David R. Braun

• PhD, Rutgers University, Anthropology
• Professor (Full) at George Washington University

Background Climate change is increasing temperatures, frequency of heatwaves, and erratic rainfall, which threatens human biology and health, particularly in already extreme environments. Therefore, it is important to understand how environmental heat stress measures are tied to human water needs and thermoregulation under increasingly hot conditio...

Objective Extreme climatic events, like droughts, are increasing in frequency and severity. Droughts disrupt community livelihoods and resources with serious implications for human biology. This study investigated how chronic stress, measured by fingernail cortisol concentration (FCC), and water insecurity status were predictive of C‐reactive prote...

Objectives In subsistence populations, high physical activity is typically maintained throughout pregnancy. Market integration shifts activity patterns to resemble industrialized populations, with more time allocated to sedentary behavior. Daasanach semi‐nomadic pastoralists living in northern Kenya face lifestyle heterogeneity due to the emergence...

Background Pastoralists live in challenging environments, which may be accompanied by unique activity, energy, and water requirements. Aim Few studies have examined whether the demands of pastoralism contribute to differences in total energy expenditure (TEE) and water turnover (WT) compared to other lifestyles. Subjects and methods Accelerometer...

Background and objectives Non-communicable disease (NCD) risk and the epidemic of cardiometabolic diseases continue to grow across the expanding industrialized world. Probing the relationships between evolved human physiology and modern socioecological condition is central to understanding this health crisis. Therefore, we investigated the relation...

In the study of Early Pleistocene stone artifacts, researchers have made considerable progress in reconstructing the technical decisions of hominins by examining various aspects of lithic technology, such as reduction sequences, hammer selection, platform preparation, core management, and raw material selection. By comparison, our understanding of...

The Miocene was a key time in the evolution of African ecosystems witnessing the origin of the African apes and the isolation of eastern coastal forests through an expanding arid corridor. Until recently, however, Miocene sites from the southeastern regions of the continent were unknown. Here, we report the first Miocene fossil teeth from the shoul...

During the middle Pliocene (~3.8e3.2 Ma), both Australopithecus afarensis and Kenyanthropus platyops are known from the Turkana Basin, but between 3.60 and 3.44 Ma, most hominin fossils are found on the west side of Lake Turkana. Here, we describe a new hominin locality (ET03-166/168, Area 129) from the east side of the lake, in the Lokochot Member...

Intentionally produced sharp-edged stone flakes and flaked pieces are our primary evidence for the emergence of technology in our lineage. This evidence is used to decipher the earliest hominin behavior, cognition, and subsistence strategies. Here, we report on the largest lithic assemblage associated with a primate foraging behavior undertaken by...

The oldest Oldowan tool sites, from around 2.6 million years ago, have previously been confined to Ethiopia's Afar Triangle. We describe sites at Nyayanga, Kenya, dated to 3.032 to 2.581 million years ago and expand this distribution by over 1300 kilometers. Furthermore, we found two hippopotamid butchery sites associated with mosaic vegetation and...

Objectives: Investigations of early childhood growth among small-scale populations are essential for understanding human life history variation and enhancing the ability to serve such communities through global public health initiatives. This study characterizes early childhood growth trajectories and identifies differences in growth patterns rela...

The analyses of the stable isotope ratios of carbon (δ¹³C), nitrogen (δ¹⁵N), and oxygen (δ¹⁸O) in animal tissues are powerful tools for reconstructing the feeding behavior of individual animals and characterizing trophic interactions in food webs. Of these biomaterials, tooth enamel is the hardest, most mineralized vertebrate tissue and therefore l...

This article quantifies Daasanach water insecurity experiences in northern Kenya, examines how water insecurity is associated with water borrowing and psychosocial stress, and evaluates if water borrowing mitigates the stress from water insecurity. Of 133 households interviewed in seven communities, 94.0% were water insecure and 74.4% borrowed wate...

The ability of humans to mediate environmental variation through tool use is likely the key to our success. However, our current knowledge of early cultural evolution derives almost exclusively from studies of stone tools and fossil bones found in the archaeological record. Tools made of plants are intrinsically perishable, and as such are almost e...

Objective Water plays a critical role in the production of food and preparation of nutritious meals, yet few studies have examined the relationship between water and food insecurity. The primary objective of this study, therefore, was to examine how experiences of household water insecurity (HWI) relate to experiences of household food insecurity (...

Globally, fire is a primary agent for modifying environments through the long-term coupling of human and natural systems. In southern Africa, control of fire by humans has been documented since the late Middle Pleistocene, though it is unclear when or if anthropogenic burning led to fundamental shifts in the region's fire regimes. To identify poten...

Humans evolved in the dynamic landscapes of Africa under conditions of pronounced climatic, geological and environmental change during the past 7 million years. This book brings together detailed records of the paleontological and archaeological sites in Africa that provide the basic evidence for understanding the environments in which we evolved....

Background Gorongosa National Park in Mozambique hosts a large population of baboons, numbering over 200 troops. Gorongosa baboons have been tentatively identified as part of Papio ursinus on the basis of previous limited morphological analysis and a handful of mitochondrial DNA sequences. However, a recent morphological and morphometric analysis o...

The late Holocene was a period of cultural change along the west coast of South Africa, with widespread archaeological evidence for shifts in settlement patterns and economic activity. With these changes, we expect variability in the movement patterns of resident populations. In this proof-of-concept paper, we use lithic assemblages from Spring Cav...

This article quantifies Daasanach water insecurity experiences in Northern Kenya, examines how water insecurity is associated with water borrowing and psychosocial stress, and evaluates if water borrowing mitigates the stress from water insecurity. Of 133 households interviewed in 7 communities, 95% were water insecure and 74.4% borrowed water thre...

To understand the ways in which past stone knappers controlled the morphology of the flakes they produced, archaeologists have focused on examining the effects of striking platform attributes on flake size and shape. Among the variables commonly considered, platform width has routinely been noted to correlate with flake size and hence used to expla...

The Greater Cape Floristic Region (GCFR) of South Africa is a biodiversity hotspot of global significance, and its archeological record has substantially contributed to the understanding of modern human origins. For both reasons, the climate and vegetation history of southwestern South Africa is of interest to numerous fields. Currently known paleo...

Objectives: Thirst is an evolved central homeostatic feedback system that helps regulate body water for survival. Little research has examined how early development and exposure to extreme environments and water availability affect thirst perception, particularly outside Western settings. Therefore, we compared two indicators of perceived thirst (...

The Miocene is a key time in the evolution of African mammals and their ecosystems witnessing the origin of the African apes and the isolation of eastern coastal forests through an expanding biogeographic arid corridor. Until recently, however, Miocene sites from the southeastern regions of the continent were unknown. Here we report discovery of th...

The aspects of hominin behavior responsible for Oldowan stone tool variation are the focus of much debate. There is some consensus that this variation arises from a combination of ecological and cultural factors. The diversity of raw material types and technological strategies present at Kanjera South, Kenya, provide an opportunity to examine the i...

The flora of the Greater Cape Floristic Region (GCFR) of South Africa is a biodiversity hotspot of global significance, and its archaeological record has contributed substantially to the understanding of modern human origins. For both reasons, the climate and vegetation history of south-western South Africa is of interest to numerous fields. Curren...

The KNM-ER 2598 occipital is among the oldest fossils attributed to Homo erectus but questions have been raised about whether it may derive from a younger horizon. Here we report on efforts to relocate the KNM-ER 2598 locality and investigate its paleontological and geological context. Although located in a different East Turkana collection area (A...

Despite advances in our understanding of the geographic and temporal scope of the Paleolithic record, we know remarkably little about the evolutionary and ecological consequences of changes in human behavior. Recent inquiries suggest that human evolution reflects a long history of interconnections between the behavior of humans and their surroundin...

Spatial information is crucial to archaeological field research. From the plane-table to the total station, recent technological advances have enabled data collection to become fully digital and highly accurate. Nevertheless, the recent expansion of archaeological expeditions to novel environments often incompatible with modern mapping equipment, e...

Stone tools represent the largest source of information about past human behaviors on the planet. Much of the information about stone tools remains untranslated because we have little understanding about what the variation in artifact form means. One component of stone tool production that has less ambiguity is the reductive nature of the technolog...

One of the greatest difficulties with evolutionary approaches in the study of stone tools (lithics) has been finding a mechanism for tying culture and biology in a way that preserves human agency and operates at scales that are visible in the archaeological record. The concept of niche construction, whereby organisms actively construct their enviro...

Water salinity is a growing global environmental health concern. However, little is known about the relation between water salinity and chronic health outcomes in non-coastal, lean populations. Daasanach pastoralists living in northern Kenya, traditionally rely on milk, yet are experiencing socioecological changes and have expressed concerns about...

The stone artifact record has been one of the major grounds for investigating our evolution. With the predominant focus on their morphological attributes and technological aspects of manufacture, stone artifacts and their assemblages have been analyzed as explicit measures of past behaviors, adaptations, and population histories. This analytical fo...

The original version of this article unfortunately contained mistake in the presentation of the author’s name.

One of the greatest difficulties with evolutionary approaches in the study of stone tools (lithics) has been finding a mechanism for tying culture and biology in a way that preserves human agency and operates at scales that are visible in the archaeological record. The concept of niche construction, whereby organisms actively construct their enviro...

The Pleistocene ungulate communities from the western coastal plains of South Africa's Cape Floristic Region (CFR) are diverse and dominated by grazers, in contrast to the region's Holocene and historical faunas, which are relatively species-poor and dominated by small-bodied browsers and mixed feeders. An expansion of grassy habitats is clearly im...

Determining the provenance of archeological material is essential for documenting the movement of ancient resources and placing fossils and artifacts in the correct stratigraphic context. Non-destructive geochemical compositional analysis with Energy Dispersive X-ray Fluorescence (ED-XRF) spectrometry has the potential to provide a versatile approa...

The three-dimensional (3D) revolution promised to transform archaeological practice. Of the technologies that contribute to the proliferation of 3D data, photogrammetry facilitates the rapid and inexpensive digitization of complex subjects in both field and lab settings. It finds additional use as a tool for public outreach, where it engages audien...

Objectives This study compared the prevalence of concentrated urine (urine specific gravity ≥1.021), an indicator of hypohydration, across Tsimane' hunter‐forager‐horticulturalists living in hot‐humid lowland Bolivia and Daasanach agropastoralists living in hot‐arid Northern Kenya. It tested the hypotheses that household water and food insecurity w...

Paranthropus boisei was first described in 1959 based on fossils from the Olduvai Gorge and now includes many fossils from Ethiopia to Malawi. Knowledge about its postcranial anatomy has remained elusive because, until recently, no postcranial remains could be reliably attributed to this taxon. Here, we report the first associated hand and upper li...

During the last decade new geospatial approaches aimed at increasing the likelihood of finding fossiliferous deposits have emerged [1]. Using satellite imagery and coordinates of previously known fossil localities, machine learning algorithms can be trained to mine spectral patterns associated with fossiliferous outcrops across a region [2]. Two ma...

This paper investigates Oldowan hominin behavioral ecology through use-wear analysis of artifacts from Kanjera South, Western Kenya. It extends development of our experimental use-wear reference collection and analysis of use-wear on the well preserved and unweathered Oldowan tools from this site to include rhyolite, a non-local material of similar...

Hominin fire use in the early Pleistocene has been debated since the early 1970s when consolidated reddened sediment patches were identified at FxJj20 East and Main, Koobi Fora, Kenya. Since then, researchers have argued for evidence of early Pleistocene fire use at a handful of archaeological sites with evidence of combustion. Some argue that morp...

Levallois technology is a hallmark of many Middle and Late Pleistocene stone artifact assemblages, but its definition has been much debated. Here we use three-dimensional photogrammetry to investigate the geometric variation among Levallois and discoidal core technologies. We created models of experimental and archaeological stone artifact assembla...

It has been suggested that a shift in diet is one of the key adaptations that distinguishes the genus Homo from earlier hominins, but recent stable isotopic analyses of fossils attributed to Homo in the Turkana Basin show an increase in the consumption of C4 resources circa 1.65 million years ago, significantly after the earliest evidence for Homo...

Significance Humans are distinguished from all other primates by their reliance on tool use. When this uniquely human feature began is debated. Evidence of tool use in human ancestors now extends almost 3.3 Ma and becomes prevalent only after 2.6 Ma with the Oldowan. Here, we report a new Oldowan locality (BD 1) that dates prior to 2.6 Ma. These ea...

Despite the numerous hominin fossils found in the Omo-Turkana Basin dating to between 2.0 and 1.4 Ma., a resolved understanding of their dietary ecology has been challenging due to limited research on similar patterns in contemporaneous large mammals. In this study, we use a sample (n = 390) of enamel δ13C values of six Bovidae, Suidae, and Equidae...

Any inference of behavior based upon the spatial distribution of archaeological material requires a detailed understanding of site formation processes. Natural agents, such as water flow, are responsible for post-depositional alteration of buried materials. These processes can result in spatial patterns that mask the behavioral processes associated...

The Koobi Fora Formation of the northeastern Turkana Basin provides a largely continuous fossil- and artifact-rich volcanoclastic sequence allowing for the investigation of morphological and behavioral evolution during the Plio-Pleistocene. Recent and ongoing archaeological investigations have documented in situ evidence of early modern humans in t...

Raw craniofacial landmark data comprising papio specimens from Gorongosa National Park and the comparative dataset from Dunn et al. 2013. 3D Mesh at: http://www.morphosource.org/Detail/MediaDetail/Show/media_id/40022

Most authors recognize six baboon species: hamadryas (Papio hamadryas), Guinea (Papio papio), olive (Papio anubis), yellow (Papio cynocephalus), chacma (Papio ursinus), and Kinda (Papio kindae). However, there is still debate regarding the taxonomic status, phylogenetic relationships, and the amount of gene flow occurring between species. Here, we...

Fluctuations of paleolake Lorenyang and volcanic activity in the Turkana Basin from 2.0-1.38 Ma caused environmental shifts during a crucial period of hominin evolution. To examine spatial variability in habitats within the Turkana basin, we studied faunal abundance data compiled from the Koobi Fora (KF), Nachukui, and Shungura Formations of the Om...

In the version of this Article originally published, the authors mistakenly included duplicate entries in the flake datasets for the new Pech de l’Azé IV and Warwasi collections, resulting in minor errors in the statistical analysis. The authors have now repeated this analysis with the correct flake datasets. As a result, in the following two sente...

KNM-ER 47000 is a fossil hominin upper limb skeleton from the Koobi Fora Formation, Kenya (FwJj14E, Area 1A) that includes portions of the scapula, humerus, ulna, and hand. Dated to ∼1.52 Ma, the skeleton could potentially belong to one of multiple hominin species that have been documented in the Turkana Basin during this time, including Homo habil...

KNM-ER 47000A is a new 1.52 Ma hominin scapular fossil belonging to an associated partial skeleton from the Koobi Fora Formation, Kenya (FwJj14E, Area 1A). This fossil effectively doubles the record of Early Pleistocene scapulae from East Africa, with KNM-WT 15000 (early African Homo erectus) preserving the only other known scapula to date. KNM-ER...

The Plio-Pleistocene fossil record of eastern Africa is generally characterized by increasing aridity and expansion of C4 grasslands. Between ~2–1.4 Ma, paleolake Lorenyang in the Turkana Basin regressed in response to climate cycling, potentially restructuring the surrounding ecosystems. Using faunal abundance of mammalian communities from the Koo...

From its initial appearance at ~1.7 Ma, the Acheulean was prevalent through a vast chronological span of hominin behavioural evolution that lasted nearly 1.5 million years. The origins and production patterns of large bifacial cutting tools ('LCTs') e the marker of the Acheulean techno-complex e and the systematic changes in this behaviour through...

From its initial appearance at ~1.7 Ma, the Acheulean was prevalent through a vast chronological span of hominin behavioural evolution that lasted nearly 1.5 million years. The origins and production patterns of large bifacial cutting tools ('LCTs') e the marker of the Acheulean techno-complex e and the systematic changes in this behaviour through...

First fossil sites from the Urema Rift, central Mozambique, and their paleoenvironmental and paleoecological contexts

Oldowan sites in primary geological context are rare in the archaeological record. Here we describe the depositional environment of Oldowan occurrences at Kanjera South, Kenya, based on field descriptions and granulometric analysis. Excavations have recovered a large Oldowan artefact sample as well as the oldest substantial sample of archaeological...

The East African Rift System (EARS) has played a central role in our understanding of human origins and vertebrate evolution in the late Cenozoic of Africa. However, the distribution of fossil sites along the rift is highly biased towards its northern extent, and the types of paleoenvironments are primarily restricted to fluvial and lacustrine sett...

The study of non-human primate tool use, particularly that of chimpanzees (Pan troglodytes sp.) has provided a comparative framework for the study of human evolution and the contexts surrounding the emergence and development of technology in our lineage. As our closest living relatives, chimpanzees, like early hominins, use stone tools for percussi...

The ecological and selective forces that sparked the emergence of Homo's adaptive strategy remain poorly understood. New fossil and archaeological finds call into question previous interpretations of the grade shift that drove our ancestors' evolutionary split from the australopiths. Furthermore, issues of taphonomy and scale have limited reconstru...

Today, the family Hippopotamidae is represented by two genera, Hippopotamus and Choeropsis. Modern representatives of these genera differ significantly from one another in terms of size, ecology, and geographical distribution. While differences in size span back as far as the Miocene, throughout their fossil record in Africa, it is unclear whether...

Fossil faunal abundance data from Plio-Pleistocene deposits are useful for understanding the spatial and temporal dynamics of the environments inhabited by hominins. Here, we synthesize faunal abundance records from the Koobi Fora, Nachukui, and Shungura formations of the Turkana Basin in northern Kenya and southern Ethiopia to better understand en...

Temporal variability in flaking stone has been used as one of the currencies for hominin behavioural and biological evolution. This variability is usually traced through changes in artefact forms and techniques of production, resulting overall in unilineal and normative models of hominin adaptation. Here, we focus on the fundamental purpose of flak...

Artifacts with varying use-lives have different discard rates and hence are represented unequally among archaeological assemblages. As such, the ability to gauge the use-lives of artifacts is important for understanding the formation of archaeological assemblage variability. In lithic artifacts, use-life can be expressed as the extraction of utilit...

A major puzzle in human origins research is the question of where, when, and under what environmental conditions our lineage originated in Africa, but answers are hampered by the scarcity of Mio-Pliocene paleontological sites. To help fill these gaps, the Paleo-Primate Project Gorongosa, a multidisciplinary research initiative on human origins, was...

The well-known South African mid-Pleistocene site of Elandsfontein has yielded an abundance of fossil fauna and artifacts (ca. 1 to 0.6 million years ago) and has produced hominin fossils, specifically a calvarium and mandibular fragment often assigned to the species Homo heidelbergensis. Elandsfontein is located within the Greater Cape Floristic R...

While lithic objects can potentially inform us about past adaptations and behaviors, it is important to develop a comprehensive understanding of all of the various processes that influence what we recover from the archaeological record. We argue here that many assumptions used by archaeologists to derive behavioral inferences through the definition...

We have learned much about tool use in nonhumans since the discovery of Oldowan stone tools. Despite the ongoing debate over whether tool use in other animals requires cultural transmission, it seems clear that, today, humans show a quantitative, if not qualitative, difference in our ability to transmit information socially through cultural transmi...