Dmitriy Gimranov

• PhD
• Researcher at Russian Academy of Sciences

The paper deals with some finds from the Imanai cave in Bashkiria, Southern Ural, which provide evidence of cave bear hunting by the Middle Palaeolithic inhabitants of the region. The excavations conducted in 2015–2016 yielded a small cave bear (Ursus rossicus) cranium with a hole in the right temporal bone. Judging by its morphology and the absenc...

The article publishes for the first time the findings of three structures from the time of the Golden Horde, which were studied in 2019 at kurgan 1 of the “Plavni” group near the village of Glinoe. Two of these structures (burials no. 8 and 20) were pits with ledges, while the third one (pit no. 1) was a round pit. Notably, the skeleton of an adult...

Twenty-seven Pachycrocuta brevirostris coprolites from Taurida Cave (Early Pleistocene) were studied. Eggs of parasitic worms were found in 6 of them (22.2%). Eggs of Trematoda species were identified among them, as were eggs of Taenia, Toxocara, and Capillaria. Toxocara eggs were the most common; they were found in 15% of the coprolites. This is t...

The cranial and mandibular fragments of the medium-sized canid from the Early Pleistocene locality of Taurida cave (about 1.8–1.5 Ma) in Crimea are described and included in the hypodigm of the species Canis etruscus Forsyth Major, 1877, a common member of the Late Villafranchian fauna of the Circum-Mediterranean region. This species is reported fo...

The article describes the first find of a bird from the Paleogene of Siberia. A tibiotarsus fragment from the Eocene Tavda Formation of the Tyumen Region (Western Siberia) is referred to Procellariiformes. The described bird is morphologically closer to Procellariidae, but comparable in size to albatrosses (Diomedeidae) and, apparently, represents...

A mandible fragment of Acinonyx pardinensis (Croizet et Jobert, 1828) is described from the Early Pleistocene locality in the Taurida cave (Crimea, Late Villafranchian, about 1.8–1.5 Ma). This is the first discovery of the genus Acinonyx in the Pleistocene of Crimea. Along with other felids, such as Homotherium, Megantereon, Panthera, Lynx, and Pum...

Описано новое местонахождение с остатками Crocuta spelaea (Goldfuss, 1823) в пещере Тип-Тугай (52°59'28.6”N, 57°00'22.3”E) на Южном Урале. На основе анализа видового состава фауны крупных и мелких млекопитающих отложения, содержащие остатки гиены, датируются морской изотопной стадией 3 (MIS 3). Исследование возрастного состава остатков гиены и хара...

The frozen mummy of a saber-toothed cat Homotherium latidens cub was found in the Upper Pleistocene permafrost of the Yedoma deposits on the Badyarikha River (the right tributary of the Indigirka River) in the northeast of Yakutia. For the first time in the history of paleontology, the external appearance of an extinct mammal species that has no an...

The frozen mummy of the large felid cub was found in the Upper Pleistocene permafrost on the Badyarikha River (Indigirka River basin) in the northeast of Yakutia, Russia. The study of the specimen appearance showed its significant differences from a modern lion cub of similar age (three weeks) in the unusual shape of the muzzle with a large mouth o...

The cranial and mandibular fragments of the medium-sized canid from the Early Pleistocene locality of Taurida cave (about 1.8-1.5 Ma) in Crimea are described and included in the hypodigm of the species Canis etruscus Forsyth Major, 1877, a common member of the Late Villafranchian fauna of the Circum-Mediterranean region. This species is reported fo...

A mandible fragment of Acinonyx pardinensis (Croizet et Jobert, 1828) is described from the Early Pleistocene locality in the Taurida cave (Crimea, Late Villafranchian, about 1.8–1.5 Ma). This is the first discovery of the genus Acinonyx in the Pleistocene of Crimea. Along with other felids, such as Homotherium, Megantereon, Panthera, Lynx, and Pum...

The structure of the second upper molar (M2) of brown (Ursus arctos) and white (U. maritimus) bears was studied, with 9 polymorphic characters identified, and their descriptions given. A west to east geographic pattern was discovered in the variability of the lingual cingulum M2 in continental U. arctos. In island populations of U. arctos, there is...

The Tip-Tugai Cave (52°59′28.6″ N, 57°00′22.3″ E) is described as a new site with cave hyena Crocuta spelaea fossils. The bone-bearing layer was dated to marine isotope stage (MIS) 3 based on the speciescomposition of the fauna of large and small mammals. Finds of abundant remains of cave hyenas of differentages and bones of large herbivores with h...

The article describes the first find of a bird from the Paleogene of Siberia. A fragment of tibiotarsus from the Eocene Tavda Formation of the Tyumen Region (Western Siberia) is assigned to Procellariiformes. The bird is morphologically closer to Procellariidae, but comparable in size to albatrosses (Diomedeidae) and is assumed to represent the ste...

The present work describes for a first time a fused fracture of the metacarpals of the medium-sized felid, Lynx issiodorensis, the putative ancestor of all Eurasian extant lynx species. The studied remains, four metacarpals from the same individual, were unearthed from the Early Pleistocene of the Taurida cave (Crimean Peninsula) and studied throug...

The Dongal settlement is located 208 km southeast of the city of Karaganda. Based on his materials, a special type of ceramics was identified, called Dongal. Pottery of this type has features that connect it both with the previous Sargary-Alekseevsky culture of the final Bronze Age, and with the monuments of the early Iron Age. This establishes a g...

A mass burial of complete and almost complete cattle and small ruminant skeletons found in a pit within the Early Iron Age settlement of Abylai in Central Kazakhstan (49°15'N, 75°07'E) has been studied. The results of archaeological research and archeozoological contexts have shown that the animals’ burial was a single event. The aim of the work wa...

Seven species of red-toothed shrews (Sorex araneus, Sorex isodon, Sorex tundrensis, Sorex caecutiens, Sorex minutus, Sorex minutissimus, Neomys fodiens), white-toothed shrews (Crocidura sp.), moles (Talpa sp.) and hedgehogs (Erinaceus sp.) were found in Late Pleistocene and Holocene deposits of the southern part of the western slope of the Ural Mou...

An Erratum to this paper has been published: https://doi.org/10.1134/S0012496623050010

An isolated tooth, of morphology unusual for cave bears, is described from Late Pleistocene deposits (MIS 3) of Pobeda Cave (54.1000° N, 56.5100° E) in the Urals. Analysis of ancient DNA showed that it belongs to the Ural cave bear Ursus kanivetz, which was widespread in the Pleistocene in the Urals. The fossil find was identified as an anomalous u...

The dentary of Mustela strandi Kormos, 1934 is described from the Lower Pleistocene deposits (Late Villafranchian, 1.8–1.5 Ma) of the Taurida cave in Crimea. It is the first finding of M. strandi in Russia. This extinct mustelid species is rarely found in the Lower and Middle Pleistocene of Central Europe.

The mandibles of two individuals of Mustela palerminea (Petenyi, 1864) are described from the Lower Pleistocene deposits of the Taurida cave in Crimea (Late Villafranchian, 1.8-1.5 Ma). This extinct mustelid species was a typical representative of the Villafranchian faunas of Europe. It is the first record of M. palerminea in Russia.

The dentary of Mustela strandi Kormos, 1934 is described from the Lower Pleistocene deposits (Late Villafranchian, 1.8-1.5 Ma) of the Taurida cave in Crimea. It is the first finding of M. strandi in Russia. This extinct mustelid species is rarely found in the Lower and Middle Pleistocene of Central Europe.

The mandibles of two individuals of Mustela palerminea (Petenyi, 1864) are described from the Lower Pleistocene deposits of the Taurida cave in Crimea (Late Villafranchian, 1.8–1.5 Ma). This extinct mustelid species was a typical representative of the Villafranchian faunas of Europe. It is the first record of M. palerminea in Russia.

The tiger (Panthera tigris) is a charismatic megafauna species that originated and diversified in Asia and probably experienced population contraction and expansion during the Pleistocene, resulting in low genetic diversity of modern tigers. However, little is known about patterns of genomic diversity in ancient populations. Here we generated whole...

The present work describes the dentognathic remains of Ursus etruscus Cuvier, 1823 from the recently discovered Taurida cave in central Crimea at the north Black Sea area. The bone-bearing layer of Taurida cave corresponds to the Psekupsian Faunal Assemblage of Eastern Europe and to the Late Villafranchian of Western Europe (ca. 1.8–1.5 Ma). Here,...

A maxillary fragment of the extinct hyena Chasmaporthetes lunensis (Del Campana, 1914) is described from the Early Pleistocene locality of the Taurida cave (Crimea, Late Villafranchian, 1.8-1.5 Ma). The species was a typical representative of the Villafranchian faunas of Eurasia. This is the first record of the genus Chasmaporthetes in the Pleistoc...

A maxillary fragment of the extinct hyena Chasmaporthetes lunensis (Del Campana, 1914) is described from the Early Pleistocene locality of Taurida cave (Crimea, Late Villafranchian, 1.8–1.5 Ma). The species was a typical representative of the Villafranchian faunas of Eurasia. This is the first record of the genus Chasmaporthetes in the Pleistocene...

The contrast between the tiger's (Panthera tigris) 2-3 My age and extant tigers' coalescence approximately 110,000 years ago suggests an ancient demographic bottleneck. Here we collected over 60 extinct specimens across mainland Asia and generated whole genome sequences from a 10,600-year-old Russian Far East (RFE) specimen (RUSA21, 8x coverage), 1...

Исследована ископаемая фауна мелких млекопитающих из отложений пещеры Иманай (53°02′ с.ш., 56°26′ в.д.) (Южный Урал, Россия). В фауне преобладают виды открытых биотопов, узкочерепная полевка (Microtus (Stenocranius) gregalis) является доминирующим видом. Выявлено два типа фауны, которые характеризуют ее состав и структуру в конце позднего плейстоце...

A total of 1250 lower first molars (m1) of voles (Arvicolini) were studied from Late Pleistocene deposits (the radiocarbon dates from rodent bones: 17 100 ± 50 IGANAMS-9117; 13 255 ± 60 IGANAMS-9116) of the Imanay Cave (southern Urals, 53°02' N, 56°26' E). Of these, 24 m1 of voles of the subgenus Stenocranius were found to have broadly connected tr...

Since the pioneering works of Björn Kurtén from the middle of XX century, the Eurasian Early Pleistocene species Ursus etruscus is considered the putative ancestor of both cave and brown bears. However, in the last two decades the remains of the late Early Pleistocene has been in a hotly phylogenetic and taxonomic debate and several authors include...

The microwear of the non-occlusal surface of incisors (I1, I2) of the small cave bear (Ursus ex gr. savini-rossicus) and Ural cave bear (Ursus kanivetz) from the Pleistocene of the Middle and South Urals is analyzed and compared. Qualitative characteristics of incisor microwear have been shown to be different in these species. In the small cave bea...

Coprolites of the hyena Pachycrocuta brevirostris from the Lower Pleistocene (Upper Villafranchian) of Taurida Cave (Crimea) were studied. One of the three hyena coprolites contained helminth eggs. These eggs were assigned to Toxocara sp. based on their size and morphology. Toxocariasis was evidently a very common infestation among extinct hyena sp...

По материалам поселения раннего железного века Абылай в Центральном Казахстане было исследовано массовое захоронение полных и почти полных скелетов домашних животных. Данная работа призвана представить результаты проведенного анализа по этим материалам. Целью работы является определение причин формирования скопления костей. Для этого были проведены...

The first record of the hyena Chasmaporthetes lunensis (Del Campana, 1914) in Crimea (Tavrida cave, Early Pleistocene)

In the rich vertebrate fauna of Imanay Cave the abundant material of the small-sized cave bears was originally assigned to the taxon Ursus savini. Teeth and metapodials of statistical amounts were compared with other cave bear faunas and the taxonomic position was determined through morphological and metric analyses. The size of teeth and metapodia...

Recent field surveys performed in the newly-discovered Taurida Cave (Crimea, Russia, Late Villafranchian, 1.8–1.5 Ma) enabled us to unearth dentognathic remains of the dirk-toothed cat Megantereon. Here we describe in detail the recovered remains further comparing it anatomically and biometrically with coeval Eurasian and African sites. The perform...

The Early Pleistocene vertebrates assemblage from the Taurida cave in the central Crimea (Belogorsky district, Zuya village) includes two saber-toothed cat species, Homotherium crenatidens and Megantereon adroveri.

The cranial and mandibular remains of two adult individuals of Lynx issiodorensis (Croizet et Jobert, 1828) are described from the Early Pleistocene locality of the Taurida cave (Crimea, Late Villafranchian, 1.8-1.5 Ma). This lynx species was a typical representative of the Villafranchian fauna of the Eastern Mediterranean. A high craniological var...

The fossil faunas of mammals from layer 10 of excavation V of the Ignatievskaya Cave, Southern Urals (54°53′ N, 57°46′ E) are described. Data on the species composition of the faunas from different horizons, as well as on the morphology and size of teeth and bones of bats, insectivorous mammals, lagomorphs, and rodents, are presented. The complexes...

The cranial and mandiblular remains of two adult individuals of Lynx issiodorensis (Croizet et Jobert, 1828) are described from the Early Pleistocene locality of the Taurida cave (Crimea, Late Villafranchian, 1.8–1.5 Ma). This lynx species was a typical representative of the Villafranchian fauna of the Eastern Mediterranean. A high craniological va...

A mandible fragment and four isolated teeth of the fossil foxes, Vulpes alopecoides (Del Campana, 1913), Vulpes cf. vulpes (Linnaeus, 1758) and Vulpes sp., are described from the Early Pleistocene locality of Taurida cave (Crimea, Late Villafranchian, 1.8-1.5 Ma). Based on the size and morphological features, a mandible fragment and two M1 are attr...

A mandible fragment and four isolated teeth of the fossil foxes, Vulpes alopecoides (Del Campana, 1913), Vulpes cf. vulpes (Linnaeus, 1758) and Vulpes sp., are described from the Early Pleistocene locality of Taurida cave (Crimea, Late Villafranchian, 1.8–1.5 Ma). Based on the size and morphological features, a mandible fragment and two M1 are attr...

An Erratum to this paper has been published: https://doi.org/10.1134/S0012496621050112

There are many Middle-Upper Pleistocene outcrop sections in the Lower Volga region (Caspian Lowland, Russia) in between cities of Volgograd and Astrakhan. The present study focuses on the Seroglazovka locality, which is among the most famous key sites within the Lower Volga region. The Seroglazovka locality is of great regional interest, because of...

The lower molar (m1) of cave bears from Late Pleistocene localities of the Urals was studied employing the methods of traditional morphometry and geometric morphometrics. On the basis of the size and shape variation of m1, the small cave bear (Ursus ex gr. savini-rossicus) was found to have been a part of the faunas from the caves Skazka, Viasher,...

Morphometric and morphotypic variability of the cave bear lower incisors from two different geographic regions (Caucasus and Urals), different stratigraphic periods (Middle and Late Pleistocene), and bearing different mitochondrial haplogroups (kudarensis (Baryshnikov, 1985) and ingressus Rabeder, Hofreiter & Withalm, 2004) was studied. Urals Ursus...

Imanay Cave is located in the Southern Urals (53°02' N, 56°26' E), at 420 m.a.s.l. A 9.5 m2 trench was exca-vated in the grotto in the inner part of the cave to examine the sediments. The accretion thickness was 1.2 m. The taphocoenosis of the Imanay Cave is of the Pleistocene age and contains about 10,000 specimens of bone re-mains of large mammal...

The Bering Land Bridge (BLB) last connected Eurasia and North America during the Late Pleistocene. Although the BLB would have enabled transfers of terrestrial biota in both directions, it also acted as an ecological filter whose permeability varied considerably over time. Here we explore the possible impacts of this ecological corridor on genetic...

Remains of a cave bear were studied from a new locality in the Prokoshev Cave in the Middle Urals (58°13´ N, 58°12´ E). Bones from all regions of the skeleton are present, bones are intact and without traces of human or animal activity. They all belong to the cave bear (Ursus kanivetz Vereshchagin, 1973). An AMS radiocarbon date of 53 375 ± 765 BP,...

The dental remains of a giant hyena Pachycrocuta brevirostris (Gervais, 1850) from the Early Pleistocene locality of the Taurida cave (Crimea, Late Villafranchian, 1.8-1.5 Ma) are described. This species was a typical representative of the Villafranchian fauna of the Eastern Mediterranean. The Taurida cave was occasionally used by hyenas and other...

This paper describes the maxillaries of adult specimens of saber-toothed cats Homotherium crenatidens (Fabrini, 1890) and Megantereon sp. from the Taurida Cave (Crimea, Late Villafranchian; age, 1.8–1.5 Ma). We discuss the morphometric variability in the morphology of the upper carnassials and the length of the С1–Р3 diastema.

The dental remains of a giant hyena Pachycrocuta brevirostris (Gervais, 1850) from the Early Pleistocene locality of the Taurida cave (Crimea, Late Villafranchian, 1.8–1.5 Ma) are described. This species was a typical representative of the Villafranchian fauna of the Eastern Mediterranean. The Taurida cave was periodically used by hyenas and other...

Впервые получены результаты комплексных минералого-геохимических исследований ископаемых ко-стей плейстоценовых млекопитающих из пещеры Иманай (Южный Урал) с использованием оптической и сканирующей электронной микроскопии, статистического объемометрического метода определения нанопористости, термического, рентгеноструктурного и рентгено-флюоресцент...

Introduction. Imanai-1 Cave is a new monument of the Middle Paleolithic in the Southern Urals. It was discovered by the authors in 2009 and is located in the west of the Ural mountain system, in the interfluve of the Belaya and Nugush Rivers, on the border of the mountain-forest and steppe zones. Goals. The paper aims to introduce preliminary resul...

Martes is one of the most widely spread genera and probably has the longest evolutionary history among the living Carnivora, yet its fossil record is incomplete, especially those related to the crown group of Martes. Here we describe a fossil species, Martes crassidens sp. nov. from the Jinyuan Cave, Luotuo Hill, Dalian Puwan Economic Zone, Liaonin...

We clarify the contribution of gene introgression in the recovery of the sable Martes zibellina, which has undergone significant population declines by the end of the nineteenth century. To verify the contribution of interspecific gene introgression, variation among the sable and the pine marten Martes martes was investigated utilizing restriction...

A fragment of a bear skull with partially preserved dentition is descibed from the Lower Pleistocene deposits of the Taurida cave (Crimea). The presence of P1–P3, the structure of P4, and sizes of the cheek teeth enable the identification of the specimen as Ursus etruscus Cuvier, 1823. The new find is the first in Crimea and the entire Russia and i...

В нижнеплейстоценовых отложениях пещеры Таврида (Крым) найден фрагмент черепа медведя с частично сохранившимися зубными рядами. Присутствие Р1–Р3, строение Р4 и размеры щечных зубов позволяют отнести находку к Ursus etruscus Cuvier, 1823. Новая находка является первой для территории Крыма и России в целом и представляет большой интерес в связи с ре...

Analysis of mammalian species composition has been performed for eight archaeological sites on the Ustyurt plateau. They date to the range from the middle Subboreal to the middle Subatlantic. The modern species composition of large mammals formed by the end of the Middle Holocene (Subboreal 2) and did not change until the early 20th century. The un...

Enamel macro- and microstructure has been studied in the teeth of Merck's rhinoceros (Stephanorhinus kirchbergensis Jäger, 1839), woolly rhinoceros (Coelodonta antiquitatis Blumenbach, 1799), and rhinoceroses from the Tetyukhinskaya (44°35' N, 135°36' E) and Sukhaya (43°09' N, 131°28' E) caves in southern Primorye. The teeth from the caves were ide...

Дано описание ископаемых остатков двух видов винторогих антилоп Gazellospira torticornis (Aymard, 1854) и Pontoceros ambiguus Vereschagin, Alexejeva, David et Baigusheva, 1971, найденных в 2018 г. в нижнеплейстоценовых отложениях пещеры Таврида в Центральном Крыму. Эти находки – первое свидетельство присутствия этих таксонов на территории полуостро...

The composition of large mammal fauna from the Imanay Cave in the Southern Urals (53°02' N, 56°26' E) is described in the present study. The paper aims to provide an preliminary description of the large mammal remains from the Imanay Cave in order to establish their taxonomic status, geological age and to detect the factors that led to accumulation...

The article describes fossil remains of two spiral-horned antelope species Gazellospira torticornis (Aymard, 1854) and Pontoceros ambiguus Vereschagin, Alexejeva, David et Baigusheva, 1971 which were found in 2018 in the Lower Pleistocene sediments of the Taurida Cave in Central Crimea. These finds are the first evidence of the occurrence of these...

Berezov is one of the first Russian towns in the north of Western Siberia (Khanty-Mansi Autonomous Area). This article is aimed at describing the household structure, the commercial activities of the population inhabiting the posad of the town on the basis of animal bone remains, as well as at determining the degree of integration of the Russian an...

A study of the association of vertebrates from the Pleistocene deposits of the Taurida karst cave discovered in 2018 in the central Crimea (Belogorsky district, Zuya village) showed its correlation with the Late Villafranchian faunas of the Eastern Mediterranean and an approximate age of 1.8–1.5 Ma.

We describe a vertebrate assemblage from the Pleistocene deposits of the Taurida karst cave discovered in 2018 in central Crimea (Zuya village, Belogorsk raion). The assemblage is correlated with Late Vil-lafranchian faunas of the Eastern Mediterranean and has an approximate age of 1.8-1.5 Ma.

For the first time in the south of the Russian Far East in the Late Pleistocene cave deposits, fragments of the giant flying squirrel of the genus Petaurista were discovered. Petaurista tetyukhensis n. sp. is described based on a fragment of the upper jaw with two teeth and five isolated teeth from two cave locations. The main differences between t...

The fossil faunas of mammals from layer 10 of excavation V of the Ignatievskaya Cave, southern Urals (54°53′ N, 57°46′ E) is described. Data on the species composition of the faunas from different horizons, as well as on the morphology and size of teeth and bones of bats, insectivorous mammals, lagomorphs and rodents are presented. The complexes of...

The article describes the species composition of bone remains of the Pleistocene age collected on the shores of Lake Yambuto located on the Gydan Peninsula (71°N, 79°E). 812 bones were collected from 4 sites on the west shore, and 603 bones were collected from 2 sites on the east shore. The bones belonged to the following species: Lepus timidus, Ca...

Morphometric and morphotypic variability of the cave bear upper incisors from two different geographic regions (Caucasus and Urals), different stratigraphic periods (middle and late Pleistocene), and bearing different mitochondrial haplogroups (kudarensis and ingressus) was studied. The specific diet of the cave bears, i.e. hard vegetables, led to...

The vertebrate fauna from the cave deposits in Imanai Cave in the Southern Urals (53°02′ N, 56°26′E) has been studied. It contains 715 bones that belonged to at least 11 individuals of fossil lion (Panthera (Leo) ex gr. fossilis-spelaea). It has been established that this is one of the largest Eurasian burial sites of fossil lions. The bones were a...

Исследовали новую выборку остатков млекопитающих из плейстоценовых отложений Игнатиевской пещеры на Южном Урале (54°53′ с.ш., 57°46′ в.д.). Среди остатков грызунов определили зубы обыкновенной белки (Sciurus vulgaris) и лесной сони (Dryomys nitedula). Коэффициент дифференциации эмали (SDQ) зубов водяной полёвки соответствовал виду Arvicola terrestr...

Фауна крупных млекопитающих плейстоцена в пещере Иманай на Южном Урале.

The study deals with a new sample of the mammalian fossils from the Pleistocene deposits of the Ignatievskaya Cave in Southern Urals (54°53' N, 57°46' E). Among the rodent fossils, the teeth of red squirrel (Sciurus vulgaris) and forest dormouse (Dryomys nitedula) have been identified. The enamel differentiation quotient (SDQ) of the water vole tee...

In the late Miocene deposits in the Taralik-Cher locality (Tuva Republic), fossil remains of a giant mustelid of the genus Eomellivora Zdansky, 1924 have been found. This finding is the first reliable evidence of the Eomellivora inhabiting the modern Russia. Teeth of the Eomelliovra from Taralik-Cher are similar in size to those of E. wimani and E....

The paper describes the bone animal remains from the excavations of the mansion of Balandins merchants, which reflect the economic culture of the Russian population of the Yeniseisk in the 17th – 19th centuries. The analysis of species composition and ratio of animal remains were carried out, and methods for cutting and utilization of domestic anim...