Errol Véla

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• PhD, Associate Professor
• Lecturer at Université de Montpellier

Greater Kabylia is a regional hotspot characterized by a multitude of ecosystems, from the coast to the high mountains. The only reference study on orchids did not take into account the entire territory of the wilaya of Tizi Ouzou, unlike Bejaia (the neighboring wilaya). The present study completes the inventory, ecology and distribution of the var...

During routine inventories of the flora of Kabylia, we were able to make the regional rediscovery in Aït Ziki (wilaya of Tizi-Ouzou) of Squilla anthericoides (syn. Charybdis anthericoides, Drimia anthericoides, Scilla anthericoides, Urginea anthericoides, Urginea maritima var. anthericoides), an endemic to northeastern Algeria that had not been see...

While Epipactis palustris has been documented for the first time in Lebanon (at Mount Lebanon Governorate), this presence in the Middle East outside its classical range questions us. The species is widely distributed through Europe until Central Asia, but this southernmost location is isolated from the Turkish nearest ones from at least 500 km. Its...

Background and aims – In spite of their proximity to the coasts, the flora of the islands and islets of the Algerian coasts has been poorly studied. This work is the first to provide an overall view of the richness and the determinants of the Algerian island flora. Material and methods – The study is based on a compilation of original floristic dat...

With the aim of updating our knowledge of our natural heritage, a study was carried out on the vegetation of the Oued Sahel valley (North Algerian), an area of Kabylia that is little studied. The sampling of the flora revealed the existence of three new species for the flora of Kabylia, sector of Djurdjura/ Bibans (Dipcadi serotinum, Silybum eburne...

This study is a contribution to the knowledge of the floristic biodiversity of the Belezma and Aures massifs, recognized as important areas for plants sensitive to climate change and anthropogenic disturbances. Our subjective sampling over several years resulted in 13 stations hosting 13 taxa of orchids, including 5 never reported in this region (D...

The genus Allium is one of the most diverse in the world, with more than 1000 accepted species yet [POWO, 2023]. But a lot of new species are regularly described (several per year), including in the Euro-Mediterranean subgenus Allium. Within the sections Allium, Codonoprasum, Cupanioscordum and Pseudoscorodon, some old described species were recent...

The orchid flora of Greater Kabylia, in particular of the Djurdjura Biosphere Reserve and its surroundings (North-Central Algeria), is still poorly known, despite its importance as an indicator of biodiversity. For this reason, it has been studied more closely in recent years. This contribution is the result of a collaborative work started 17 years...

Introduction : Les premières aires protégées ont vu le jour en Algérie à partir de l'année 1983 dans les soucis de protéger et de préserver la biodiversité notamment la conservation de la flore, de la faune, des espèces à haute valeur patrimoniale et la préservation du potentiel génétique. Les missions officielles assignées aux gestionnaires des ai...

Accurate species delimitation is critical for biodiversity conservation. Integrative taxonomy has been advocated for a long time, yet tools allowing true integration of genetic and phenotypic data have been developed quite recently and applied to few models, especially in plants. In this study, we investigated species boundaries within a group of t...

Inference of recent population divergence requires fast evolving markers and necessitates to differentiate shared genetic variation caused by ancestral polymorphism and gene flow. Theoretical research shows that the use of compound marker systems integrating linked polymorphisms with different mutational dynamics, such as a microsatellite and its f...

Sixteen new Asteraceae are added to the non-indigenous flora of Tunisia. Six of them (Dimorphotheca ecklonis,Gaillardia pulchella, Gazania linearis var. linearis, Guizotia abyssinica, Rudbeckia triloba and Tithonia diversifolia) are also new for the flora of North Africa. Chrysanthemummorifolium is here first reported for the Mediterranean Basin. S...

Résumé.– La région de Constantine fait partie de la zone de Numidie qui couvre une superficie d’environ 10000 km2, réputée par une richesse floristique remarquable et une importante diversité écologique. Avec Guelma et Souk-Ahras, toutes les trois régions disposent un caractère nettement continental. Ce travail nous a permis de rassembler les ancie...

Rebbas, K., Bougaham, A.F., Rebbas, N. & E. Véla (2021): Inventory of Orchids in the Wilaya of Béjaïa (Little Kabylia, Algeria) over seventeen years.- J. Eur. Orch. 53 (2-4): in press... This article represents a collaborative work initiated 17 years ago (2004-2020) on the inventory, illustration, cartography and taxonomy of Orchids in the 52 comm...

Rear-edge populations represent reservoirs of potentially unique genetic diversity but are particularly vulnerable to global changes. While continental-scale phylogeographic studies usually do not cover these populations, more focused local scale study of rear-edge populations should help better understand both past evolutionary history and its con...

Some tree species have distributions on both sides of the Strait of Gibraltar and the Strait of Sicily. It is a challenge to determine whether such distributions result from the Tertiary or Pleistocene or from more recent dispersal related to human activities. Tetraclinis articulata (Cupressaceae) is a gymnosperm that offers an ideal model to deal...

The section Melanocrommyum Webb & Berthel. of the genus Allium L., with an Iranian-Anatolian center of diversity, has long been reduced to one species - the archeophytic segetal Allium nigrum L. - in western Europe and in the Maghreb. The discovery of Allium cyrilli Ten. in Algeria confirms the co-occurrence of two species in Maghreb, as well as in...

This is the thirteenth of a series of miscellaneous contributions, by various authors, where hitherto unpublished data relevant to both the Med-Checklist and the Euro+Med (or Sisyphus) projects are presented. This instalment deals with the families Amaryllidaceae (incl. Alliaceae), Apocynaceae, Caryophyllaceae, Chenopodiaceae, Compositae, Crassulac...

Résumé : Avec l'engouement qu'ont suscité les orchidées algériennes depuis une vingtaine d’années, et les progrès réalisés concernant leur taxonomie (De Belair & Boussouak 2002, De Bélair et al. 2005, Rebbas & Vela 2008, Rebbas & Vela 2013, Kreutz et al. 2013, 2014, Martin et al. 2020, etc.), des travaux de terrain à vocation floristique ont été mu...

The Mediterranean low-altitude limestone cliffs are known for their exceptional diversity and richness in endemics, often insufficiently assessed in Algeria. The present study is an analysis and a synthesis of individual IUCN assessments of three endemic Algerian chasmophytic species, Silene sessionis, S. aristidis and S. auriculifolia. The aim is...

Among Mediterranean trees, “ Juniperus turbinata” remains a little-known taxon. Described in 1844 (J. turbinata Guss.) it was ignored in many floras and checklists until very recently despite the fact that since the 1980s morphology and phytochemistry studies have confirmed its taxonomic individuality. The conclusions of recent molecular studies al...

Revue électronique annuelle de la Société botanique du Centre-Ouest-Evaxiana n°8-2021 Tison J.M., Abdulhak S., Bock B., Boudrie M., Fridlender A., Roccia A., Van Es J., Véla E., 2021-Combinaisons nouvelles requises dans la seconde édition de Flora. - Résumé. 73 combinaisons et noms nouveaux sont proposés en vue de changements taxonomiques dans la s...

Details on the distribution of Malva stenopetala (Malvaceae) in Algeria. Bull. Soc. linn. Provence, 71: 229-233 (2020). ISSN: 0373-0875. In this note, we report new field observations and update from herbaria the known geographic distribution of Malva stenopetala in Algeria. These observations began to reinforce as desirable the knowledge on thi...

Discovery of Epipactis microphylla (Orchidaceae) in Djebel Babor (north-eastern Algeria), a new orchid for North Africa. — Epipactis microphylla is a European-Hyrcanian, forest mycoheterotrophic species, widespread in temperate and submeridional zones, from Andalusia to the Caspian Sea. This orchid has been discovered in Djebel Babor, in northeast...

The coast of Mostaganem (Algeria), an "important area for plants" (ZIP) as much neglected as threatened. – The flora of the coastline of Mostaganem still till nowadays remains very little known, even though it has been historically very well studied, like the whole Oran’s coastal area. The objective of our study is to inventory the local plant dive...

The diversity of habitats in the Cévennes national Park is home to a rich flora, comprising more than 2400 species (angiosperms, gymnosperms, and ferns). A good knowledge of this flora is essential for the development of adapted management strategies. However, human resources for this purpose are limited, and the support of residents and visitors c...

La systématique de Quercus faginea Lam. (s.l.) (Fagaceae) reste encore indéfinie, d’autant plus qu’elle s’hybride avec d’autres espèces et qu’elle est polymorphe. Quatre sous-espèces ont été identifiées et présentement reconnues en péninsule Ibérique et au Maroc, or, les données disponibles ne permettent pas de trancher entre les taxons présents en...

This species is endemic to the "Kabylies-Numidia-Kroumiria" plant diversity hotspot (Véla and Benhouhou 2007) in northern Algeria. The extent of occurrence (EOO) is estimated at 2,623 km2 and the area of occupancy (AOO) is 32 km2. The species is assessed as Vulnerable (A3c; B1ab(iii,iv,v)+2ab(iii,iv,v); C2a(i)) because of its natural restricted pop...

The species is recorded from North Africa. The extent of occurrence is large, however the area of occupancy (AOO) is likely to be restricted; it is estimated at 176 km2 based on mapped localities, although this is an underestimate. The species is still locally abundant in Tunisia, but rare elsewhere, and populations are considered to be declining a...

This species is endemic to a very restricted area of the coastal submaritime cliffs from Djebel Murdjadjou to Djebel Santa Cruz massif close to the city of Oran, Algeria, and from another locality (Djebel Krichtel) now presumed extinct. The species is classified as Critically Endangered (CR: B1ab(iv,v)+2ab(iv,v); C2a(i,ii); D) because of its very...

Vascular plants new or rare for Tunisia occurring on the islands (Galite, Zembra, Kuriat, Monastir, Kerkennah, Kneiss, Djerba). – This study concerns the indigenous vascular plants newly reported for Tunisia and discovered on the islands during some field trips performed since 2014. Five native taxa are new to Tunisia: Bellis perennis, Carthamus cr...

Abstract. – Orchids in the northern part of the Setif province (North-East of Algeria). As a follow-up to the previous study on the orchids of the wilaya of Setif and continuing to explore the different potential wider sites that could shelter these plants, this inventory brings other observations of orchids in an area that ranges from Setif to Khe...

The same as https://www.researchgate.net/publication/340878201_Inventaire_des_Orchidees_de_la_partie_septentrionale_de_la_wilaya_de_Setif_nord-est_de_l%27Algerie

The Mesopotamian Region is often recognised as the origin of the Western European (W European) segetal plants that have been coevolving with winter cereals since the emergence of agriculture. This eastern origin assumption is based on the close association between cereal domestication and arable weed occurrence. However, it has never been properly...

This document is a summary of recent data about orchids from north-eastern Algeria. The areas concerned are the Kabylia in the broad sense and the historic Algerian Numidia with its adjacent mountains of Hodna and Aurès. They are very diverse territories in terms of geography (from the coast up to + 2000m altitude), bioclimate (from the semi-arid s...

This report includes the results of a botanical and phytoecological survey carried out in spring 2019 on the island of Zembra and on some of the islets of this archipelago, as part of a mission of the PIM Initiative (International NGO for Mediterranean Small Islands). This work includes the checklists of the vascular plants observed to date by the...

La diversité des habitats du Parc national des Cévennes héberge une flore riche, composée de plus de 2400 espèces (angiospermes, gymnospermes et fougères). Une bonne connaissance de cette flore est essentielle pour le développement de stratégies de gestion adaptées. Les ressources humaines étant cependant limitées, l’appui des résidents et visiteur...

VÉLA E., 2019 – Allium baeticum. The IUCN Red List of Threatened Species, 2019 (3): e.T172150A19094919. The species is widespread in Morocco and the southern Iberian Peninsula, although less common in Portugal. The plant is assessed as Least Concern in view of its relatively wide distribution and the fact that it grows in a variety of habitats an...

YOUSSEF S., MAHMOOD A., HUSSEIN W., & VÉLA E., 2019 – Allium noeanum. The IUCN Red List of Threatened Species, 2019 (3): e.T135303925A135303932. Historically, this species had been found in 27 localities but recently 45 other new localities of this species often has been discovered in the 2005-2017 period, mainly in southeastern Anatolia in the T...

VÉLA E., 2019 – Cephalanthera epipactoides. The IUCN Red List of Threatened Species, 2019 (3): e.T176038A21336941. This orchid is endemic to eastern Greece, western Turkey and southern Bulgaria. Populations are more or less localised but can sometimes be abundant. It has a wide distribution, with an extent of occurrence exceeding 100,000 km2 and...

LANSDOWN R.V. & VÉLA E., 2019 – Desmazeria sicula. The IUCN Red List of Threatened Species, 2019 (3): e.T13165092A18610018. The species is endemic to the Mediterranean Hotspot region, with a wide distribution, but is known from very few sites (AOO 62 km2), many widely separated. It is described as locally abundant but it occurs in a habitat which...

YOUSSEF S. & VELA E., 2019 – Viola pachyrrhiza. The IUCN Red List of Threatened Species, 2019 (3): e.T135527906A135527918. The species is naturally fragmented in cliff habitats considered as stable, straddling two countries, through the Iraqi and Persian Zagros Mountains, from 1,500 to 3,500 m asl. No threats are currently known in these kinds of...

VÉLA E., 2019 – Allium duriaeanum. The IUCN Red List of Threatened Species, 2019 (3): e.T116540535A116540538. This species is endemic to northern coastal areas of Tunisia and Algeria. It has a restricted area of occupancy (AOO) that is estimated to between 24 and 40 km2, however this estimate does not include potential new location not yet discov...

VÉLA E., 2019 – Allium polyanthum. The IUCN Red List of Threatened Species, 2019 (3): e.T172193A129513554. The species as here circumscribed is considered Least Concern because of its frequency, abundance and dynamic populations in many synanthropic environments (road embankments, fields, orchards, rural gardens for example) at least in southern...

MAHMOOD A., YOUSSEF S., & VÉLA E., 2019 – Allium trachycoleum. The IUCN Red List of Threatened Species, 2019 (3): e.T136006215A136006220. Allium trachycoleum is an endemic wild leek of the eastern Mediterranean and Irano-Anatolian hotspots with a disjoint range restricted to Mt. Hermon in Israel/Syria and the Taurusian-Zagrosian Mountain range in...

VÉLA E., REBBAS K. & LANSDOWN R.V., 2019 – Allium trichocnemis. The IUCN Red List of Threatened Species, 2019 (3): e.T13145944A18610859. This species is endemic to North Africa, where it is known from Algeria, with presence in Morocco now rejected. The species has a small but well-conserved subpopulation in the Gouyara National Park, but a restri...

VÉLA E., 2019 – Prospero pulchellum. The IUCN Red List of Threatened Species, 2019 (3): e.T136106211A136106213. Because of the strong uncertainty about its taxonomic delimitation, the geographical range is not well defined, varying from a possible northwestern Algerian endemic to a possibly widespread western Mediterranean species, but most likel...

MESBAH M., BEKDOUCHE F., VÉLA E. & SAHNOUNE M., 2019 – Silene sessionis. The IUCN Red List of Threatened Species, 2019 (3): e.T72458759A72458763. This species is endemic to a very restricted area of coastal sea cliffs on the Yemma Gouraya Massif near the city of Bejaia, Algeria. The species is assessed as Endangered (EN: D) because of its natural...

YOUSSEF S., MAHMOOD A. & VÉLA E., 2019 – Allium pustulosum. The IUCN Red List of Threatened Species, 2019 (3): e.T136106093A136106119. Allium pustulosum is officially known since 1882 from southeastern Turkey, then after in Iraq and Iran under the synonymy “Allium qaradaghense”. Recent field data from Turkey produced a lot a new localities, but a...

This is the eleventh of a series of miscellaneous contributions, by various authors, where hitherto unpublished data relevant to both the Med-Checklist and the Euro+Med (or Sisyphus) projects are presented. This instalment deals with the families Anacardiaceae, Asparagaceae (incl. Hyacinthaceae), Bignoniaceae, Cactaceae, Compositae, Cruciferae, Cyp...

The Brachypodium distachyon complex includes (at least) three annual species, whose taxonomic identities have recently been demonstrated by cytological, molecular and morphological data (Catalan et al. 2012-Ann. Bot., 109: 305-405): B., stacei, B. distachyon, B. hybridum. The initial taxonomic studies contained no nomenclatural revision of the fift...

Rediscovery of Allium scaberrimum J. Serres (A. pardoi Loscos syn.) in North Africa (Algeria) Abstract. – A species lost for more than 80 years in North Africa, Allium scaberrimum J. Serres (Amaryllidaceae) has been observed in several localities in northern Algeria, including the three historical regions of presence in the country. Despite its lon...

The Brachypodium distachyon complex includes three annual species, whose taxonomic identities have recently been demonstrated by cytological, molecular and morphological data. The initial taxonomic studies contained no nomenclatural revision of the twenty or so heterotypic synonyms at species rank, while the authors needed to describe two species a...

Les secteurs et sous-secteurs phytogéographiques de l’Algérie de Quézel & Santa (1962) sont consacrés par l’usage, mais n’ont pas fait l’objet de description précise ni d’analyse critique. L’objectif est de replacer la sectorisation de Quézel & Santa parmi les travaux antérieurs (Lapie, 1909 ; Maire, 1926) ou postérieurs (Barry & Celles, 1974 ; Alc...

Christella dentata (= Cylosorus dentatus, Thelypteridaceae, Pteridophyta), native to the tropical and subtropical regions of Africa and Asia, has been observed between Jijel and Bejaia in northeastern Algeria. Distribution, ecological notes and comparison with related taxa as Cyclosorus interruptus (Dryopteris gongylodes p.p.) are also provided. Th...

This illustrated electronic book is about the natural history of the Kurdish autonomous region of Iraq and its surroundings. It focuses on the Orchidaceae botanical family as a windows on the biodiversity of the North-Western Zagros, part of the Irano-Anatolian biodiversity hotspot. A general introduction presents the main aspects of orchids as the...

De tous temps, l'inventaire et la description de la biodiversité se sont heurtés à des difficultés diverses. Aujourd'hui le besoin de conservation de cette biodiversité sujette aux menaces des activités humaines nous oblige à reconsidérer ces difficultés pour mieux les dépasser. En Algérie la connaissance de la flore a connu des sommets juste avant...

Concluding messages 1. In order to halt and reverse the continuous decline of plant diversity, specific action plans and programmes addressing particular plant conservation challenges must be incorporated in regional and national conservation programmes and strategies. 2. Enhanced cooperation between botanists, practitioners and communities is esse...

The current Red List assessment of Mediterranean plants aims to assess all endemic and near-endemic monocotyledon taxa of the Mediterranean Basin Hotspot, in order to identify those that are threatened with extinction. Currently, more than 600 species or subspecies have been assessed (including more than 150 orchids), while hundreds of synonyms wer...

This species has a current extent of occurrence of around 30,000 km² after an historical reduction, an area of occupancy comprised between 64 and 100 km² and it could be found in between 15 and 30 locations. After the destruction of several historical locations by agriculture and urbanisation, now the species is not severely fragmented, but the dec...

This species is listed as Least Concern because it is widespread and common where it occurs and its population is considered stable and currently experiencing a lack of major threats to itself and to its habitats.

This Mediterranean orchid occurs mostly in North Africa, from Morocco to western Tunisia. In Europe, it is only known from a small number of localities in the Andalusian province of Malaga, Spain. The extent of occurrence (EOO) is large, however, the area of occupancy (AOO) is estimated at around 200 km2. The global population is estimated between...

This orchid is endemic to the Mediterranean region in Algeria and Tunisia, where it is relatively widely distributed. The limited population information available shows that some sites host only a few mature individuals and the total population size is suspected to be less than 10,000 mature individuals, with none of the estimated subpopulations in...

The species is endemic to the northern coastal area of the Cyrenaica region of Libya. There are few recent records and it is not known if older localities, especially those close to human settlements, still exist. Based on known recent records and bibliography, the extent of occurrence (EOO) and area of occupancy (AOO) are estimated at >500 and >20...

This rare Mediterranean orchid is endemic to northeastern Algeria and northwestern Tunisia mountain oak forests. It is assessed as Endangered (EN B2ab(ii,iii,iv,v)) in view of its restricted range, with an area of occupancy (AOO) estimated at 20 km2, considering only recent and confirmed records. A continuing decline in the extent and quality of it...

This eastern Mediterranean orchid is listed as Least Concern because it has a wide distribution with bundant subpopulations in part of its range, and does seem not face any major threats at present.

This southwestern Mediterranean orchid is native to Sicily, Tunisia and Algeria. It is listed as Least Concern because its population is considered stable and no facing major threats at present. In Tunisia, this taxon is very rare and localised and, therefore, there is a need for further research on its distribution and population status in this co...

Mediterranean regional assessment: Least Concern (LC). In the Mediterranean, this orchid is rather local and sometimes abundant with quite a large distribution area. The population is decreasing in parts of its range due to habitat loss but can also increase due to new habitat colonisation. Furthermore, the existing threats are unlikely to cause th...

Mediterranean Regional Assessment: Least Concern (LC). In the Mediterranean, like in the global area, this orchid is rather local and sometimes abundant with quite a large distribution area. Nevertheless, favourable habitat has been destroyed by urbanisation and agriculture intensification and the population could also locally decrease due to speci...

Mediterranean regional assessment: Least Concern (LC) This Mediterranean orchid native to the Iberian Peninsula and North Africa is listed as Least Concern due to its widespread distribution, abundance and absence of major threats to its population, at least in Spain where are present a very large majority of its population.

Mediterranean regional assessment: Vulnerable (VU) This orchid has a disjunct distribution in the Mediterranean region, with two distinct subpopulations, one in North Africa and the other in northwestern Italy. The extent of occurrence (EOO) is large, however, the area of occupancy (AOO) is restricted and has been estimated at 1,260 km2. The popula...

The authors present the results of more than 5 years of observations of orchids in the western Tell Atlas of Tiaret (Algeria). These taxa were inventoried at the regional level, according to subjective sampling, and the ecological conditions at each of the stations collected. The identification of taxa was made through different flora, with a taxon...

Prospero seisumsianum (Rukšans & Zetterl.) Yıldırım, a species recently described from SE-Turkey, is recorded for the first time to the flora of Iraq based on samples collected from Amadya and Duhok areas (N-Iraq). We contribute here to know better its description, cytology and chorology. The specimen collected from the Mesopotamian dry plain diffe...

Arrival of Senecio angulatus (Asteraceae) in Algeria. — In this contribution, the authors report for the first time the presence of Senecio angulatus L. (Asteraceae) in Algeria. Field observations have shown the presence of this taxon in five localities in the center of the country. The identification was checked with particular regard to closely r...

In this contribution, the authors report for the first time the presence of Senecio angulatus L. (Asteraceae) in Algeria. Field observations have shown the presence of this taxon in five localities in the center of the country. The identification was checked with particular regard to closely related taxa (e.g. Senecio mikanioides Harv.). This repor...

This species is assessed as Least Concern in view of its wide distribution, presumed large overall population, and because it is unlikely to be declining fast enough to qualify for listing in a more threatened category.

This species is is considered rare and declining in Italy and Greece but there is no information on population trends in North Africa, consequently the information available is not adequate to assess the conservation status of the species and it is therefore classed as Data Deficient, there is an urgent need to collect information on its population...

This species is considered widespread and sometimes locally abundant. It is known from records scattered throughout mainland Spain, Morocco, Tunisia, Algeria, Libya, and the islands of Corsica in France, Sardinia and Sicily in Italy, and the Balearic Islands in Spain. Nevertheless, since the beginning of the 20th century, there has been a continuou...

This species is relatively widespread along coastal areas of the Mediterranean suitable habitat. Some localised threats are reported but they are not affecting the species at a rate to justify a threatened category, and it is assessed as Least Concern in view of its wide distribution and local abundance in many islands.

This species is endemic to mountains areas in Crete, where it is found in forest habitats, with localised sub-populations, and the population trend is declining. The extent of occurrence is below 2,000 km2 and the area of occupancy is 56 km2 (historically 80 km2). The population has been estimated to be as low as c.450 individuals but is possibly h...

This species is endemic to southern France and eastern Spain. It is assessed as Least Concern because it has a widespread distribution, occurs in several protected areas throughout its range and its population is not expected to significantly decline in the near future. Further studies on the genetic structure of its population and trends in popula...

The species is not yet well known from a taxonomic and biogeographic point of view. Nevertheless, the frequency and local abundance of the species at least in the central part of northern Algeria, from sea level until middle mountain altitudes, including both natural and synanthropic habitats, allows us to consider it as Least Concern.

This plant is present in the western Mediterranean countries of Spain and Portugal. It has dense populations in part of its range, some of them being located in protected areas. Although it is impacted by overgrazing and urban development in North Africa, and there is a potential threat of future over-exploitation of wild subpopulations for commerc...

This species is near-endemic to the Mediterranean region (95%), only extending outside the region in northwestern Italy. It has a well-understood distribution in northeastern Spain (Aragón), and southern France (Durance basin and Herault department), however its current distribution, and population size and trend in North Africa is not known; it ha...