Enzo Colonnelli

Enzo Colonnelli
  • Sapienza University of Rome

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Publications (93)
Article
Full-text available
During the last years, we had the opportunity of extensive collecting of Curculionoidea Latreille, 1802 in Friuli region, northeastern Italy. The surveyed area, centered mainly in Val Resia, a relatively wide west-east axis river valley surrounded by quite high mountains peaking in Monte Canin (m 2587 a.s.l.), is located at the border between Italy...
Article
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Two new species of Coniatus are described with taxonomic remarks and a key to species.
Article
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During the last years, we came across two unusual species of Coniatus Germar, 1817 and, although they are both represented by a few individuals, we decided to describe them and to tentatively clarify the intricate taxonomy of both the tribe to which they belong and of species of the genus.
Book
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An annotated catalogue of Coleoptera Curculionoidea of “Monte Grappa” Biosphere Reserve is presented. The 528 species surveyed, divided into 7 families and 283 genera and subgenera, include the 265 species already known, plus the 263 are reported for the first time for the area. Faunistic, chorological and ecological information is given for all th...
Book
Full-text available
The present catalogue is an improved and updated edition of the one published in November 2017.It contains 1627 valid genera, 681 valid subgenera, 15,531 valid species, 1,013 valid subspecies and9,098 names of invalid or nomenclaturally unavailable species-group taxa belonging to the sevenfamilies of Coleoptera Curculionoidea recognised in the Pala...
Preprint
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This is the preview of the volume dedicated to the Curculionoidea beetles of the Man and Biosphere Reserve (MaB) of Monte Grappa (NE-Italy). The book, edited by World Biodiversity Association, will be officially released the 24th of February 2023.
Article
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The taxonomical status of some species of Otiorhynchus is clarified, with a description of a new species group and a new endemic species from central Appenines.
Article
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The trophic ecology of Chiloneus weevil species from Sicily was unknown until recently when the host plant of these closely related species was discovered. The opportunity to collect many samples of these elusive species allowed us to re-evaluate the morphology and the genetic relationships among the three closest of the four species inhabiting Sic...
Article
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A new genus of Entiminae, an endogean weevil of the tribe Otiorhynchini, Giavarhynchus Bellò, Osella & Ruzzier, gen. n., and its type species Giavarhynchus amicorum Bellò, Osella & Ruzzier, sp. n. are described. The new taxon is readily distinguished from all other members of the tribe due to the unique combination of lack of eyes, elongate rostrum...
Article
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Contribution to the knowledge on the weevil-fauna of the Algarve, Portugal (Coleo ptera: Curculionoidea). During two excursions to the Algarve region in southern Portugal, 216 Curculionoidea were collected. Among them, 31 species are new records for Portugal (9 Apionidae, 1 Brachyceridae, 19 Curculioni-dae, 2 Nanophyidae). Among Curculionidae, the...
Article
An annotated synopsis of Iranian Brentidae (Coleoptera: Curculionoidea) is provided as a basis for future studies, with notes on distribution and plant associations. In total, 123 species from 41 genera and 3 subfamilies, Apioninae (109 species within 32 genera), Brentinae (3 species within 3 genera) and Nanophyinae (11 species within 6 genera) are...
Article
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The unexpected finding of an Otiorhynchus Germar, 1822 sensu stricto on the Mount Amiata, central Italy, led to designate a neotype for Curculio clavipes Bonsdorff, 1785 and to discover several overlooked synonymies among the species group to which it belongs. The new synonymies are as follows: Otiorhynchus clavipes (Bonsdorff, 1785), stat. rev. [=...
Article
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The recently published catalogue of Palaearctic Curculionoidea (Alonso-Zaraza ga et al. 2017) lists for the genus Otiorhynchus Germar, 1822 O. (Podonebistus) phasma Rottenberg, 1872 and O. (Metopiorrhynchus) garibaldinus Solari & Solari, 1909, both distributed in Sicily and in Calabria (Sparacio 1999, Osella et al. 2004, Abbazz i & Maggini 2009). T...
Chapter
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An annotated catalogue of the 394 species (divided into 203 genera or sub-genera) of Coleoptera Curculionoidea thus far known from Tuscan Archipelago, a group of islands and islets in the Tyrrhenian Sea located between the Sardinian-Corsican islands complex and central Italy, is presented. Faunistic, chorological and ecological information is given...
Chapter
Full-text available
The following new synonymies in Coleoptera Curculionidae are established: Sitona sulcifrons (Thunberg, 1798) [ = Sitona argutulus Gyllenhal, 1834; syn. n.]; Entomoderus laesirostris (Fairmaire, 1859: 268) [= Rhytirhinus loesirostris Fairmaire, 1859: 278, alternative original spelling, here rejected; syn. n.]; Aulacobaris caerulescens (Scopoli, 1763...
Article
The fauna of Iranian Anthribidae and Nemonychidae (Coleoptera: Curculionoidea) is summarized in this paper. In total, 13 species within nine genera of Anthribidae (Allandrus LeConte, 1876, Enedreytes Schoenherr, 1839, Platyrhinus Clairville, 1798, Platystomos D.H. Schneider, 1791, Dissoleucas Jordan, 1925, Noxius Jordan, 1936, Araecerus Schoenherr,...
Article
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Is described the new genus Viridistylus n. gen. and its type species Viridistylus robustus n. sp. from Tajikistan belonging to the entimine tribe Mesostylini. A key to the genera of Mesostylini thus far known, and illustration of the new species are included.
Article
An overview of Iranian Attelabidae (Coleoptera: Curculionoidea) is given. In total, 30 species within 12 genera are listed as belonging to the fauna of Iran (three species of Attelabinae, and 27 species of Rhynchitinae). Two Rhynchitinae species, Temnocerus coeruleus (Fabricius, 1798) and Mecorhis (Pseudomechoris) aethiops Bach, 1854 are new record...
Article
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Based on specimen examination and previous records, we provide a checklist including 146 species from 51 genera and 10 tribes of the weevil subfamily Ceutorhynchinae in China. Of these species, the following 18 are reported in China for the first time: Amalus scortillum (Herbst, 1795), Ceutorhynchus filiae Dalla Torre, 1922, Dieckmannius ingensColo...
Article
An annotated list of the single species of Cleridae and Anthribidae, and two of Curculionidae described by Ponza from the environs of Saluzzo in northwestern Italy, partly overlooked by subsequent authors, is provided. The following new synonymy in Curculionidae is established: Hylobius (Callirus) transversovittatus (Goeze, 1777) [= Curculio brunon...
Conference Paper
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The wild olive seed weevil, Anchonocranus oleae Marshall, 1912 was not found after its description and the study of its biology by Silvestri in 1915. Recently, A. oleae larvae were found alive inside the kernels of wild olives (O. europaea subsp. cuspidata) collected from trees between 2009 and 2012 in the Western Cape, South Africa. No weevil larv...
Article
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Here we report the first finding of Tychius amplicollis Aubé, 1850 in central Italy, and its association with the Fabaceae Vicia sativa subsp. macrocarpa (Moris) ArcAng. This is to date the first observed host plant for a species of the subgenus Apeltarius Desbrochers Des Loges, 1873.
Article
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У даній роботі розглянуто результати фауністичних досліджень, проведених на території Ірану. Матеріал був зібраний з використанням різних ентомологічних методик. Крім того, були використані матеріали ентомологічних колекцій інститутів та університетів. Всі матеріали зберігаються у колекціях авторів публікації. В роботі для кожного виду надано інфор...
Article
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The new genus Gibbopterus gen. nov. is included among Scleropterini to accommodate the new species of wingless weevils Gibbopterus guizhounus sp. nov. and Gibbopterus tenuis sp. nov., both from southwest China. The new genus's relationships with its three close relatives - Scleropteroides, Scleropterus and Proscleropterus - are discussed.
Article
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We provided a list of 224 specimens belonging to 53 ceutorhynchine species (Coleoptera, Curculionidae), which were intercepted from 1984 to 2016 at the Narita Sub-station, Yokohama Plant Protection Station, Japan. The specimens examined were found by import inspections of vegetables and cut flowers from Africa, Europe, East Asia, Oceania, and North...
Book
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The present catalogue contains 1,611 valid genera, 678 valid subgenera, 15,407 valid species, 1,042 valid subspecies and 9,381 names of invalid or nomenclaturally unavailable species-group or infrasubspecific taxa belonging to the six families of Coleoptera Curculionoidea recognised in the Palaearctic Region. New combinations are: Mecorhis holzschu...
Poster
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Phytophagous weevils of the genus Trichosirocalus include 17 species with Palaearctic distribution that mainly feed on Plantaginaceae and Asteraceae. In particular, T. horridus shows a distribution from Iberian Peninsula to Caucasus and an association to vegetal species belonging to the genus Carduus, Cirsium and occasionally Onopordum (Asteraceae,...
Article
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Chiloneus hoffmanni (González, 1970) (Coleoptera Curculionidae) thus far considered a Maltese endemic, is here recorded for the first time from Lampedusa island (Italy). Several adults of this species and of C. solarii Pesarini, 1970 were found feeding on leaves of Charybdis pancration (Asparagaceae). These observations constitute the first data on...
Article
Full-text available
Chiloneus hoffmanni (González, 1970) (Coleoptera Curculionidae) thus far considered a Maltese endemic, is here recorded for the first time from Lampedusa island (Italy). Several adults of this species and of C. solarii Pesarini, 1970 were found feeding on leaves of Charybdis pancration (Asparagaceae). These observations constitute the first data on...
Article
Full-text available
A new species Mogulones neslihanae Korotyaev, Gültekin & Colonnelli sp. nov. is described from Northeastern Turkey where it lives on Nonea caspica. The new species is very closely related to M. tristis (Korotyaev, 1981) from the Hakkari and Şırnak provinces of Southeastern Turkey; M. neslihanae sp. nov. differs primarily in the smaller body size an...
Article
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On the types of Otiorhynchus binaghii Luigioni, 1931 (Coleoptera: Curculionidae) The discovery of two genuine syntypes of Otiorhynchus binaghii Luigioni, 1931 is reported after checking the Luigioni collection preserved in the Museo Civico di Zoologia, Rome, Italy. The authorship of the species was incorrectly attributed by Luigioni to Ferdinando S...
Article
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A new species of Glocianus REITTER, 1916 from Iran, with notes on the taxonomy of this genus, and a new synonymy (Coleoptera: Curculionidae) Abstract Glocianus martini sp.n. (Coleoptera: Curculionidae) is described from Iran. Characters to differentiate it from G. herbsti (FAUST, 1895) are provided. A lectotype is designated for Ceutorhynchus herbs...
Article
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Riassunto. Brachycerus normandi Desbrochers des Loges, 1898, finora noto solo del Nordafrica, viene riportato per l'isola di Lampedusa, rappresentando la prima segnalazione per l'Italia e per l'Europa. Viene anche fornita una chiave dicotomica per il riconoscimento delle 11 specie di Brachycerus italiani. Abstract. Brachycerus normandi Desbrochers...
Article
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Otiorhynchus (Tecutinus) lefkaoriensis sp. nov. from the Lefka Ori mountains in Chania province of Crete Island is described and illustrated. The new species belongs in the kindermanni Stierlin, 1861 species group and represents the first certain discovery of a member of the subgenus Tecutinus Reitter, 1912 outside Western Asia.
Article
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Intensive sampling in Rawdhat Khorim National Park located in the central part of the Kingdom of Saudi Arabia yielded additional new country records of Curculionoidea. Twelve species of Curculionidae and one of Nanophyidae are reported for the first time from the Kingdom of Saudi Arabia.
Article
Trichosirocalus horridus sensu lato has been used as a biological control agent of several invasive thistles (Carduus spp., Cirsium spp. and Onopordum spp.) since 1974. It has been recognized as a single species until 2002, when it was split into three species based on morphological characters: T. horridus, Trichosirocalus briesei and Trichosirocal...
Article
Full-text available
First record of Listroderes costirostris Schoenherr, 1826 (Coleoptera: Curculionidae) for Italy and Tunisia Listroderes costirostris Schoenherr, 1826 is recorded for the first time for Italy (Rome) and Tunisia (Port El Kantaoui). This weevil of South American origin, severely injurious to many cultivated plants, is now spreading in several countrie...
Article
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A collection of weevils was randomly carried out in various regions of Pakistan. Six of the species collected, belonging to three subfamilies of Curculionidae, result new for Pakistan: Megamecus sciurus (Olivier, 1807) (Entiminae); Asproparthenis punctiventris (Germar, 1824), Menecleonus virgatus (Schoenherr, 1832) and Temnorhinus brevirostris (Gyl...
Article
Full-text available
is described and illustrated the parthenogenetic species Otiorhynchus (Eprahenus) cooteri sp. n. from Kazakhstan and Uzbekistan, close to the amphigonic O. aksudshabaglinus Bajtenov, 1974 from an adjoining area of Kazakhstan. The new species is allied with other central asiatic O. (Eprahenus), with all of which is shortly compared, whereas other me...
Article
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Study of the type specimen of Otiorhynchus ambrosii Bertolini, 1893 allowed to reinstate the synonymy: Dodecastichus mastix (Olivier, 1807) [= Otiorhynchus (Dodecastichus) ambrosii Bertolini, 1893; syn. n.]. As a consequence, the following synonymy is also established: Dodecastichus consentaneus (Boheman, 1842) [= Otiorhynchus (Dodecastichus) conse...
Article
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Basing on the study of the type specimen of Curculio pericarpius Linnaeus, 1758 preserved in the Linnean Collection in London, and here designated as the lectotype, it was found that this name was misapplied to another close species of the genus Rhinoncus Schoenherr, 1825 starting from the Paykull 1792 misidentification. As a consequence, the follo...
Article
Full-text available
Anthonomus eugenii Cano (Coleoptera: Curculionidae) was observed for the first time in Italy and the Mediterranean Region attacking sweet pepper (Capsicum annum L.; Solanaceae) in greenhouses and in fields in the coastal area of the Lazio Region of Italy. The incursion, detected in Oct 2013, was a heavy infestation of sweet pepper buds and not full...
Article
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This contribution deals with a total of 71 species of the curculionoid families Apionidae, Nanophyidae, Brachyceridae and Curculionidae (except for Scolytinae) recorded from Socotra Island, based on both literature and new records. The following twelve new genera of Curculionidae are described: Armifemur gen. nov. (type species A. pusillus sp. nov....
Article
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Curculionoidea of Monti Prenestini, with description of a new species of Otiorhynchus (Coleoptera, Curculionoidea) The 468 species found on Monti Prenestini, a mountain range of moderate extent and elevation situated in Latium east of Rome, are listed. The new edaphic Otiorhynchus (Lixorrhynchus) hystricis n. sp., differing from all hitherto known...
Article
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A new species Otiorhynchus (Aranihus) venarum n. sp. from the central Apennines is described. It differs from the close O. ligneus (Olivier) and O. frescati BOheman by its longer rostrum, shorter second funicular joint, less erect elytral setae, and acutely crenulate tibiae. Otiorhynchus colasi rOudier, 1854 is elevated to species rank from subspec...
Article
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Cyllorhynchites sarahae sp. nov. is described upon some specimens recently collected in Cyrenaica (Libya). The genus Cyllorhynchites Voss is new for the Mediterranean Basin: it was so far known only from Asia.
Article
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Parascythopus saltinii n. sp. from Syria is described and illustrated. The new species cannot be confused with any other by its apical part of rostrum reddish and its lack of pronotal keel in combination with the elongate funicular and tarsal segments. A key to species of Parascythopus Desbrochers is included. New combination is: Parascythopus intr...
Article
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The name Megorchestes Kojima, 2011 is here selected as the correct original spelling over Magorchestes (alternative original spelling, here rejected) and Megorchsetes (alternative original spelling, here rejected) of a recently described genus of Oriental Rhamphini.
Article
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The 195 species of Curculionoidea (Coleoptera) thus far indicated from Golestan province (northern Iran) according to both literature and original records are listed in this paper. New data for 61 species belonging to 18 genera collected during this research are also given. Besides the 42 species newly reported for the Golestan province, the follow...
Article
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Seven Curculionoidea described from Lombardy (northern Italy) by Virginio Betta in 1857 and overlooked by subsequent authors are here treated. Since the Betta collection where their types are supposed to have been preserved was never traced, neotypes of all of them are designated with the purpose of clarifying their taxonomic status. Revised combin...
Article
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Riassunto - Gli autori forniscono dati precisi sulla distribuzione in Italia dell’antribide Eusphyrus vasconicus (Hoffmann & Tempere, 1954) e ne tracciano la storia nomenclaturale. Sono caratterizzate le stazioni di raccolta ed elencati i sistemi di cattura utilizzati. Il taxon viene illustrato con fotografie e inserito in una tabella dicotomica pe...
Article
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Are described and illustrated two new Italian species of Ceutorhynchus. The first of them, C. apenninus n. sp. from central Italy, collected on the montane crucifer Isatis allionii P. W. Ball., is close to C. peyerimhoffi Hustache from Spain, Italy and Algeria, also living on Isatis. The second, C. magnanoi n. sp. from southern Italy is very close...
Article
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Four new species of Palaearctic Ceutorhynchinae are described: Thamiocolus rufitarsis sp. n. from Iran close to T. comptus Colonnelli, 1997 from Southern Turkey; Datonychus terminassianae sp. n. from Northern China and Datonychus kostali sp. n. from Iran both rather isolated among their genus, and Datonychus gultekini sp. n. from Northeastern Turke...
Article
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A checklist of Curculionoidea is given from southern Turkey (Adana, Osmaniye and Hatay provinces). The known localities and ecological notes on each of them were reported based on literature records and on recently collected material. Lixus farinifer Reitter is newly recorded from Turkey, and 35 species (2 Rhynchitidae, 5 Apionidae and 28 Curculion...
Article
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Otiorhynchus (Sulcorhynchus) emrei Avgın & Colonelli sp. n. is described from the Amanos mountains (Hatay), southern Turkey. This is the first record of the subgenus Sulcorhynchus Magnano, 1998 outside the Caucasian area.
Article
Urometopus angelinii sp. nov. from northern Turkey (Ordu province) is described and illustrated. Description and illustration of the male of U. anatolicus Smreczynski, 1977 with new locality records are also given. Omias imereticus Reitter, 1888, is here designated as the type species of Urometopus Formanek, 1904. Taxonomy and position of this genu...
Article
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The Curculionoidea of the families Anthribidae, Rhynchitidae, Apionidae, Nanophyidae, Brachyceridae, Curculionidae, Erirhinidae, Raymondionymidae, Dryophthoridae and Scolytidae from the Maltese islands are reviewed. A total of 182 species are included, of which the following 51 species represent new records for this archipelago: Araecerus fascicula...
Article
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Sono riportati i dati relativi a una prima parte di Curculionoidea di Sicilia conservati nella collezione di Francesco Vitale. Vengono prese in esame le specie di Brachyceridae, Curculionidae (Bagoinae, Baridinae, Ceutorhynchinae, Conoderinae, Cossoninae, Curculioninae, Cyclominae, Lixinae, Molytinae, Phytonominae), Dryophthoridae ed Erirhinidae. S...
Article
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Torneuma curtulum vastum Stüben, 2007 (Curculionidae) is raised to species level (stat. rev.). Choragus aurolineatus Abeille de Perrin, 1893, Melanopsacus grenieri (C. Brisout, 1867) (Anthribidae), Hemitrichapion pavidum (Germar, 1817), Holotrichapion barbirostre (Hoffmann, 1952) (Apionidae), Ceutorhynchus resedae (Marsham, 1802), Tychius cuprifero...
Article
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Seventy one species of Tychiini Tychiina and Mecinini collected in South Africa are here reported, among which seven are new: Tychius densevestitus Caldara n. sp.; Tychius saetosipennis Caldara n. sp.; Sibinia incompta Caldara n. sp.; Gymnetron claviger Caldara n. sp.; Gymnetron globiscapus Caldara n. sp.; Gymentron filum Caldara n. sp.; Gymnetron...
Article
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The beetle Thamnurgus euphorbiae Küster, whose larvae feed in the inner stems of Euphorbia characias L., found in Italy, was selected as a candidate agent for the biological control of Euphorbia esula-virgata Waldstein and Kitaibel in the United States. Its adaptation to and survival on several E. esula-virgata ecotypes from North America justified...
Article
Il 3 luglio 2009, nel giorno del suo ottantaquattresimo compleanno, Luigi Magnano ci ha lasciati per sempre, dopo una non lunga ma penosa malattia che lo aveva costretto da alcuni mesi ad interrompere la sua intensa attività.
Article
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Six new species of the genus Heisonyx Marshall, 1947 from South Africa (H. barclayi sp. nov., H. danielssoni sp. nov., H. fuscus sp. nov., H. giustocaroli sp. nov., H. hexarthrum sp. nov., and H. jelineki sp. nov.) are described, illustrated and keyed. The genus is redescribed and its taxonomic position is briefly discussed. Heisonyx and Lalagetes...
Article
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A survey conducted in central Italy found 126 species of phytophagous insects from five orders and 25 families on dyer’s woad, Isatis tinctoria L., a biennial or short-lived perennial. About 75% of the species found attacking this weed were polyphagous, 20% were restricted to the family Brassicaceae, and only 5% were restricted to the genus Isatis....
Chapter
These proceedings contain the full-length papers and abstracts of papers presented at the symposium. Subjects covered include: ecology and modelling in biological control of weeds; benefits, risks and cost analysis of biological weed control; target and biological control agent selection; pre-release specificity and efficacy testing; regulations an...
Article
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We collected specimens of Paramecops sinaitus (Pic, 1930) (Curculionidae: Molytinae) from south Sinai in Egypt, which enabled us to make the first complete description of this species. We also include some taxonomic remarks on the genus. Paramecops solenostemmatis (Peyerimhoff, 1930) is a synonym of Paramecops sinaitus. We propose the new combinati...
Chapter
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A b s t r a c t : The Arabian species of the genus Ocladius Schönherr, 1825 are revised. The following new species are described: O. abdeldayemi n. sp. (type locality: Egypt, Sinai, Wadi El Hage), O. aidhabensis n. sp. (type locality: Egypt, Marsa Halaib), O. amharicus n. sp. (type locality: Ethiopia, Dire Dawa), O. eremorum n. sp. (type locality:...
Article
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New data are provided on distribution and ecology of the rare Liparus mariai, endemic of central Apennines, Italy. The weevils have been found while feeding on Meum athamanticum, a montane plant of the family Apiaceae very probably its host. Notes about the correct spelling of the weevil name are included
Article
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Ranunculiphilus pseudinclemens (DIECKMANN, 1969) is recorded for the first time from Poland and Slovakia. Short notes on taxonomy, ecology and distribution of this species, as well as a key to all members of the genus Ranunculiphilus DIECKMANN, 1969 are also given.
Article
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Graduate School of Bioresource and Bioenvironmental Sciences, Kyushu University The genus Ceutorhynchoides Colonnelli (Ceutorhynchini, Ceutorhynchinae) is recorded for the first time from Japan, with descriptions of two new species: C. squamatus sp. noc. (Ryukyus) and C. styracis sp. nov. (Honshu, Shikoku, and Kyushu). In addition, Hainokisaruzo ge...
Article
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Ceutorhynchus varius Rey, 1895, formerly considered to be a variety of C . hirtulus Germar, 1824, is revealed to be a valid species and redescribed. It lives on Arabidopsis thaliana (L.) Heynh. (Brassicaceae) and is recorded from France, Poland, Czech Republic, Slovakia, Austria and Bulgaria. Lectotypes are designated for C. varius Rey and C. hirtu...
Article
A list of Curculionoidea (Nemonychidae, Anthribidae, Rhynchitidae, Attelabidae, Brentidae, Apionidae, Nanophyidae, Brachyceridae, Curculionidae, Erirhinidae, Raymondionymidae, Dryoph-thoridae, Scolytidae, Platypodidae) thus far known from Italy is drawn up, updating that by Abbazzi et al. published in 1995. Distributional data of each species are g...
Article
Coeliastes rustemi sp. n., the third species of the genus Coeliastes Weise from Turkey is described. The new species differs from the congeners in the larger size, denser pubescence formed by very large white scales, less distinct elytral pattern, more strongly rounded sides of pronotum and apical elytral prominences, and more coarse sculpture of e...
Article
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The classification of Ochthebius (Calobius) is revised using isozyme analysis combined with a preliminary morphological study of population samples from 69 Mediterranean and Macaronesian localities. Seven taxa are recognized, including 3 new taxa, provisionally designated as Ochthebius species A from Italy, O. heeri subspecies A from the western Ca...
Article
Ceutorhynchus trimaculatus (F.) and Trichosirocalus (= Ceuthorhynchidius auctt. nee du Val) horridus (Panzer) are distributed in central and southern Europe and the Caucasus area of the USSR. C. trimaculatus is also present in North Africa. Host plants of adult C. trimaculatus are Carduus and Cirsium spp., and host plants of adult T. horridus are C...
Article
RIASSUNTO Vengono descritti Brachycerus gilvipes n. sp. ed Hypohypurus litoralis n. sp., entrambi raccolti sul litorale della Somalia durante le ricerche organizzate dal Centro di Studio per la Faunistica ed Ecologia Tropicali del Consiglio Nazionale delle Ricerche.

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