basin. This level is positioned in the Y2 local zone (MN2), of Agenian age (early Miocene). The footprints are small, tridactyl or tetradactyl, with...

Le site d’empreintes de dinosaures d’Anza au nord de la ville d’Agadir ; se situe sur un estran et de ce fait le gisement est couvert pendant les marrées hautes. Le site offre deux affleurements distants d’environ 10 mètres. C’est l’affleurement situé au Nord Ouest du site qui montre un très grand nombre d’empreintes à la surface de niveaux peu épa...

A new dinosaur tracksite (La Rueda) with ten small tridactyl footprints (the length ranges between 9 and 15 cm) from the Urbión Group (Cameros Basin, Lower Cretaceous, La Rioja, Spain) is described. The footprints are approximately as long as wide and have high divarication angles between digits II-IV (~80º), some pad impressions on each digit, cla...

http://journals.plos.org/plosone/article?id=10.1371/journal.pone.0115477 Background Large ornithopod tracks are known from the Upper Jurassic to the uppermost Cretaceous rocks of all continents but Antarctica. They include the tracks historically called Iguanodon footprints, iguanodontid footprints, hadrosaur/hadrosaurid footprints, and other lar...

During the Early Cretaceous, lakes, meandering streams, and flood plains covered the region where the current foothills of Rioja now exist. Today the area is known for its wine and for the dozens of sites where footprints and trackways of dinosaurs, amphibians, and even pterosaurs can be seen. The dinosaurs that lived here 120 million years ago lef...

Recientemente se ha descubierto un nuevo bloque con huellas fósiles de aves (14CN) en la localidad de Cenicero (La Rioja, España). Se conservan en el techo de un bloque de arenisca de la unidad de transición entre las formaciones Nájera y Haro de la Cuenca del Ebro y de edad Ageniense (Mioceno temprano). Las huellas son muy pequeñas (17-35 mm de lo...

A new site has been found in the Central High Atlas, in the Issil-n-Aït Arbi near the Dadés Gorges. This paper is the first study presented of this local-ity, with the description of a quadruped trackway of pentadactyl pes and tri-dactyl manus, which we have attributed to Otozoum. The characters of these prints and trackway are described, compared,...

A new site has been found in the Central High Atlas, in the Issil-n-Aït Arbi near the Dadés Gorges. This paper is the first study presented of this local-ity, with the description of a quadruped trackway of pentadactyl pes and tri-dactyl manus, which we have attributed to Otozoum. The characters of these prints and trackway are described, compared,...

This article describes the sedimentological characteristics of dinosaur ichnites at three sites within the municipal area of El Castellar (in the Province of Teruel, Aragón, Spain): El Castellar (CT-1), El Pozo (CT-2), and Camino El Berzal (CT-3). These sites possess large concentrations of footprints made by quadrupedal (sauropod, stegosaurid and...

In La Rioja the widespread ichnological record of dinosaurs ranges from the Berriasian to the Aptian age (Early Cretaceous). There, due to the high palaeodiversity, several evidences of gregarious behaviour can be observed in different types of footprints (sauropods, theropods and ornithopods). Moreover, due to the huge dimensions of some sites, di...

In this paper thyreophoran and theropod footprints contained in two of four new sites found in Ansous are described. In one of the sites there are also preserved sauropod tracks. Although there have been cited three other references to thyreophoran footprints in Morocco, this is the first of such quadrupedal footprints from the Lower Jurassic age (...

The Cameros Basin is a privileged place to study dinosaur footprints. It is estimated to contain more than 250 tracksites, although the total number depends on if the tracksites with a single footprint, the missing ones, the inventoried, cataloged or considered the BIC are counted (Pérez-Lorente, 2003). The Cameros Basin is traditionally divided in...

The timing of development of the magnetic fabric is a major issue in the application of anisotropy of magnetic susceptibility (AMS) as a strain marker. Analysis of AMS in unconcealed synsedimentary structures can be a sound approximation to this task. In this work, three types of early compactional structures (ECS) were studied by means of AMS, sin...

Se completa con este trabajo la cartografía del cuaternario de La Rioja en la Depresión del Ebro, ya iniciada en 1979, y se extraen algunas conclusiones de la distribución, evolución e interacción entre los diferentes niveles de erosión y depósito recientes.

Eight vertebrate trackways were found near the Imi-n-Tanout village, in the proximity of the two Marrakesh-Agadir roads: three theropod, two sauropods trackways; and three of an unknown bipedal trackmaker. The outcrop is the first Early Cretaceous dinosaur tracksite found in Morocco and the second (Barremian-early Aptian) site of Africa. This preli...

A new bird ichnotaxon found in Cenicero (La Rioja, Ebro Basin, Spain) is described here. The footprints are preserved in sandstone beds in a central-distal alluvial fringe with a mud-dominated floodplain, located in the transition unit between the Nájera and Haro formations. This level is positioned between the Y and Z local Agenian biozones (lower...

Son numerosos los aspectos del comportamiento de los dinosaurios que se han estudiado en los últimos años y que aún continúan interesando a la comunidad científica. Uno de los más polémicos es el comportamiento social (social behaviour), utilizado en muchos casos como sinónimo de comportamiento de grupo o gregario (Lucas, 2007). Según Currie & Eber...

La Cuenca de Cameros es un territorio con abundante registro de icnitas de dinosaurio en sedimentos de edades que comprenden desde el Berriasiense al Albiense basal (Cretácico Inferior). El sector riojano de dicha cuenca es la zona con mayor número de icnitas encontradas y estudiadas de España, así como una de las más importantes del registro fósil...

LOWER MIOCENE BIRD AND MAMMAL FOOTPRINTS FROM LA RIOJA (EBRO BASIN, SPAIN). A new site carrying vertebrate footprints is described at Cenicero (Ebro Basin, Spain). The footprints are preserved in sandstone beds located within the “Transition Unit”, between the Agenian (early Miocene) Nájera and Haro formations. This level is placed between local bi...

Weems parameter. Abbreviations: see Materials and Methods. (TIF)

Tables with data from all trackways. Abbreviations: see Materials and Methods. (DOC)

Measurements of the footprints and trackways. Abbreviations: see Materials and Methods. (TIF)

Trackways with all the footprints tetradactyls. (TIF)

New tetradactyl theropod footprints from Upper Jurassic (Oxfordian-Kimmeridgian) have been found in the Iouaridène syncline (Morocco). The tracksites are at several layers in the intermediate lacustrine unit of Iouaridène Formation. The footprints were named informally in previous works "Eutynichnium atlasipodus". We consider as nomen nudum. Boutak...

A study of the undescribed part of the Las Cerradicas tracksite in Galve (Teruel, Spain) has revealed 6 trackways made by small sauropods showing gregarious behaviour (parallel trackways). Las Cerradicas is located towards the top of the Villar del Arzobispo Formation (early Berriasian). The wide-gauge condition, the manus–pes distance, the kidney-...

Resumen: La participación activa de España en diferentes proyectos internacionales de Geoconservación, así como la ratificación de normativas internacionales que aconsejan la protección de la Geodiversidad, entran en contradicción con la ausencia de regulación de la preservación de elementos geológicos en la legislación vigente sobre Protección del...

Resumen Una de las características del sinclinal de Iouaridène, situado en el Alto Atlas Central, en la provincia de Azilal (unos 100 km al este de Marrakech) es la gran cantidad de yacimientos con icnitas de dinosaurio. En el miembro inferior de la Formación de Iouaridène se han inventariado más de 40 yacimientos (Boutakiout et al., 2009b) de edad...

El pueblo de Enciso alberga 18 yacimientos estudiados con icnitas: La Gilera, Corral de Valdefuentes, Navalsaz, Barranco de Valdeño, Barranco de Valdecevillo, Valdecevillo Este, La Virgen del Campo, El Barranco de la Sierra del Palo, Barranco de Valdegutierrez, Totico, Del Rio, La Cuesta de Andorra, El Villar-Poyales, La Senoba, Las Losas, Corral d...

Los campos de trabajo y cursos universitarios que se desarrollan en La Rioja desde hace 30 años han supuesto un importante impulso para muchas actividades relacionadas con las huellas de dinosaurio realizadas en la región. Se han excavado y estudiado más de 140 yacimientos, y se han rehabilitado varios de ellos. Se han construido también varios cen...

Lorente, F. 2009. Rastrilladas de icnitas terópodas gigantes del Jurásico Superior (Sinclinal de Iouaridène, Marruecos). [Gigantic theropod footprints from Upper Jurassic trackways (Iouaridène syncline, Morocco).] Revista Española de Paleontología, 24 (1), 31-46. ISSN 0213-6937. ABSTRACT The dinosaur ichnological research in the Upper Jurassic lacu...

In the proximity of Jumilla (Murcia, Spain) there is a site called Hoya de la Sima with mammal footprints dating from the Upper Miocene. This site is an old gypsum quarry where footprints of Hipparion, Pecoripeda, Carnivora, Ursidae, and Proboscidea were discovered. While the site was being cleaned and conditioned after the discovery to protect the...

Recibido el 9 de enero de 2008, aceptado el 6 de noviembre de 2008 Resumen La clasificación es uno de los principales problemas del estudio de huellas fósiles de dinosaurio. La forma de las icni-tas está condicionada principalmente por: el tipo de dinosaurio y su comportamiento (causas intrínsecas); las propiedades físicas del substrato, las interf...

The dinosaur ichnological research in the Upper Jurassic lacustrine sediments of the Iouaridène syncline provide data of interest. One of them includes the discovery of gigantic digitigrade theropod footprints, with a not des-cribed size. The exposures of ichnitic levels are continuous in the zone and the continuity of stratigraphic beds permits th...

A fundamental question remaining unanswered in dinosaur behavior is whether they had the ability to swim. We report the discovery of an exceptional swimming dinosaur trackway, with 12 consecutive footprints, in lacustrine nearshore sediment from the Early Cretaceous Cameros Basin, La Rioja, Spain. The singular morphology of these footprints strongl...

Cabezon de Cameros is a well known place for paleoichnological workers because one of the first described iguanodontid cuadrupedal trakways was found there. Up to now, this was the only site known in this place. In this paper we describe five new sites found near the one mentioned above. They not only widen the paleoichnological record of the regio...

In 1BO, one of the outcrops of Bin el Ouidane sites, there are theropod footprints with narrow toemarks that are small in size. The prints, even those from the same trackway, are formed in two superposed levels (from an earlier and later time) of finely laminated sandstones separated by some three centimeters of sediment. The upper level cannot be...

RESUMEN Se describen dos nuevos lugares con huellas en el término de Enciso, que se engloban en el yacimientos del Corral del Totico. Se han llamado 1TT y 2TT según las normas de nomenclatura seguidas en la Fundación Patrimonio Paleontológico de La Rioja. Son dos afloramientos distintos tanto por su contenido icnítico como por su extensión y propie...

The Las Losas site (approximately 500 m2) was cleared in July 1998 and revealed 375 theropod dinosaur footprints. Statistical surveys of their biomorphical and morphometrical data are congruent with a continuous morphological transition among the theropod footprints--a transition which does not allow us to differentiate ichnogroups.Certain footprin...

En la Cuenca de Cameros se han encontrado 156 yacimientos con icnitas de dinosaurio, repartidas en un registro estratigráfico que comprende parte del Jurásico superior (Kimmeridgiense-Titónico) y Cretácico inferior (Berriasiense-Albiense). Se analiza por una parte la distribución geográfica y por Grupos litoestratigráficos de los yacimientos, y por...

Se han analizado las icnitas terópodas de La Rioja de dos formas: aisladamente cada una de ellas una a una, y en la relación con las asociaciones de huellas que hay en los afloramientos en los que se encuentran. Al aplicar los criterios para su reconocimiento se observa que algunas icnitas se ajustan a las normas de identificación establecidas mien...

Calderón (1886) und Sánchez Lozano (1894) haben die ersten allgemeinen Arbeiten über die Geologie der Rioja veröffentlicht. Sie beziehen sich auf eine Reise, die J.U. Pereda und I. Aguirre y Muniain auf der Suche nach Mineralen unternommen hatten. Davon existiert noch ein Manuskript aus dem Jahr 1783. Weil Berichte über Bergbau-Tätigkeit bis 1536 z...

In southern Spain, within the Chiclana de Segura Formation of red detrital fades, enigmatic structures were found whose origins are discussed here. Each structure is composed of the union of elongated traces, two laterals and a shorter central one. All are concave epireliefs at the top of a sandstone bed whose age is assumed to be Ladinian (Triassi...

Palabras clave: Pisadas fósiles, Mamíferos, Mioceno superior, Jumilla, Murcia. Resumen: En la cuenca terciaria de Jumilla-Ontur (Murcia) se han encontrado huellas de mamífero fosilizadas en una cantera de yesos. El estudio de las mismas ha permitido distin-guir icnitas de Hippipeda y Megapecoripeda dejadas por grupos de équidos y de pecorinos, de c...

In this work we study part of the thrust between the Iberian Chain and the Ebro Basin. We applied criteria wich allow us to contribute arguments to the discussion about the mechanism and kinematics of the mentioned thrust. The conclusion wich we have reached seem to indicate that the tectonic instability is reflected only in the compressive phases...