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Introduction
I am a systematist interested in assessing global biodiversity of overlooked small eucaryotes, particularly marine snails and orchids. I enjoy various light and scanning electron microscopy techniques, and writing books.
Current institution
Additional affiliations
February 2004 - present

Position
- Curator of Malacology
Description
- Research into systematics of marine snails (Gastropoda) and orchids (genus Oberonia). Oversee collection of 2-3M Invertebrates from Porifera through Chordata. In charge of Scanning Electron Microscopy facility (Zeiss EVO 10LS, LaB6, SE, HDBSD, C2DX, beam deceleration, cold stage, VP-E-SEM, EDS, N2), CPD, sputter coater. Work and oversee volunteers and students/post-docs. Former department head. Retired as of Aug 1 2024.
January 2016 - October 2023
Editor roles

Zootaxa Senior subject editor Mollusca 2003-2013
Position
- Associate Editor

Malacologia editorial board 2008-present
Position
- Associate Editor

Molluscan Research associate editor 2004-present
Position
- Associate Editor
Education
January 1994 - December 1999
September 1987 - December 1993
Publications
Publications (129)
We report on curation and digitization experiences of a large collection of Bryozoa (>19,759 of estimated 38,000 lots digitized, >1,000 type lots). Storage of one to two histology slides in capped glass vials and larger series in twenty-five-slot histology boxes is optimal. Bulk dry material is stored in acid-free containers (cardboard, plastic). F...
Three new records of Bathysciadium pacificum Dall, 1908 from off Oregon and Washington extend the range of the species from Peru by ~7,300 km to the temperate northeastern Pacific. The preserved and relaxed animal shows that the mantle extensions are not sensory papillae, but mantle folds forming the periostracum rays on the shell. Examination of r...
The genus Hippeophyllum Schltr. is synonymised under Oberonia Lindl. The following names are synonymised under Oberonia scortechinii Hook.f.: Hippeophyllum alboviride J.J.Sm., H. biakense J.J.Sm., H. celebicum Schltr., H. halmaherense J.J.Sm., O. hamadryas Ridl., O. longifolia Ridl., H. micranthum Schtr., H. papillosum Schltr., H. sulense J.J.Sm. a...
The rivers of southeastern Australia host a species complex within the carp gudgeon genus Hypseleotris that includes parental species and hemiclonal hybrid lineages. These hemiclones can be difficult to distinguish from their parent taxa, making delineation of species unusually difficult. We approach this historical taxonomic problem by using singl...
A checklist of all species of bryophytes, lycophytes, ferns, gymnosperms and angiosperms that are found in the wild (native, naturalised and casual) in Singapore is presented. We have attempted to account for all names of species and infraspecific taxa that have ever been recorded for Singapore, along with the pertinent publications that reported e...
A few taxonomic and nomenclatural aspects of the genus Oberonia Lindl. relating to the Flora of Singapore are resolved. First, no subgeneric names are recognised in the genus Oberonia. Oberonia subg./sect. Apothemnophyllum Schltr. and Oberonia subg./sect. Menophyllum are synonyms of Oberonia s.s. The autonymous subgenus is a non-mandatory rank, hen...
Cultivation and flowering of Oberonia maxima. Images of flowers by light microscopy, scanning electron microscopy (SEM), and ultraviolet reflectance imaging (simulated insect vision). SEM of seeds.
A checklist of all species of bryophytes, lycophytes, ferns, gymnosperms and angiosperms that are found in the wild (native, naturalised and casual) in Singapore is presented. We have attempted to account for all names of species and infraspecific taxa that have ever been recorded for Singapore, along with the pertinent publications that reported e...
The new species of Lichtenstein’s catalog (1794: part 2) were discussed by Geiger (2003). Here a facsimile edition and full translation of the work is given. Additional insights into select Lichtenstein species are provided. Tentative identifications include Mya papyracea Lichtenstein, 1794 as a member of Thraciidae, Helix porphyrostoma Lichtenstei...
We describe two new species in the genus Gobiomorphus, a radiation of fresh and brackish water gudgeons known from Australia and New Zealand. These species are a prominent component of New Zealand's freshwater ichthyofauna and most are widely distributed throughout both the North and South Islands. Two of the inland species, G. breviceps and G. bas...
Preface for special issue on Orchid Systematics in Rheedea 31(3)
The following synonyms are introduced or confirmed for Oberonia brachystachys Lindl.: O. recurva Lindl., O. setifera Lindl., O. demissa Lindl., O. gardneriana Thwaites, O. parvula King & Pantl. [including O. myriantha var. parvula (King & Pantl.) Tang & Wang], O. croftiana King & Pantl. (second-step lectotypification herein), O. lingmalensis Blatt....
A new species of Scissurella (Gastropoda, Scissurellidae) from Cuba. Una nueva especie de Scissurella (Gastropoda, Scissurellidae) de Cuba. Iberus 39: 225-229.
The reasons for excess names in microfloral orchids such as Oberonia Lindl. can be traced to poor scholarship (e.g., failure to review the literature, ignoring expert advice), typological thinking, and erroneous assumption of microendemism. Some extraordinarily poor descriptions, including some from the 21st century, can be termed “taxonomic vandal...
The following 24 new synonyms in the orchid genus Oberonia are proposed, correct name (synonym(s)): O. aporophylla (O. longirepens); O. bertoldii (O. gracillima, O. laeta); O. bifida (O. celebica, O. fissiglossa); O. brunoniana (O. wallichii, O. saintberchmansii); O. costeriana (O. vulcanica); O. ensiformis (O. vesiculifera); O. griffithiana (O. kh...
A combined morphological-molecular investigation identifies ten new synonyms of Oberonia equitans (G.Fost.)Mutel: Oberonia affinis Ames & C.Schweinf. in O.Ames, Oberonia aurea Schltr., Oberonia ciliolata Hook.f., Oberonia enoensis J.J.Sm., Oberonia equitans var. chaperi Finet, Oberonia lampongensis J.J.Sm., Oberonia mcgregorii Ames, Oberonia murkel...
Page proof corrections were incompletely carried out by Magnolia Press / Zootaxa. The LSID number urn:lsid:zoobank.org:pub:8D3B9B4C-5EA7-4746-9987-CBE75B771D0E should have been used instead of the place holder. This is an error beyond the control of the authors.
Two new fissurellid limpets (Mollusca: Gastropoda: Fissurellidae), Fissurella? stantoni n. sp. and Sceli- dotoma aldersoni n. sp., are described from Miocene deposits in southern California. Fissurella? stantoni is described from a single specimen from the middle Miocene Topanga Canyon Formation in the Santa Monica Mountains, Los Angeles County, Ca...
Two new fissurellid limpets (Mollusca: Gastropoda: Fissurellidae), Fissurella? stantoni n. sp. and Sceli- dotoma aldersoni n. sp., are described from Miocene deposits in southern California. Fissurella? stantoni is described from a single specimen from the middle Miocene Topanga Canyon Formation in the Santa Monica Mountains, Los Angeles County, Ca...
Characters for the identification of the orchid genus Oberonia are discussed, including the rather extensive intraspecific variability.
Conference presentation on micro mollusk diversity. Paper published in The Nautilus 2018.
Poster on the McLean Memorial volume published in Zoosymposia 13, in full open access format
Description of the recently obtained deep-sea and hydrothermal vent molluscan collection from Robert Vrijenhoek of the Monterey Bay Research Institute (MBARI), now housed at the Santa Barbara Museum of Natural History (SBMNH)
The following 24 new synonyms in the orchid genus Oberonia are proposed, correct name (+ synonym(s)): O. bicornis (+ O. tenuis); O. brunoniana [lectotype here designated] (+ O. santapaui lectotype and epitype here desig- nated); O. caulescens (+ O. katakiana); O. complanata (+ O. flavescens); O. costeriana (+ O. elmeri, + O. obesa, + O. gigantea);...
Ultraviolet reflectance photography of Oberonia species only shows limited patterning on flowers and rachis. It suggests that pollinators have relatively poor vision, which makes Diptera a likely group. Spatial scales of the patterns suggest that visual cues only play a role at distances of 2–50 cm. Attraction to flowers over larger distances could...
The abalone of the northeastern Pacific are reviewed. The taxonomy and distribution remains unchanged. McLean’s work on abalone is discussed.
The microscopic slit bearing gastropods of the northeastern Pacific are reviewed. Since the global monograph of Geiger (2012), one additional species has been described (Anatoma georgii Geiger, 2017). No significant material of other species has changed distributional records or range limits of the known taxa.
The family Seguenziidae, including the genus Adeuomphalus, of the northeast Pacific are here revised. Seguenzia megaloconcha Rokop, 1972 is here synonymized under S. cervola Dall, 1919, S. quinni McLean, 1985 is here synonymized under S. gioviae Dall, 1919, S. catalina Dall, 1919 and S. certoma Dall, 1919 are synonymized under S. stephanica Dall, 1...
A biography of the late James H. McLean, former Curator of Malacology at the Natural History Museum of Los Angeles County is provided. It is complemented with a full bibliography and list of 344 taxa named by him and co-authors (with type information and current status), as well as 40 patronyms.
James H. McLean Memorial Volume (Table of Content)
James H. McLean Memorial Volume (Cover)
Foreword The present volume has a history spanning approximately 60 years. It began with James (Jim) Hamilton McLean's dissertation, commenced in the early 1960's and completed in 1966. Right from the beginning, Jim wanted to revise the entire gastropod fauna of the northeast Pacific. This initial dissertation proposal was tempered in scope by the...
Four statistical metrics (Chao1, Chao2, ACE, and rarefaction) were used to estimate the total number of species of shelled micromollusks based on composition of nine benthic samples collected from the sublittoral zone off Wailea, Maui, Hawaiʻi. There was a total of 250 species in the nine samples analyzed, and the estimated total number of species...
The Convention on Biological Diversity (CBD) commits its 196 nation parties to conserve biological diversity, use its components sustainably, and share fairly and equitably the benefits from the utilization of genetic resources. The last of these objectives was further codified in the Convention's Nagoya Protocol (NP), which came into effect in 201...
Violin rosins are marketed with metal and other additives. Scanning Electron Microscopy - Energy Dispersive Analysis of X-rays SEM-EDAX/EDS was used to assess the composition of a variety of rosins. None contained any of the alleged additives at the detection threshold of 0.01%.
Orchid flowers are amongst the most conspicuous attractions that plants have generated over evolutionary epochs. However, orchid flowers have been and are still under a long-term discussion about organ homology in particular of androecium and gynoecium organs. A contribution to the clarification of organ identity orchid floral aberrations has tradi...
Four new species of small to minute Vetigastropoda from two
families are described: in the Anatomidae, Anatoma georgii new species from the intertidal of Alaska, with its radula illustrated; in the Seguenziidae, Carenzia golikovi new species, Asthelys careyi new species, and Seguenzia macleani new species, all from abyssal depth of the northeastern...
The present article describes Fissurella hendrickxi, a new deepsea
gastropod species found off the Pacific coast of Mexico (Baja
Peninsula; Gulf of California), at 650–837 m depth, collected by
the TALUD XV and TALUD X projects. Fissurella hendrickxi was
classified according to shell sculpture, radula, epipodium, and
ctenidial structure. It is rema...
Oberonia aureolabris sp. nov. from western Java is described, having been studied using light and scanning electron microscopy. It is distinguished by a panduriform inflated disc, relatively short and widely separated epichile lobes on the lip, more or less equal length of the floral bracts over the entire floriferous portion of the inflorescence a...
The malacofauna of Hawaii is rather well-known, owing to Kay (1979) and Severns (2011). Both works stand out because they include a large number of the generally under-represented micromollusks (<5 mm). Here a striking new genus and species of a microsnail is reported from that region.
Oberonia maxima is here recognized as the senior synonym and correct name of Oberonia integerrima and Oberonia orbicularis. Various misidentifications in the literature are corrected. Light and scanning electron micrographs of the flower are provided. Oberonia “maxima” found in the horticultural trade is identified as Oberonia ensiformis.
A taxonomic revision of the family Anatomidae from Brazil led to the discovery of nine species, four of which are new. This contrasts with only two species previously recorded: Anatoma aedonia (Watson, 1886) and Anatoma proxima (Dall, 1927). The lectotype designation for A. aedonia by Geiger (2012) has clarified the identity of this taxon, which is...
New information on the scissurellids fauna, Scissurellidae and Anatomidae (Gastropoda Vetigastropoda), in the Mediterranean Sea is presented. Scissurella azorensis Nolt, 2008, is confirmed from several localities in the Tyrrhenian Sea. Sinezona semicostata Burnay et Rolán, 1990, a species until now known from Cape Verde and Canary Islands, is repor...
With over 2 million mollusk specimens and 2,000 primary type lots, the Santa Barbara Museum of Natural History (SBMNH) houses one of the largest collections of mollusks in North America, with collections dating back to the mid-1800’s. The collection is housed in a modern well-lit facility, with expansive space for visiting researchers.
The SBMNH P...
Qualitative and quantitative data are given for wood anatomy of six of the 11 recognized species of Forchhammeria (Mexico, Central America, West Indies), a genus formerly placed in Capparaceae. Though still in Brassicales, the genus has been excluded, along with several other genera, from the major recognized families of that order on the basis of...
The new bathyal gastropod species, Cornisepta guzmani new species, collected at a depth of 846 metres from a methane seepage site in the subduction zone off Concepcio´n (36�220 S; 73�430 W), central Chile, is described. The new species is most similar to Cornisepta pacifica (Cowan, 1969), described from offshore Alaska and Oregon, from which it dif...
Thirteen new species of Scissurellidae (Scissurella regalis n. sp., Sinezona mechanica n. sp., Sinezona platyspira n. sp., Sinezona enigmatica n. sp., Sinezona wanganellica n. sp., Satondella azonata n. sp., Satondella bicristata n. sp.), Anatomidae (Anatoma amydra n. sp., Anatoma kopua n. sp., Anatoma megascutula n. sp., Anatoma tangaroa n. sp.),...
monograph of abalone.
This book is available from booksellers including Amazon, ConchBooks and the Santa Barbara Museum of Natural History. There is no digital copy.
Monograph of Scissurellidae Anatomidae Larocheidae Depressizonidae Sutilizonidae
**** Please do NOT ask for pdf/full-text. I will not make it available. No exception. ****
The book is in print. It can be purchased form Santa Barbara Museum of Natural History, ConchBooks, mal de mer books, NHBS, and amazon.
The species of Anatoma Woodward, 1859 from Norwegian waters and from the Norwegian Sea are treated taxonomically. Anatoma crispata (Fleming, 1828) has until quite recently been regarded as the sole representative of this family in north European waters, but constant conchological differences make it evident that at least three species have been lum...
This revision of the scissurellid genus Satondella Bandel, 1998 is mainly based on shell characters due to the availability of only a few live collected specimens. There are seven Recent species (two described as new) and one Eocene fossil. Satondella minuta Bandel, 1998, the type species from Indonesia, is redescribed and its range extended to New...
Geiger, Daniel L., Fernández-Garcés, Raúl (2010): Anatoma rolani n. sp. and new records of rare Caribbean Anatoma species (Gastropoda: Vetigastropoda: Anatomidae). Zootaxa 2488: 65-68, DOI: 10.5281/zenodo.195568
Anatomidae McLean, 1989, is a family of small (1–11 mm), slit-bearing Vetigastropoda (see Geiger et al. 2008) occurring from the shallow subtidal to the abyss in all the world’s oceans. They are distinguished from Scissurellidae by the peripheral location of the slit. Three genera have been described: Anatoma Woodward, 1859, Thieleella Bandel, 1998...
Anatoma fujikurai sp. nov. is described from the hydrothermal-vent environment at Myojin Knoll, southern Japan. The shell of the species is characterized by the predominant axial sculpture on the shoulder and base and by the undulating selenizone. The animal lacks eyes and shows a radular structure not seen in any other . anatomid species examined...
New Scissurellidae and Anatomidae from the geographic and political Americas are described, with many long-standing misidentifications being corrected, and some comparisons to described species complementing the contribution. Sinezona kayae n. sp., previously mis-identified as the South African Sin. insignis, and Sin. hawaiiensis n. sp. without a s...
The microscopic scissurellids are being currently revised; scissurellids is used here for the small Vetigastropoda formerly classified in Scissurelloidea, but which has been shown to be a polyphyletic assemblage (e.g., Geiger 2008, Geiger & Thacker 2005). Geiger (2003) presented an overview, where the new monotypic subfamily, genus, and species wer...
Three new Scissurellidae and one new Anatoma (Anatomidae) are described from Cape Manazuru, Sagami Bay, and Okinawa, Japan. Sinezona costulata n. sp. from Sagami Bay has a protoconch with strong axial cords, the apertural varix is not connected to the embryonic cap, and the teleoconch sculpture is initially of strong axials, which are replaced on t...
A systematic revision on the shallow and abyssal species of Scissurellidae and Anatomidae occurring in Sub-Antarctic and Antarctic waters was carried out. This study is based on material from several field trips and museum collections. The shell morphologies of all the species are redescribed and illustrated with scanning electron micrographs. Addi...
Four new species of Scissurellidae, Anatomidae, and Larocheidae are described from the Austral Archipelago, French Polynesia and the Indo-Malayan Archipelago: Sinezona danieldreieri new species, S. wiley new species (both Scissurellidae), Anatoma rapaensis new species (Anatomidae) and Trogloconcha lozoueti new species (Larocheidae). One of the spec...
Scissurella dalli Bartsch, 1903, from the southwestern Atlantic Ocean has been reported at least six times in the malacological literature. However, the species is thus far known only from the original description; it was never fi gured; and the type material is missing from the original repository, the United States National Museum, Smithsonian In...
This chapter investigates the evolution and phylogeny of the gastropod clade of Vetigastropoda. It explains that Vetigastropoda, introduced in 1980, is a clade of basal marine snails that include abalone, keyhole limpets, and slit shells. It provides a summary of the major groups within Vetigastropoda including their key characters and morphologica...
Four new species of Anatomidae are described: Anatoma herberti n. sp. with strong axial sculpture on the shoulder and base; A. austrolissa n. sp. with almost smooth sculpture except for axial cords in the adumbilical half of the base; A. boucheti n. sp. with sunken protoconch and selenizone that starts after more than one teleoconch I whorl; and A....
The present volume contains the proceedings from the symposium Micromolluscs: Methodological Challenges – Exciting Results held during the 16th UNITAS Malacologica World Congress of Malacology in Antwerp, Belgium, July 15–20, 2007. For the purpose of this volume, micromolluscs were defi ned as molluscs no larger than 5 mm. Such forms play an import...
Proceedings from the micromollusc symposium of the 16th UNITAS Malacologica World Congress of Malacology July 15–20, 2007 in Antwerp, Belgium
The use of molecular techniques is pervasive in contemporary systematics. We stress the importance of depositing voucher material, particularly for the less well-known micromolluscs, along with images from which specimens can be identifi ed. Appropriate storage is paramount to ensure long-term survival of those voucher specimens. We detail methods...
The last taxonomic monographs of micromolluscs (size less than 5 mm) of worldwide scope date back to the turn of the 19th/20th century, prior to the invention of the scanning electron microscope (SEM). In the present study, experiences from the preparation of a monograph on worldwide Scissurellidae s.l., a gastropod taxon comprising minute members...
Edited volume, conference proceedings from UNITAS symposium on micro mollusks.
Download pdfs from
http://www.vetigastropoda.com/micromolluscs
Micromolluscs are small-sized molluscs (< 5 mm), and include the great majority of undescribed molluscan taxa. Such species require special collecting, sorting and handling techniques and different storage requirements to those routinely used for larger specimens. Similarly, the preparation of shells, opercula, radulae and animals poses some challe...
Anatoma janetae n. sp. from the bathyal of the northeastern Pacific is described based on two incompletely preserved specimens, one including the animal. The new species is characterised by the marked switch in teleoconch sculpture from distinct raised axial cords to fine spiral steps. The species is compared to all north Pacific species. The eyes...
Sasakiconcha elegantissima, a new genus and species from the Western Pacific is described. The specimens are most similar to members of Anatomidae (Anatoma, Thieleella) with a peripheral selenizone, but close the slit to a foramen, and lack a columella because of a disjointly coiled base resulting in an umbilical trough. Additionally, an aberrant s...
Eight new species of Scissurellidae and Anatomidae are described: Scissurella kaiserae new species from the Panamic; Scissurella lorenzi new species from the Indo-Malayan archipelago; Scissurella maraisorum new species from South Africa; Sinezona garciai new species from the Caribbean; Sinezona globosa new species from the tropical Western Pacific;...
Triassurella carnica n. gen. n. sp., a new scissurelloid limpet-shaped gastropod is described from the Late Triassic (early Carnian) Cassian Formation. It represents one of the few Triassic members of the slit-bearing non-nacreous vetigastropods of the Scissurelloidea-Fissurelloidea-group and suggests that diversity and disparity of this,group are...
A molecular phylogeny of 40 basal gastropod species representing 14 families (including eight species in four families from chemosynthetic habitats) was constructed, based on 3038 characters (Histone 3, COI, 18S). Unambiguous fossils indicated minimum ages of four major lineages. It suggests that chemosynthetic habitats were colonized several times...
Phylogenetic relationships of 12 species of the vetigastropod family Haliotidae were analysed on the basis of partial haemocyanin coding genes. Both protein-coding and genomic sequence data, of which the latter includes one ancient intron, were analysed. We were able to distinguish two genetically different monophyletic groups with high bootstrap s...
A molecular phylogeny using cytochrome c oxidase subunit 1, Histone H3, and 18S rRNA, comprising 3038 aligned charac-ters, was developed for 38 ingroup vetigastropods and two neritimorph outgroups. Vetigastropoda is diagnosed by the presence of bursicles and their presence and absence in putative vetigastropod groups is discussed. The molecular dat...
Three new species of Scissurellidae from Australia are described: Incisura auriformis n. sp., Scissurella quadrata n. sp., and Sci. spinosa n. sp. They are compared to other species occurring in Australian waters: I. remota (Iredale, 1924) [+ I. vincentiana ( Cotton, 1959)], I. rosea (Hedley, 1904), Sci. cyprina Cotton & Godfrey, 1938, Sci. declina...
The Australian members of the vetigastropod family Anatomidae are revised and two new species are described. The family has thus far been treated as a subfamily of Scissurellidae, but recent molecular evidence ( Geiger & Thacker, unpubl. data) indicates that Scissurellinae plus Anatominae is not monophyletic, and full family rank is warranted for a...
Twenty-five nominal generic names have been proposed for the approximately 150 Recent species described in the family Scissurellidae (Mollusca : Gastropoda : Vetigastropoda). The generic diagnosing characters have not been uniformly applied and the value of these characters for scissurellid classification remains to be demonstrated. Here, the chara...
Anton August Heinrich Lichtenstein (1753–1816) was a member of a dynasty of natural scientists from Hamburg. He wrote an overlooked sales catalog of mollusks (1794) in which two new bivalve and 13 new gastropod taxa were described. The species’ identities are evaluated based on the original text written in German and Latin. Only two of his new spec...
Two types of anchoring rootlets on stolons of Bowerbankia imbricata (Adams, 1798) (Bryozoa: Ctenostomata) are discussed. Lateral anchoring rootlets form near the growing tip of the stolon carrying zooids, but become lateral structures as the stolon grows. These anchoring rootlets may be resorbed or additional ones may be formed. Terminal anchoring...
Questions
Questions (3)
In botanical nomenclature, is there a term for recognizing that a previously considered correct name is in fact a nom. illeg./superfl./inq./dubium ?
Stat. nov. only applies to changes in rank according to ICN, although the term would describe the change in status from correct to in-correct name very well. Such changes are nomenclatural novelties; it would be good to flag them.
Thanks for your wisdom!
Re typification of a plant genus if only a single species is/was contained in that genus (ICN Art. 40.3): Is that mode of designation "by original designation" or "by monotypy"? I am not sure whether the concept of typification by monotypy exists in botany (glossary has zero hits on "monotypy").
Even if typification by monotypy is not explicitly stated in ICN/Thurland, is it used in systematic botany? Or is this another distinction between ICN-Botany and ICZN-Zoology?
Please note, that the question is not the description of a genus with a single species (monotypic genus), but the mode of typification.
Thanks!
I have a bunch of Scyphozoa and other jellyfish that are have been in 5-10% formalin for at least a decade. Is it safe to transfer those to 70% ethanol? Any downsides?
I do know that jellies need to be strongly fixed in formalin (or some other aldehyde), but once they are fixed, can they be transferred to the less toxic ethanol?