Evgeny V. Popov

Saratov State University · Department of Historical Geology and Paleontology

The fossil record of chondrichthyans (sharks, rays and chimaeroids) principally consists of isolated teeth, spines and dermal denticles, their cartilaginous skeleton being rarely preserved. Several Late Jurassic chondrichthyan assemblages have been studied in Europe based on large bulk samples, mainly in England, France, Germany and Spain. The firs...

This article describes Moskovirhynchus robustus gen. et sp. nov., a new genus and species of elephant fish from two Upper Jurassic (Boreal middle Volgian Substage, virgatus Zone) localities of Moscow and the Moscow region, based on isolated mandibular, palatine and vomerine dental plates. The dentition of the new genus bears a morphological resembl...

The ancestry of chimaeriform chondrichthyan fishes can be traced back to the Late Triassic (∼220 Ma). To date, only one chimaeriform suborder, the Echinochimaeroidei, has been recognized from the Palaeozoic. The origin and evolution of the chimaeriforms has been a matter of debate for more than a hundred years. This problem is aggravated by the sca...

A new elasmobranch assemblage including 14 shark taxa, mainly lamniformes (9 taxa), is described from the Lower Eocene (NP 10–11) Kalinino Formation of the Alai River Basin, Baltai District, Saratov Region of Russia. The Nikulinka elasmobranch assemblage may have characterised inner neritic environments of the eastern part of the sub-boreal Europea...

An assemblage of chondrichthyans is described from the new locality near Serafimovich town (the western part of Volgograd region). More than 2500 fossils were collected from the basal gravel-pebble bed and phosphorite-bearing layer from the lower part of the Buzinovka formation (upper Paleocene). There are reworked Cretaceous and Paleocene chondric...

По отдельным мандибулярным, небным и сошниковой зубным пластинам из двух верхнеюрских местонахождений (средневолжский подъярус, зона virgatus) Москвы и Подмосковья описывается новый род слоновых химер Moskovirhynchus robustus gen. et sp. nov. Озубление нового рода морфологически близко к структуре зубной системы современных слоновых химер рода Call...

A chimaeroid species, Edaphodon eyrensis Long, 1985 (Holocephali, Chimaeroidei), from the Lower Cretaceous Bulldog Shale of the Eromanga Basin, South Australia, is reassessed as Ptyktoptychion eyrensis (Long, 1985), comb. nov. This is the oldest representative of the endemic Australian chimaeroid genus Ptyktoptychion Lees, 1986. An ancestor of this...

This paper presents a brief biography of the Professor Maxim G. Minich, a Soviet and Russian specialist in fossil fish and stratigraphy of the Permian and Triassic of the Russian Plate and the Cis-Uralian peripheral depression. He was a brilliant representative of the middle generation of the Saratov school of vertebrate researchers, the head of th...

A chimaeroid mandibular dental plate from the nearshore upper Oligocene Sooke Formation west of Victoria, Vancouver Island, British Columbia, is described here as a new genus of chimaerid fish (Holocephali, Chimaeridae), Canadodus suntoki, gen. et sp. nov. The new genus differs from mandibular plates of both Recent and Cenozoic Chimaeridae (Chimaer...

Observations on two specimens of Elasmodectes from Morocco and Germany.

Observations on two specimens of Elasmodectes from the late Cretaceous of Morocco and late Jurassic of Germany.

An incomplete chimaeroid (Holocephali, Chimaeroidei) mandibular dental plate from the early Bajocian (Humphriesianum Zone) of Ottange-Rumelange, on the Luxembourg-French border, is described as a new genus and species of callorhinchid fish Ottangodus lotharingiae gen. et sp. nov. Comparison of the described plate with other Jurassic chimaeriform fi...

Chimaeropsis paradoxa is a myriacanthoid chimaeriform fish (family Chimaeropsidae) from the Late Jurassic Plattenkalk of Eichstätt in Bavaria, described from a single specimen by Karl Alfred von Zittel in 1887. The material was never adequately figured, and the unique holotype was destroyed in World War II. A holomorphic specimen of the fish, sligh...

A nearly complete right mandibular tooth plate of Ischyodus bifurcatus Case (Holocephali: Chimaeroidei) is reported from the Point Loma Formation (upper Campanian) of the Upper Cretaceous Rosario Group in southern California, USA. The individual is estimated to have measured nearly 1 m in total body length. Remains of I. bifurcatus have been report...

A new complex of elasmobranchs from basal horizon of Turonian of the Samara Bend of the Volga River. The complex differs from the late Cenomanian communities of chondrichthyan fishes of the adjacent territories by a taxonomy and paleoecology. Chondrichthyan fossils show clear traces of reworking.

Late Cretaceous and Paleocene chimaeroid fishes from the South Pacific region consist of three previously reported taxa: Ischyodus brevirostris Egerton, 1843 (= I. thurmanni Pictet & Campiche, 1858) and Callorhinchus hectori Newton, 1876 (both are from the Piripauan (= Santonian) of Amuri Bluff, South Island); additionally the recently described Ed...

A new locality near Serafimovich town, west of Volgograd Region, Russia, has yielded сhondrichthyans from the Bazki Regional Horizon with two fossiliferous levels, both of late Paleocene (Thanetian) in age. The fish remains were collected mostly by surface prospecting and by a small bulk sample (ca. 50 kg,) sieved between 3 and 1 mm mesh sizes. The...

Clastic strata of the eastern German Saxonain Elbtal Group, deposited under marine conditions during the Late Cretaceous (late Cenomanian–early Coniacian), have yielded various fish remains. The diverse ichthyofauna is clearly dominated by chondrichthyans. Altogether, 19 species have been identified yet. While holocephalians and hybodonts form just...

Zverkov, N.G., Averianov, A.O. & Popov E.V., April 2017. Basicranium of an elasmosaurid plesiosaur from the Campanian of European Russia. Alcheringa March. ISSN 0311-5518 The braincase of elasmosaurid plesiosaurs is poorly known. Here, we describe the exceptionally well-preserved elasmosaurid basicranium from the Rybushka Formation (lower Campanian...

Ornithuromorph birds (the clade which includes modern avian radiation) first appeared in the Early Cretaceous in Asia and achieved a great diversity during the latest ages of the Late Cretaceous (Campanian and Maastrichtian). The evolutionary history of orithuromorphs during the first 17 MYAs of the Late Cretaceous (Cenomanian to Santonian ages) re...

Sand- and siltstones, marls and limestones of the Saxonian Elbtal Group, deposited under marine conditions during the Late Cretaceous (late Cenomanian–early Coniacian), are mainly known for their rich invertebrate fauna, but have also yielded various vertebrate remains. Fish fossils have been collected from several localities at least since the ear...

The braincase of elasmosaurid plesiosaurs is poorly known despite numerous relatively complete and well-studied skeletons. Thereby a new find of an elasmosaurid basicranium from the Rybushka Formation (mid-Campanian) of Saratov Region, Russia, is of great interest. The specimen, attributed to an elasmosaurid plesiosaur, possess an elongated plate-l...

This paper consists of biographical data for well-known Soviet palaeoichthyologist Leonid S. Glickman (1929– 2000). His life is divided into several stages: a childhood and evacuation during WWII (1929–1945), life in Saratov (1945–1950), Leningrad stage (1950–1970), working in the Russian Far East (1970–1982) and his life in Leningrad / Saint Peter...

The paper presents a short biography and an annotated bibliography of the well-known Soviet palaeoichthyologist and evolutionary morphologist Leonid S. Glickman (1929–2000). His bibliography consists of 46 titles, including 2 monographs, 3 book chapters, 33 research papers, 4 popular papers, and 4 unpublished research reports and dissertations, dev...

Elasmobranchs from the Cenomanian Melovatka Formation of the Volga region (Russia) have been known since the mid-19th century and were studied by L. Glickman between 1950 and 1970. Up to now, only one middle- and numerous late Cenomanian elasmobranch assemblages have been recognized. Recent collecting and bulk sampling from a single early Cenomania...

Eorhincodon casei from Russia and Megachasma comanchensis from the United States are two Cretaceous taxa initially described as putative planktivorous elasmobranchs, but the type specimens of these two taxa were subsequently reinterpreted to represent taphonomically abraded teeth of an odontaspidid, Johnlongia Siverson (Lamniformes: Odontaspididae)...

The breakup of Pangea and onset of growth of the Pacific plate led to several paleoenvironmental feedbacks, which radically affected paleoclimate and ocean chemistry during the Jurassic. Overall, this periodwas characterized by intense volcanic degassing from large igneous provinces and circum-Panthalassan arcs, new oceanic circulation patterns, an...

Chimaeroid fishes of the genus Elasmodectes represent one of the more easily recognizable 'edaphodontids' from the Jurassic and Cretaceous. The genus is represented in the fossil record mainly by isolated small-sized dental plates bearing a number of 'archipelagic' tritors, which formed a thin cutting dentition. Isolated dorsal fin spines with a sp...

Eorhincodon casei from the lower Cenomanian of Belgorod Province in European Russia and Megachasma comanchensis from the mid-Cenomanian of Colorado in the United States are two Cretaceous nominal taxa initially described as possible planktivorous elasmobranchs. However, the type specimens of these two taxa were subsequently re-interpreted to repres...

Articulated specimens of chimaeroids are extremely rare in rocks of Cretaceous age, being known from a handful of partial skeletons from the British and Lebanese chalk. A near complete and articulated specimen of Elasmodectes willetii from Morocco represents the best preserved chimaeroid skeleton from the Cretaceous and the first holomorphic specim...

Articulated specimens of chimaeroids are extremely rare in rocks of Cretaceous age, being known from a handful of partial skeletons from the British and Lebanese chalk. A near complete and articulated specimen of Elasmodectes willetii from Morocco represents the best preserved chimaeroid skeleton from the Cretaceous and the first holomorphic specim...

A dorsal vertebra referred to as Azhdarchidae indet. from the Rybushka Formation (Upper Cretaceous, Lower Campanian) of the Beloe Ozero locality in the Saratov Region is described. Its vertebral centrum has a hypapophysis and, at the base of the neural arch, there is a large pneumatic foramen. The vertebra possibly belongs to Volgadraco bogolubovi...

The breakup of Pangea and onset of growth of the Pacific plate led to several paleoenvironmental feedbacks, which radically affected paleoclimate and ocean chemistry during the Jurassic. Overall, this period was characterized by intense volcanic degassing from large igneous provinces and circum-Panthalassan arcs, new oceanic circulation patterns, a...

Remains of Late Albian chimaeroid fishes (Holocephali, Chimaeroidei) are described from the mid-Cretaceous condensed succession of the Annopol anticline, Poland. The described material consists of dental plates, dorsal-fin spines and a single frontal clasper. These remains are assigned to five genera and seven species of ‘edaphodontids’, namely Isc...

Spectacular whole-bodied and articulated Mesozoic chimaeroid fossils are known from the Upper Jurassic Plattenkalk Lagerstatten of Southern Germany: the Upper Kimmeridgian of Nusplingen (Wuerttemberg) and the Upper Kimmeridgian and Lower Tithonian of the Solnhofen Archipelago (Bavaria). Despite high fidelity preservation, usually including elements...

Chimaeroid fishes of the genus Elasmodus Egerton, 1843 represent one of the most typical and recognizable ‘edaphodontids’ from the late Mesozoic and early Cenozoic. The genus is represented in the fossil record by isolated dental plates bearing a number of laminated tritors (etymological origin of the name), which formed a unique breaking-cutting d...

Prior to revision, the chimaeroid fishes from the Lower Oxford Clay (mainly middle Callovian) of Cambridgeshire consisted of four nominal tooth-based species (Brachymylus altidens [Woodward, 1892]; Pachymylus leedsi [Woodward, 1892]; Ischyodus egertoni [Buckland, 1835]; I. beaumonti [Egerton, 1843]). During the revision, all available specimens at...

Based on new finds of mandibular tooth plates of the chimaeroid Stoilodon aenigma Nessov &Averianov, 1996 (Holocephali, Chimaeroidei) from the Lower Cretaceous of central Russia, the diagnosis of the genus is modified and validity of the taxon is confirmed. New record of mandibular plate of the species from the Ryazanian regional stage (boreal Berr...

A study of unpublished material of Neogene chimaeroid fishes (Chimaeroidei) from California totaling 120 specimens, in the collection of the Natural History Museum of Los Angeles County, shows the presence of 3 genera and species. Both Miocene (Arikareean: Jewett Sand and Vaqueros Formations; Clarendonain: Monterey Formation; Hemphillian: Capistran...

Previously published fossil chimaeroid assemblage of Australia (Kemp, 1991; Stahl, 1999; Bartolomai, 2008) consisted of 4 genera and 8 nominal species from the Cretaceous and Mio-Pliocene. In 2010 about 100 specimens were studied by the author in the museums in Brisbane, Perth, Adelaide and Melbourne. The Mid-Cretaceous fauna of the Great Artesian...

Kimmeridgian (= Kim. below) chimaeriform fishes are known only from S England (Dorset), NW France (Normandy) and SW Germany (Württemberg) (Chimaeroidei: 3 genera, 11 species prior revision). A number of museum and private collections were recently studied in the UK, France and Germany. In addition some microvertebrate bulk sampling was undertaken (W...

Prior to revision, the chimaeroid fishes from the Middle Bathonian (Procerites progracilis zone) Stonesfield Slate (Sharp Hill Formation) of Oxfordshire consisted of ten nominal tooth-based and two fin spine-based species, all of which were described in the mid 19th Century (Agassiz, 1843; Egerton, 1843, 1847). During the revision, all available sp...

The preliminary results of a revision of the British Cretaceous chimaeroid fishes (Holocephali, Chimaeroidei) based on a study of museum collections are presented. The taxonomic composition (2 families, 9 genera, 20 species) and stratigraphic distribution of Cretaceous chimaeroid fishes are summarized. Some taxa, including Callorhinchus, Elasmodus,...

The genus Lebediodon Nessov and Averianov 1996 was described from, and restricted to, the late Albian of Belgorod Province (2 localities), European Russia. The type species L. oskolensis is known from cutting ‘Rhinochimaera’-like mandibular and palatine tooth plates where they represent 7% of all chimaeroid plates collected. During a revision of th...

Finds of tooth plates of chimaeroid fishes (Holocephali, Chimaeroidei) from six Jurassic localities of the European Russia (Moscow, Ryazan’, Kursk and Bryansk regions) are described and figured. Some plates were assigned to callorhinchids: Brachymylus cf. altidens Woodward, 1892 from the Lower-Middle Callovian (elatmae-coronatum zonal interval) of...

Fossils preserving traces of soft anatomy are rare in the fossil record; even rarer is evidence bearing on the size and shape of sense organs that provide us with insights into mode of life. Here, we describe unique fossil preservation of an avian brain from the Volgograd region of European Russia. The brain of this Melovatka bird is similar in sha...

Cerebavis cenomanica gen. et sp. nov. from the Middle Cenomanian of the Volgograd Region (Russia) is described based on a brain mold. The brain of Cerebavis is characterized by a mosaic combination of primitive and advanced features. The brain weight is estimated as approximately 1 g. The cerebrum is relatively very large, but lacks sulci. The brai...

Two incomplete chimaeroid tooth plates from the lower (?) Vvolgian (= middle-lower Tithonian, upper Jurassic) deposits of the new locality Ib-Kargort (Komi Rrepublic) are described. The plates are referred to “Edaphodontidae” gen. et sp. indet. (fragment of mandibular plate) and Ischyodus cf. schuebleri Quenstedt, 1858 (palatine plate). Kargort is...

A small collection of elasmobranchs teeth (Chondrichthyes, Elasmobranchii) from the lower Callovian Sigaloceras calloviense ammonite zone (calloviense subzone) (Middle Jurassic) of the vicinities of Saratov city is described. Teeth of Hybodus cf. H. reticulatus Agassiz, 1837, Sphenodus (?) sp., Paraorthacodus cf. P. kruckowi (Thies, 1983), Paraorth...

New materials of Hesperornis rossicus Nessov et Yarkov, 1993 (one complete and one fragmentary tarsometatarsus) from the lower Campanian Karyakino locality in Saratov Province, Russia, show for the first time distal tarsometatarsal morphology for this species and allow reconsideration of its diagnosis. H. rossicus is the most derived species of the...

Duffinodus nikolaii gen. et sp. nov., a new genus of elephant fishes (Chimaeroidei, Callorhinchidae) based on the right mandibular dental plate and the left vomerine plate from the Middle Jurassic (Upper Callovian, Quenstedticeras lamberti Ammonite Zone) from the Dubki quarry in the vicinities of Saratov (Saratov Province, Russia) is described. An...

Five Jurassic sharks teeth localities in the Volga River Basin were discovered for the first time as a result of collecting activity during last years. All of these localities are situated in the vicinity of Saratov city and contain deposits from (?) Bathonian to Volgian (= Thitonian) stratigraphical inter� vals. A locality near Dubki village has y...

A new species of giant chimaeroid Edaphodon eolucifer sp. nov. (family Edaphodontidae) from the basal horizon of the Beryozovaya Beds (Lower Paleocene, Danian) is described based on the lower jaw (“mandibular”), and anterior maxillary (“vomerine”) dental plates hypothetically associated with the former. Accompanying assemblage of elasmobranch fish...

A new shark species Galeorhinus glickmani sp. nov. (Carcharhiniformes, Triakidae) from the Upper Cenomanian (Upper Cretaceous) of Saratov is described. Teeth of similar morphology (“Corax jaekeli”, Paracorax unilateralis) from the Santonian and Campanian of Lithuania, Russia, and Kazakhstan are conditionally referred to the genus Galeorhinus Blainv...

Left lower jaw («mandibulare») dental plate of large chimaeroid Edaphodon mantelli (Buckland, 1835) from the Lower Santonian “sponges layer” of Saratov Province is described. It’s the first record of Edaphodon remains from the Lower Santonian of Volga river basin and the first true record of the species from the Upper Cretaceous of the Russia. From...

A study of representatives of the benthic and nektonic fauna that form the basis for the Cretaceous and Paleogene marine assemblages of the southeast of the East European Platform revealed the stages of the biota formation and its “provinciality” within the European Paleobiogeographic Region. A wide variety and abundance of the sublittoral fauna an...

Amylodon karamysh sp. nov. (Chondrichthyes, Holocephali, Chimaeridae) is described from the Early Campanian (Late Cretaceous) of the Saratov Region, Russia. It is based on a single mandibular plate with a relatively short outer margin and four small median tritors. The morphology of the mandibular plate of A. karamysh appears to have evolved from t...