Fabien Knoll

Fabien Knoll
ARAID · Fundación Conjunto Paleontológico de Teruel

PhD

About

160
Publications
29,086
Reads
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Introduction
Fabien Knoll is an ARAID Senior Researcher at the "Fundación Conjunto Paleontológico de Teruel" in Spain. He received his PhD in vertebrate palaeontology from the Muséum national d’Histoire naturelle, Paris. His research interests focus on the palaeobiology and evolution of archosaurs.
Additional affiliations
January 2017 - present
Teruel Palaeontological Complex Foundation
Position
  • Researcher
January 2014 - December 2016
University of Manchester
Position
  • Researcher
January 2009 - December 2014
National Museum of Natural Sciences
Position
  • Ramón y Cajal Researcher

Publications

Publications (160)
Article
Full-text available
The end-Triassic mass extinction events mark a pivotal period in archosaur history, and have been proposed to contribute to the rise and dominance of dinosaurs throughout the Mesozoic. In southern Africa, the Triassic–Jurassic boundary is contained within the richly fossiliferous fluvio-lacustrine-aeolian deposits of the upper Stormberg Group in th...
Article
The cover image is based on the Research Article Palaeoneurology of the early cretaceous iguanodont Proa valdearinnoensis and its bearing on the parallel developments of cognitive abilities in theropod and ornithopod dinosaurs by Fabien Knoll et al., https://doi.org/10.1002/cne.25224.
Article
Proa valdearinnoensis is a relatively large‐headed and stocky iguanodontian dinosaur from the latest Early Cretaceous of Spain. Its braincase is known from three specimens. Similar to that of other dinosaurs, it shows a mosaic ossification pattern in which most of the bones seem to have fused together indistinguishably while a few (frontoparietal,...
Article
The upper Stormberg Group of Lesotho has an abundant and diverse ichnological record that dates to the Early Jurassic. Herein, we re-evaluate a known ichnosite in the lowermost Clarens Formation (Karoo Supergroup) at Tsikoane in northern Lesotho. The site was reported to preserve tridactyl tracks as natural casts in the ceiling of overhangs and tru...
Article
Full-text available
Narambuenatitan palomoi is a titanosaurian sauropod from the Upper Cretaceous of North Patagonia. Considered initially as a basal titanosaur, this taxon has uncertain phylogenetic relationships within the clade. An X-ray Computed Tomography (CT) scan of the holotype braincase allowed the first reconstruction of the brain and inner ear of this taxon...
Article
In birds, the brain (especially the telencephalon) is remarkably developed, both in relative volume and complexity. Unlike in most early‐branching sauropsids, the adults of birds and other archosaurs have a well‐ossified neurocranium. In contrast to the situation in most of their reptilian relatives but similar to what can be seen in mammals, the b...
Article
Agaresuchus fontisensis is an allodaposuchid crocodile from the Campanian–Maastrichtian (Late Cretaceous) of Lo Hueco (Cuenca, Spain). Allodaposuchidae is a clade of European basal eusuchians, which is considered a part of the stem group of Crocodylia. The holotype and paratype skulls of Agaresuchus fontisensis were scanned on computed tomography a...
Article
Full-text available
Despite continuous improvements, our knowledge of the palaeoneurology of sauropod dinosaurs is still deficient. This holds true even for Titanosauria, which is a particularly speciose clade of sauropods with representatives known from numerous Cretaceous sites in many countries on all continents. The data currently available regarding the palaeoneu...
Article
Full-text available
Recent extensive field prospecting conducted in the Upper Miocene of Lebanon resulted in the discovery of several new fossiliferous localities. One of these, situated in the Zahleh area (Bekaa Valley, central Lebanon) has yielded a particularly diverse vertebrate fauna. Micromammals constitute an important part of this assemblage because not only d...
Article
Full-text available
Recent progress has been made in paleontology with respect to resolving pigmentation in fossil material. Morphological identification of fossilized melanosomes has been one approach, while a second methodology using chemical imaging and spectroscopy has also provided critical information particularly concerning eumelanin (black pigment) residue. In...
Article
Full-text available
The genus Sivameryx (Cetartiodactyla: Anthracotheriidae), found in both Asia and Africa, is considered of Asian origin. Recent excavations in the Negev region of southern Israel led to the discovery of a new early Miocene site called Kamus Junction. Among the fossils recovered at Kamus Junction is an upper molar of Sivameryx palaeindicus. Although...
Article
A well-preserved braincase of the basal alligatoroid Diplocynodon tormis was found in the middle Eocene site of ‘Teso de la Flecha,’ Salamanca, Spain. The specimen was scanned using computed tomography (CT), and its inner cavities were digitally rendered in three dimensions. Most bones of the left side are missing, so the reconstruction was based m...
Article
Despite continuous improvements, our knowledge of the palaeoneurology of sauropod dinosaurs is still deficient. This holds true even for Titanosauria, which is a particularly speciose clade of sauropods with representatives known from numerous Cretaceous sites in many countries on all continents. The data currently available regarding the palaeoneu...
Article
Full-text available
Monitor lizards (genus Varanus) inhabited Europe at least from the early Miocene to the Pleistocene. Their fossil record is limited to about 40 localities that have provided mostly isolated vertebrae. Due to the poor diagnostic value of these fossils, it was recently claimed that all the European species described prior to the 21st century are not...
Data
List of references consulted to score characters for the phylogenetic analysis. (PDF)
Data
Matrix used in the phylogenetic analyses. (NEX)
Data
Lists of the apomorphies of V. marathonensis, of the synapomorphies of the clade including V. marathonensis and the subgenus Indovaranus, and of Varanus (based on analysis 1B). (PDF)
Data
List of extant comparative specimens of Varanus. (PDF)
Data
New characters added to the original list of Conrad et al. [11]. (PDF)
Data
Results of the phylogenetic analysis 1A. (PDF)
Data
Results of the phylogenetic analysis 2A. (PDF)
Article
The Late Cretaceous fossil site of Lo Hueco (Cuenca, Spain) has yielded a large collection of tetrapod remains, in which crocodyliforms are one of the best represented groups. The crocodyliforms from Lo Hueco helped establish Allodaposuchidae as one of the sister-groups of Crocodylia. A complete skull of the holotype of the allodaposuchid Lohuecosu...
Chapter
Crocodiles and birds are the only living representatives of Archosauria, a once diverse clade of vertebrates that mastered terrestrial, aerial and aquatic environments during the Mesozoic. Because the braincases of archosaurs are largely ossified, the group has particularly benefited from advances in non-destructive visualisation of endocranial str...
Article
Karydomys is a rare and little diversified democricetodontine, of which only six species are currently recognized. This group of rodents is first recorded in the early Miocene (MN3) in China and spread quickly thereafter to Kazakhstan and Greece (MN4). Karydomys reached south‐western and central Europe by early middle Miocene times (MN5), from wher...
Article
Full-text available
Fossils of juvenile Mesozoic birds provide insight into the early evolution of avian development, however such fossils are rare. The analysis of the ossification sequence in these early-branching birds has the potential to address important questions about their comparative developmental biology and to help understand their morphological evolution...
Article
Full-text available
A palaeosurface with one megatheropod trackway and several theropod tracks and trackways from the Lower Jurassic upper Elliot Formation (Stormberg Group, Karoo Supergroup) in western Lesotho is described. The majority of the theropod tracks are referable to either Eubrontes or Kayentapus based on their morphological characteristics. The larger mega...
Conference Paper
Full-text available
Over the last decade, our knowledge of the palaeoneurology of titanosaurian sauropods has dramatically improved. While the endocranial morphology of titanosaurian sauropods has long been known only in Jainosaurus septentrionalis from the Maastrichtian of India, a considerable amount of new data on the endocranial and inner ear morphology in Late Cr...
Article
Full-text available
The end-Triassic mass extinction and the transition and explosive diversification of fauna over the Triassic-Jurassic boundary is poorly understood and poorly represented in the rock record of the Southern Hemisphere. This is despite the rich diversity in both body and trace fossils of Triassic-Jurassic age in southern Africa, which is not found in...
Article
Full-text available
Fragmentary caudal ends of the left and right mandible assigned to Lesothosaurus diagnosticus, an early ornithischian, was recently discovered in the continental red bed succession of the upper Elliot Formation (Lower Jurassic) at Likhoele Mountain (Mafeteng District) in Lesotho. Using micro-CT scanning, this mandible could be digitally reconstruct...
Data
Left maxilla of NM QR 3076 in (A) medial, (B) lateral, and (C) ventromedial view
Data
Georeferences of fossil localities for a total of 46 Lesothosaurus diagnosticus and other Lesothosaurus cf. diagnosticus specimens in the upper Elliot Formation of Lesotho and South Africa Abbreviations NHMUK, The Natural History Museum, London BMNH, British Museum of Natural History, London MNHM, Muséum National d’Histoire Naturelle, Paris BPI, Be...
Data
MicroCT scan images of the left dentary of Lesothosaurus BP/I/7853 showing transverse view of replacement tooth
Data
Locality information pertaining to the 46 Lesothosaurus diagnosticus and Lesothosaurus cf. diagnosticus specimens in the upper Elliot Formatio NHMUK, The Natural History Museum, London BMNH, British Museum of Natural History, London MNHM, Muséum National d’Histoire Naturelle, Paris BPI, Bernard Price Institute of Palaeontology, Johannesburg, South...
Data
The nine preserved dentary alveoli from tooth positions 10 –18 of BP/1/7853 in comparison to the adapted illustration of Sereno’s (1991: Fig. 13H) The nine preserved dentary alveoli from tooth positions 10 –18 of BP/1/7853 in comparison to the adapted illustration of Sereno’s (1991: Fig. 13H) reconstruction of the mandible in Lesothosaurus. The nin...
Data
MicroCT scan images of the left dentary of Lesothosaurus BP/I/7853 showing transverse view of replacement tooth behind tooth position 12
Data
3D image reconstruction and segmentation of teeth in left dentary of Lesothosaurus BP/I/7853 showing buccal and lingual view of replacement teeth (shown in blue) behind and underlying teeth at positions 3 and 8
Article
Full-text available
In Gondwana, Early Jurassic dinosaur track sites are especially concentrated in Lesotho. Despite intensive investigations during the third quarter of the twentieth century, a limited number of vertebrate track sites of this country have been studied with rigorous ichnological and sedimentological methodology. Here, we present a previously mentioned...
Article
Four isolated theropod teeth from the ?Bathonian “Argiles de l’Irhazer” in Niger are described. The teeth were found in association with the holotype of the basal sauropod Spinophorosaurus nigerensis. These specimens have been assigned to two different taxa by independent analyses, such as direct comparison with teeth previously described in the li...
Article
Full-text available
Despite continuous improvements, our knowledge of the neurocranial anatomy of sauropod dinosaurs as a whole is still poor, which is especially true for titanosaurians even though their postcranial remains are common in many Upper Cretaceous sites worldwide. Here we describe a braincase from the uppermost Cretaceous locality of ‘‘Lo Hueco” in Spain...
Article
Full-text available
Ctenodactylinae (gundis) is a clade of rodents that experienced, in Miocene time, their greatest diversification and widest distribution. They expanded from the Far East, their area of origin, to Africa, which they entered from what would become the Arabian Peninsula. Questions concerning the origin of African Ctenodactylinae persist essentially be...
Article
El neurocráneo del holotipo de Narambuenatitan palomoi Filippi, García y Garrido fue tomografiado permitiendo la reconstrucción virtual tridimensional del endocráneo y oído interno. Como en otros saurópodos, el endocráneo es bulboso y anteroposteriormente corto, con una pituitaria bien desarrollada y proyectada posteroventralmente. Como en otros ti...
Article
Three sites at the Calvero de la Higuera complex (Camino Cave, Navalmaíllo Rock Shelter, and Buena Pinta Cave), near the village of Pinilla del Valle (Madrid, Spain), are known for their record of Neanderthals and other Late Pleistocene mammals. Occasionally, they also yield much more ancient reworked remains, which come from the Upper Cretaceous d...
Article
Three sites at the Calvero de la Higuera complex (Camino Cave, Navalmaíllo Rock Shelter, and Buena Pinta Cave), near the village of Pinilla del Valle (Madrid, Spain), are known for their record of Neanderthals and other Late Pleistocene mammals. Occasionally, they also yield much more ancient reworked remains, which come from the Upper Cretaceous d...
Article
Full-text available
The Late Cretaceous (Campanian-Maastrichtian) fossil site of Lo Hueco was recently discovered close to the village of Fuentes (Cuenca, Spain) during the cutting of a little hill for installation of the railway of the Madrid-Levante high-speed train. To date, it has yielded a rich collection of well-preserved Cretaceous macrofossils, including plant...
Article
Full-text available
The Late Cretaceous (Campanian-Maastrichtian) fossil site of Lo Hueco was recently discovered close to the village of Fuentes (Cuenca, Spain) during the cutting of a little hill for installation of the railway of the Madrid-Levante high-speed train. To date, it has yielded a rich collection of well-preserved Cretaceous macrofossils, including plant...
Article
Full-text available
The modern Asian monsoonal systems are currently believed to have originated around the end of the Oligocene following a crucial step of uplift of the Tibetan-Himalayan highlands. Although monsoon possibly drove the evolution of many mammal lineages during the Neogene, no evidence thereof has been provided so far. We examined the evolutionary histo...
Article
A fragment of a maxilla and isolated theropod teeth from the (?) Middle Jurassic Tiourarén Formation are described. The specimens come from Tadibene, in the rural community of Aderbissinat, Thirozerine Department, Agadez Region, Niger. They were identified through direct comparison with teeth previously described in the literature as well as on the...
Article
Full-text available
The preservation of fossils reflects the interplay of inorganic and organic chemical processes, which should be clearly differentiated to make interpretations about the biology of extinct organisms. A new coliiformes bird (mouse bird) from the 50 million year old Green River Formation (Wyoming, USA) has here been analysed using synchrotron X-ray fl...