David Rider

• Doctor of Philosophy
• Professor (Full) at North Dakota State University

The known distribution and habitat information of the three described species of the genus Thaumastella Horváth, 1896 (Hemiptera: Heteroptera: Pentatomomorpha: Pentatomoidea: Thaumastellidae) are reviewed. In addition a fourth, undescribed species is reported from South Africa (Western Cape). Thaumastella aradoides Horváth, 1896, is recorded for th...

A new species in the monotypic genus Nocheta Rolston, 1980, N. alfordi new species (Hemiptera: Heteroptera: Pentatomidae: Pentatominae: Pentatomini), is described from specimens collected in Bolivia, Brazil, and Peru. The new species is described, illustrated, and compared to N. adda Rolston, 1980. Additional information on N. adda is also provided...

We provide corrections to 17 recent papers (published since 2013). The following new combination is proposed: Dabessus indicus Kushwaha and Jahan, 2023 = Neojurtina indica (Kushwaha and Jahan, 2023), new combination. The following corrections are provided: Miridae: Tytthus chinensis sensu Al-Anbaki et al. (2023) = Maurodactylus nigrigenis (Reuter,...

the monotypic oriental genus Vitruvius Distant, 1901 (hemiptera: heteroptera: Pentatomidae: Pentatominae), having an intricate and confusing taxonomic history, is revised. A new species, Vitruvius dobby roca-Cusachs, rider & Kment new species, is described from Cambodia and thailand. Notes on the hypothesized classification of the genus based on bo...

The following species are recorded from the Comoro Islands for the first time: 12 species of Pentatomidae: Pentatominae—Anoano pronotalis Cachan, 1952 (from Mayotte), Antestiopsis clymeneis cf. galtiei (Frappa, 1934) (Mayotte), Bagrada hilaris (Burmeister, 1835) (Mayotte), Bathycoelia rodhaini Schouteden, 1913 (Mayotte), Coquerelia ventralis Horvát...

The genus Eurypharsa Stål 1873 is reviewed. Tingis circumdata Blanchard 1842 [not 1847] has priority over Tingis nobilis Guérin-Méneville 1844 and is transferred to Eurypharsa. Eurypharsa phyllophila Drake 1922 is subjectively synonymized with Eurypharsa circumdata [new combination, revised status]. Herein we designate lectotypes for Tingis circumd...

The Enicocephalomorpha of North Dakota is documented with the report of the unique-headed bug, Systelloderes biceps (Say, 1832) from North Dakota for the first time. We also discuss the geographic distribution of S. biceps and Systelloderes culicis (Uhler, 1892), provide color photographs, provide diagnoses for the separation of these species from...

Euschistus Dallas is distributed throughout the Western Hemisphere and is composed of 67 species in three subgenera: Euschistomorphus Jensen-Haarup, Lycipta Stål, and the nominate subgenus. Euschistus (Euschistus) includes several economically important species. Due to the lack of a phylogenetic analysis for the subgenus Euschistus, its groupings a...

Descripción de la ninfa V de Tolono confusus Carvajal, Rider y Faúndez, 2015 Abstract. The fifth instar nymph of Tolono confusus Carvajal, Rider & Faúndez, 2015 is described and illustrated based on two specimens from Napo Province, Ecuador. The need for further work on South American pentatomoid immatures stages is briefly discussed. Resumen. Se d...

The genus Halyomorpha Mayr, 1864 (Hemiptera: Heteroptera: Pentatomidae: Pentatominae: Cappaeini) was established based on a single species, Halys timorensis Westwood, 1837 (currently a junior subjective synonym of Halyomorpha picus (Fabricius, 1794)). The examination of the voucher specimens identifi ed as Halyomorpha timorensis by Mayr, a syntype...

The species of Pentatomidae known to occur north of Mexico, comprising 223 species in 68 genera, are enumerated with taxonomic notes and updated and annotated distributions. Included in this update are 126 new state records reported for 62 pentatomid species in 30 genera. The copious annotations in these distributions and attendant bibliography ser...

A new synonymy in South American Carpocorini

A new genus, Graziasternum gen. nov., with two new species, Graziasternum graziae sp. nov. and Graziasternum joceliae sp. nov. are described. According to external thoracic and genital structures, this new genus is considered to be the closest extant relative known to the genus Placosternum Amyot & Serville. Discussion of its possible phylogenetic...

The new Neotropical hyaliodine Colimacoris joceliae Kim & Jung, sp. nov., is described from Panama. Morphological information, including a description and diagnosis of the new species is presented with photographs. A key for the identification of the species of Colimacoris is also provided.

The genus Acrophyma Bergroth currently contains two species, the Andean A. cumingii (Westwood) and the Neotropical A. bicallosa (Stål). The examination of types and new material revealed that among the Neotropical species the Brazilian population differs from the Colombian, and the first one belongs to a new species in this group. A morphological a...

Two new species of Tingidae are described from Panama: Acanthotingis deltoides sp. nov. and Stephanitis joceliae sp. nov. Descriptions and diagnosis for both Acanthotingis Monte and Stephanitis Stål are provided, and a key is given for the separation of the two known species of Acanthotingis. This is the first record of both genera from Panama. A c...

The pentatomids (Hemiptera: Heteroptera) are the third most speciose family within the Heteroptera or the true bugs. The family occurs worldwide and comprises around five thousand valid species within 950 genera. Padaeus Stål belongs to a complex of other genera of Carpocorini Mulsant and Rey related to Euschistus Dallas. These genera present simil...

Pentatomidae is the third largest family of true bugs, comprising over 40 tribes. Few tribes have been studied in a phylogenetic context, and none of them have been examined using molecular data. Moreover, little is known about the evolution of key morphological characters widely used in taxonomic and phylogenetic studies at multiple levels. Here,...

This paper presents the first record of Chinavia acuta in the Cape Verde archipelago.

Resumen: Se analizan los ejemplares tipo de géneros monotípicos y poco conocidos de Acanthosomatidae, Cochabambia Pirán (Ditomotarsinae) y Tunaria Pirán (Blaudusinae: Blaudusini) de Bolivia. Se redescriben en base a caracteres conocidos y nuevos y se discute su posición taxonómica. Tunaria es considerado sinónimo menor de Blaudus Stål, resultando l...

Sibaria Stål is a genus of Pentatomidae occurring exclusively in the Neotropical region. Since its description, little information has been added to the knowledge of the taxon. The hypothesis of monophyly of Sibaria was recently supported by a phylogenetic analysis based on morphological and molecular data. The purpose of this study is to provide a...

Pentatomidae is the fourth largest family of true bugs, comprising nine subfamilies and over 40 tribes. Few tribes in the family have been studied in a phylogenetic context, and none of them have been examined using molecular data. Here, we conduct a phylogenetic study of the tribe Chlorocorini (Pentatominae) combining 69 morphological characters a...

The aeptine Mariomella singularis gen. et sp. nov. is described and illustrated from Western Australia. With this new addition, the Aeptini diversity increases to 21 species in nine genera.

The first records for Aeliosoma weberi Baehr are given since its original description in 1985. This species is newly recorded from the Australian States of South Australia and Western Australia. Additionally, a teratological case of antennal oligomery is described.

The genus Rhyncholepta Bergroth, 1911 (Hemiptera: Heteroptera: Pentatomidae: Pentatominae: Chlorocorini) is redescribed and five species-group taxa are recognized, keyed, their diagnostic characters illustrated, and the distribution reviewed. Among the five taxa, two species and one subspecies are recognized as new: Rhyncholeptagrandicallosagrandic...

The first records for Aeliosoma weberi Baehr are given since its original description in 1985. This species is newly recorded from the Australian States of South Australia and Western Australia. Additionally, a teratological case of antennal oligomery is described.

Pentatomids are highly diverse in the Neotropics, occurring in all biomes. Most of this diversity is distributed in the Atlantic Forest, from which 201 species occur in the state of São Paulo. Here we expand this sample by reporting for the first time the following four stink bug species to the state of São Paulo: Adoxoplatys gallardoi Kormilev, 19...

Se entregan nuevos registros del Asopino Alcaeorrhynchus grandis (Dallas) en Paraguay, en los departamentos de Canindeyú, San Pedro, Cordillera y Alto Paraná. Adicionalmente se describe un caso teratológico correspondiente a una oligomeria tarsal simple.

Resumen.-Se entregan nuevos registros del Asopino Alcaeorrhynchus grandis (Dallas) en Paraguay, en los departamentos de Canindeyú, San Pedro, Cordillera y Alto Paraná. Adicionalmente se describe un caso teratológico correspondiente a una oligomeria tarsal simple. Palabras clave: Asopinae, faunística, teratología, Paraguay. Abstract.-New records of...

Zootaxa 4407 (2): 275-280 Abstract The Aeptini (Pentatomidae: Pentatominae) genus Menestheus Stål, 1868, is redescribed. The original misidentification of the type species for Menestheus is corrected by action of first reviser herein by establishing Paramenestheus nercivus non Dallas, 1851 sensu Stal (1868) = Menestheus cuneatus Distant, 1899 as th...

In this chapter, we provide a taxonomic history for each family and subfamily (if applicable) and, in addition, for the tribes and generic groups within the Pentatomidae, that, perhaps, will give some perception of the taxonomic position of each group. We also discuss the systematic classification within each of these groups and provide general dia...

Kaytuesso flavolateralis gen. and sp. nov. is described from Papua New Guinea. The new genus is remarkable in that the only included species is one of the smallest species known within the Oncomerinae. The relationships among Kaytuesso and the related genera Agapophyta Guerin, Erga Walker and Musgraveia Leston and Scudder are discussed.

The African cluster bug Agonoscelis puberula Stål, 1854 (Hemiptera: Heteroptera: Pentatomidae: Pentatominae: Agonoscelidini), an Afrotropical stink bug previously introduced to the New World, is recorded from Cuba for the first time. The specimen was collected in 1978 and represents the oldest record in America, suggesting the possibility of its in...

In accordance with the rules in the International Code of Zoological Nomenclature, the type species for the genus Edessa Fabricius, 1803 is now recognized to be Cimex antilope Fabricius, 1798 rather than the previously recognized Cimex cer-vus Fabricius, 1787. Edessa antilope is redescribed, as well as the following four species that have in the pa...

Resumen: Se revisa el estatus de Neoacledra Faúndez, 2010 (Heteroptera: Pentatomidae), dándole rango génerico a este taxón. El concepto de Neoacledra es ampliado, el género es redescrito y se describen también dos nuevas especies, Neoacledra viserion sp. nov. y Neoacledra drogon sp. nov., ambas de Chile. Adicionalmente, Jalla flavomaculata Blanchar...

The nymphal instar V of Parajalla sanguineosignata (Spinola, 1852) is described. First formal records with specific locality data are given for Aysen and Magallanes Regions in Chile. San Gregorio becomes the southernmost locality known for this species.

Although the knowledge of South American Acanthosomatidae has increased in the past decade, still little is known about the features or natural history of several genera. Pseudosinopla canaliculus (Reed, 1898) is a rare Acanthosomatid, known so far only from its type. In this work, rediscovery of P. canaliculus in the regions Araucania and Los Rios...

The first record of Copeocoris truncaticornis (Stål, 1865) from Entre Ríos Province, Argentina is given, filling the distributional gap in the country. In addition, a cephalic atrophy is described. Morphological sternal characters are described, including a sulcate mesosternum and basal tubercle of mesosternum. The significance of these characters...

Features: • Presents a brief history of past classifications, a summary of present classification, and speculation on how the classification may evolve in the future • Includes keys for the identification of families and subfamilies of the Pentatomoidea and for the tribes in the Pentatomidae • Explains transmission of plant pathogens and concepts o...

Two teratological cases are described in the pentatomid Hillieria acuminata Distant, 1910, from Southern Australia. The two cases belong cephalic atrophy. Both cases are illustrated and discussed.

Dictyla ainsliei (Drake and Poor) is transferred to the genus Ambotingis Drake and Ruhoff, redescribed, and reported from Costa Rica for the first time. Ambotingis senta (Drake and Hambleton) is also diagnosed, and a key for the separation of the two species is provided.

The brown marmorated stink bug, Halyomorpha halys (Stål, 1855) (Heteroptera: Pentatomidae), is recorded for the first time as established species in Chile, outside of interception facilities in the entrance gates of the country. The new records are based on adult specimens collected in homes in three different places in the city of Santiago, Metrop...

A new genus and new species, Graphorn bicornutus (Heteroptera: Pentatomidae), is described from Chaco Province, Argentina. This new taxon is provisionally placed in the tribe Carpocorini. Relationships of the new genus and close genera are discussed.

New data on the biology and immature stages of H. abbreviatus Uhler, 1872 is provided. The external morphology of the egg and egg batches of H. abbreviatus are illustrated for the first time. Differences between H. abbreviatus and H.limbolarius first instar nymphs are discussed. The data here provided is useful on pest management, as some species o...

A new monotypic Australian genus of Pentatomidae in the tribe Aeptini is described based on an also new species from Western Australia. The new genus is closely related to the genera Hillieria Distant and Pseudaelia Distant, from which can be differentiated by characters of the head, pronotum and legs. A phylogentic analysis is presented for select...

Atheas mimeticus Heidemann is recorded from Canada (Manitoba) for the first time. Known biological information is discussed and an updated host list for the species of the genus is provided.

The painted bug Bagrada hilaris (Burmeister, 1835) (Heteroptera: Pentatomidae) is recorded for the first time from South America. Specimens were collected in Quilicura, Metropolitan Region of Chile. The large amounts of observed and collected specimens, as well as the biological observations, suggest that the species is well established. Identifica...

A short historical review of pentatomoid classifications is presented. This includes notes from the early day describers (Amyot and Serville 1843, Dallas 1851, Stal 1872-1876) through the middle day catalogers (Lethierry and Severin 1893, Kirkaldy 1909) to the later day systematists (Gross 1975, 1976, Linnavuori 1982, Gapud 1991, Schuh and Slater 1...

Introduction: Poecilotoma Dallas, 1851 is an Australian genus of Pentatomidae (Pentatominae: Caropocorini), that currently comprises 4 valid species. This genus is very distinguishable in its general aspect with several ivory ridges on the dorsum, a sulcate mesosternum, and having only 4 antennal segments. Because of this, it has been placed close...

Resumen: Se estudia la identidad de Acledra (Acledra) gregalis Berg, 1878 (Hemiptera: Heteroptera: Pentatomidae). Se aporta una diagnosis revisada, con ilustraciones de caracteres de importancia taxonómica, y se define el concepto de la especie. Una nueva especie, Acledra (Acledra) nazgul sp. nov., es descrita de Argentina. Se discute la relación e...

Resumen: Se analiza la situación del complejo de Acledra (Acledra) albocostata (Spinola, 1852) (Hemiptera: Heteroptera: Pentatomidae). Se concluye que este complejo esta formado por tres subespecies: la subespecie nominal, del centro-sur de Chile; Acledra (Acledra) albocostata carinata Breddin, 1914 n. stat., del sur de Chile; y Acledra (Acledra) a...

Aeliavuori linnacostatus gen. and sp. nov. is described from the Democratic Republic of the Congo. A brief discussion of its tribal placement is provided.

First description of the 5th instar nymph of the Acanthosomatid bug Tolono confusus

The pentatomid bug Mormidea speciosa is recorded for the first time from French Guiana, togther with the first record from the state of Pará, Brazil. The record from French Guiana is the first report of this species outside of Brazil.

The Afrotropical genus-group taxa Dymantis Stal, 1861 and Eomyrochea Linnavuori, 1982 (Hemiptera: Heteroptera: Pentatomidae: Pentatominae: Myrocheini) are confirmed as objective synonyms, both sharing the same type species: Dymantis subvittata Stal, 1861. As a result, Dymantis (= Eomyrochea, syn. confirm.) is downgraded to a subgenus of Myrochea Am...

The first specific locality data within New Guinea is provided for Tamolia ramifera (Walker, 1868). The male genitalia of this species is described for the first time. Tamolia ancalagon n. sp. is described from Karimui, New Guinea.

In this contribution, we present new distributional records from Argentina of 63 pentatomid species; three genera (. Caonabo, Cromata and Taurocerus) and 14 species (. Marmessulus brasiliensis, Podisus crassimargo, Cromata ornata, Acledra haematopa, Caonabo pseudocylax, Dichelops avilapiresi, Euschistus cribarius, E. paranticus, Mormidea maculata,...

Resumen Se describe Planois smaug sp. nov. de Isla Navarino, Región de Magallanes, y se entrega una lista actualizada de los Acantosomátidos de la Región de Magallanes, indicando su distribución por provincias. Se añade una clave de identificación para las especies de la familia presentes en Magallanes. Palabras clave: Heteroptera, Acanthosomatidae...

The acanthosomatid genus Tolono Rolston and Kumar is revised. The generic description is modified slightly to accommodate the two new species described herein: Tolono confusus n. sp. is described from Ecuador and Colombia and T. pallidus n. sp. is described from Colombia. The first host plant record, Baccharis floribunda (Ruiz and Pav.) (Asteraceae...

Acledra (Acledra) bichromocornis n. sp. from the highlands of Argentina and Bolivia is described. A checklist of the species of Acledra s. str. is provided, including the distribution by country. A key to the species of Acledra s. str. is included.

Pentamyrmex spinosus new genus, new species is described, and a new tribe is erected for this remarkable insect. This is the first report of ant mimicry in the Pentatomidae.

New distributional data is provided for Ea australis Distant, 1911 in Chile and Argentina. The presence of this species in the Magallanes Region of southern Chile is confirmed. On the other hand the northernmost record for this species is provided, from Neuquén Province in Argentina. The first hostplant records (Nothofagus pumilio and Nothofagus sp...

Pentatomidae have been relatively poorly documented in Minnesota. Based on literature and museum records, an annotated checklist of the Pentatomidae of Minnesota was created. State-level and county-level records for Minnesota and the distribution of each species in North America are provided. Fifty-one species of Pentatomidae (12 Asopinae, 37 Penta...

The following new synonymies within the family Pentatomidae, subfamily Pentatominae, are established: Aesula Stål, 1876 = Pseudacrosternum Day, 1965, syn. nov. (Nezarini), Aesula viridissima (Signoret, 1861) = Aesula signoretiana Kirkaldy, 1909, syn. nov. = Pseudacrosternum cachani Day, 1965, syn. nov. = Pseudacrosternum cachani var. scutellatum Da...

Archaeoditomotarsus crassitylus, gen. and sp. nov. of Acanthosomatidae (Ditomotarsinae: Ditomotarsini), is described and illustrated. The biogeography of the new taxon is discussed and a key to the Chilean genera of Ditomotarsinae is included.

Thestral incognitus, new genus and species of Pentatomidae (Pentatominae: Carpocorini) is described from central Chile. The relationships among this new genus and other South American genera are discussed. Also, similarities are noted between Thestral and the Australian genus Poecilotoma. Poecilotoma is tentatively placed in the Carpocorini. A key...

Seven families of Pentatomoidea including Acanthosomatidae, Dinidoridae, Pentatomidae, Plataspididae, Scutelleridae, Thaumastellidae, and Thyreocoridae are discussed in this catalog. The Iranian fauna of Pentatomoidea (excl. Cydnidae) is represented by 99 genera and 271 species and subspecies. Elasmostethus interstinctus (Linnaeus, 1758) (Acanthoso...

Cochabambia Marcuzzi (1985) (Coleoptera: Tenebrionidae) is a junior homonym of Cochabambia Pirán, 1959 (Hemiptera: Heteroptera: Acanthosomatidae). Allotriocochabambia is proposed as a new name for the tenebrionid genus, and as result Allotriocochabambia kulzeri (1985) is proposed as new combination. The placement of Cochabambia Pirán within the Aca...

A new extinct family of Pentatomoidea, Primipentatomidae fam. nov., with four new genera and five new species (Primipentatoma peregrina gen. et sp. nov., Primipentatoma fangi gen. et sp. nov., Breviscutum lunatum gen. et sp. nov., Oropentatoma epichara gen. et sp. nov. and Quadrocoris radius gen. et sp. nov.), is described from the Upper Mesozoic d...

Resumen Se revisa el estado actual del conocimiento del género Chinavia Orian, 1965 en Chile, se aportan nuevos datos de distribución y plantas hospedadoras, se describe Chinavia perezi n. sp. y se incluye una clave para las especies de este género presentes en Chile. Synopsis of the genus Chinavia Orian, 1965 (Heteroptera: Pentatomidae) in Chile A...

The Pentatomoidea fauna for North Dakota is documented. There are 62 species of Pentatomoidea known from North Dakota: Acanthosomatidae (2), Cydnidae (4), Pentatomidae: Asopinae (9), Pentatomidae: Pentatominae (34), Pentatomidae: Podopinae (2), Scutelleridae (6), and Thyreocoridae (5). Of this total, 36 represent new state records for North Dakota....

An extinct new family of Pentatomomorpha, Venicoridae Yao, Ren & Cai fam. nov., with 2 new genera and 2 new species (Venicoris solaris Yao, Ren & Rider gen. & sp. nov. and Clavaticoris zhengi Yao, Ren & Cai gen. & sp. nov.) are described from the Early Cretaceous Yixian Formation in Northeast China. A cladistic analysis based on a combination of fo...

Geographical and stratigraphic distribution of the known fossil species in Pentatomomorpha from the Mesozoic. (T3 = Late Triassic, J1 = Early Jurassic, J2 = Middle Jurassic, J3 = Late Jurassic, K1 = Early Cretaceous; * indicates amber specimens). (DOC)