
Dale R. CalderRoyal Ontario Museum · Natural History
Dale R. Calder
Ph.D.
About
152
Publications
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Introduction
I'm currently Curator Emeritus at the Royal Ontario Museum, Toronto, Canada, having taken early retirement in 2003. While retired, I continue to do research on alpha-taxonomy of hydrozoans. Projects currently underway include taxonomic studies on hydroids of the Cocos Islands, Costa Rica, with colleagues from the USA, Ecuador, and Costa Rica; and an account on intertidal hydroids of Newfoundland and southern Labrador, Canada, as sole investigator. However, my health is now in decline.
Additional affiliations
November 2000 - June 2006

International Commission on Zoological Nomenclature
Position
- Commissioner
December 1983 - December 2002
September 1981 - December 2003
Education
August 1964 - August 1968
September 1960 - April 1964
Publications
Publications (152)
Fourteen species of hydroids, collected during August 2019 by ROV SuBastian of the Schmidt Ocean Institute, are reported from the Emperor Seamount chain in the western North Pacific Ocean. Two others, Candelabrum sp. and Eudendrium sp., were observed only on videos taken by the ROV. From collections and video observations, eight species of hydroids...
Forty-two species of hydroids, excluding stylasterids, are reported in the present collection from the Northwestern
Hawaiian Islands. Of these, four are anthoathecates and 38 are leptothecates. Among the latter, Sertularella affinicostata
and Monotheca gibbosa are described as new species. The binomen Halopteris longibrachia is proposed as a new
re...
The hydroids of Cocos Island (Isla del Coco), Costa Rica, have received scant attention and are poorly known. Only 11 species have been reported from there previously, with five of them being stylasterids. Hydroids examined here were collected during 2019 in a search for invasive species, as part of a fouling survey. Fourteen species – three anthoa...
Laura Roscoe Thornely was born on 6 March 1860 in Blundellsands, Lancashire, England. On approaching age 30, in the late 1880s, she became interested in the taxonomy of hydroids and bryozoans under Prof. W.A. Herdman at University College, Liverpool. Her studies during much of the 1890s were based largely on the relatively well-known fauna of the b...
Tima nigroannulata sp. nov. is described from medusae collected in shallow waters of four prefectures on the Pacific coast of Japan (Miyagi, Fukushima, Kanagawa, and Miyazaki), as well as from cultures maintained at two aquaria (Enoshima Aquarium, Kanagawa Prefecture; Tsuruoka City Kamo Aquarium, Yamagata Prefecture). Adult medusae differ from thos...
Massive accumulations of pelagic species of Sargassum have generated recent social, economic and ecological problems along Caribbean shores. In the Mexican Caribbean, these events have prompted the study of diverse biological and ecological aspects of these macroalgae. However, studies on their associated biota, including Hydrozoa, remain scarce. T...
This report is based on a small collection of hydroids from the Hawaiian Islands, in the central Pacific Ocean. Most of the examined material was obtained by staff of the Bernice Pauahi Bishop Museum, Honolulu, during surveys for nonindigenous marine species in shallow, sheltered, inshore or nearshore waters, and especially in harbours, bays, and l...
Sixty species of hydroids, assigned to 24 families and 39 genera, are recognized and discussed in a collection of material from the southwest coast of Florida. One new species (Clytia joycei) is described from turtlegrass (Thalassia testudinum) and reported as well from the Caribbean coast of Panama. Under provisions of the First Reviser Principle...
An account is given of hydroids collected in 2015 and 2016 from port and harbor fouling communities in the Galápagos Islands. Also included is the hydroid of Ectopleura media, discovered on the wreck of the tanker Jessica near Isla San Cristóbal in 2001. Among 20 species reported herein were six anthoathecates and 14 leptothecates. Most common in t...
Twenty-eight species of hydroids are now known from Japanese tsunami marine debris (JTMD) sent to sea in March 2011 from the Island of Honshu and landing between 2012 and 2016 in North America and Hawai‘i. To 12 JTMD hydroid species previously reported, we add an additional 16 species. Fourteen species (50%) were detected only once; given the small...
Two new species of hydroids, Eudendrium bleakneyi and Halecium praeparvum, are described from the Bay of Fundy. Fourteen others, Tubularia acadiae Petersen, 1990, Coryne pusilla Gaertner, 1774, Sarsia lovenii (M. Sars, 1846), Zanclea implexa (Alder, 1856), Corydendrium dispar Kramp, 1935, Rhizogeton fusiformis L. Agassiz, 1862, Bougainvillia muscus...
Prior to 1800, exploring expeditions were usually of a commercial and political nature, undertaken to colonize or expand relations with new countries and territories. Among those to be explored as the 19th century progressed was Brazil, the largest nation in South America. This account is a brief historical overview of some 19th and early 20th cent...
One new genus (Schizoplumularia) and three new species (Schizoplumularia vervoorti, S. geniculata and S. elegans) of plumulariids are recognized and described from large collections of plumularioid hydroids collected in New Cale-donia and vicinity during several French expeditions. During taxonomic studies of these hydroids, colonies were com-pared...
The binomen Nemopsis gibbesii McCrady, 1859, originally applied to a species of hydromedusa and its supposed hydroid from South Carolina, USA, has been known for more than a century to encompass two species. The medusa stage is conspecific with that of Nemopsis bachei L. Agassiz, 1849, while the hydroid stage is referable to the genus Margelopsis H...
George James Allman (1812-1898), acclaimed for pioneering studies of Hydrozoa and Bryozoa, was born in Cork, Ireland. He earned B.A. (1839) and M.B. (1843) degrees at Trinity College, Dublin, and an M.D. (1847) from Trinity College and Oxford University. Choosing academia over medicine, he served as Professor of Botany, University of Dublin (1844-1...
The hydroid Similiclava nivea, gen. nov., sp. nov., is described from colonies collected in nearshore waters of southern British Columbia, Canada. It has been observed by divers, and recorded as Clava sp., several hundred times at locations between southeast Alaska and southern Oregon, USA. While resembling the hydractiniid Clava multicornis, tenta...
Fourteen species of hydroids, including two anthoathecates and 12 leptothecates, are reported from the west coast of North America on debris from the tsunami that struck Japan on 11 March 2011. Six species were found on a dock that stranded at Agate Beach, Newport, Oregon, five from a boat at Gleneden Beach, Oregon, four from a dock in Olympic Nati...
Axel Elof Jäderholm was born in Söderhamn, Sweden, on 24 July 1868. In 1888 he entered Uppsala Universitet, earning undergraduate (1892) and doctorate (1898) degrees. His doctoral dissertation was based on an anatomical study of South American Peperomia (Piperaceae). While a graduate student he commenced research on hydroids in collections at the u...
The shallow-water hydroid Clava multicornis is reported from the west coast of North America for the first time in nearly six decades. A North Atlantic species often occurring on intertidal fucoid algae, it had been found elsewhere only in heavily-invaded San Francisco Bay, California, with records dating from the late 19 th to mid-20 th centuries....
The shallow-water hydroid Clava multicornis is reported from the west coast of North America for the first time in nearly six decades. A North Atlantic species often occurring on intertidal fucoid algae, it had been found elsewhere only in heavily-invaded San Francisco Bay, California, with records dating from the late 19 th to mid-20 th centuries....
Five species of aglaopheniid hydroids (Aglaophenopsis cornuta, Cladocarpus diana, C. formosus, C. integer, and Nematocarpus ramuliferus) were collected from the Flemish Cap, Flemish Pass, and Grand Banks of Newfoundland during surveys with bottom trawls, rock dredges, and scallop gear. All are infrequently reported species, with C. diana being disc...
Zyzzyzus rubusidaeus, sp. nov., is described from inshore waters near the northern tip of Vancouver Island, British Co-lumbia, Canada. Specimens were collected on rocky bottoms amongst barnacles, sponges, and compound ascidians at a depth of 18 m in Weynton Passage, Broughton Strait, during March, July, and October 2012. Polyps tend to grow in dens...
This paper gives a systematic account of 67 species, referable to 22 families and 40 genera, identified in a small collection of hydroids from the central Atlantic coast of Florida between Melbourne and Palm Beach. The fauna mostly comprises an assemblage of tropical western Atlantic species ranging northwards along the southeastern coast of the Un...
The leptothecate hydroid Sertularella mutsuensis Stechow, 1931 is reported on debris from the 2011 Japanese tsunami that came ashore on 5 June 2012 at Agate Beach north of Newport, Oregon. Its discovery on a barnacle (Semibalanus cariosus) from a derelict floating dock originating at Misawa, Honshu, confirms the capability of successful transoceani...
Harry Beal Torrey was born on 22 May 1873 in Boston, Massachusetts. Two years later his family moved to Oakland, California. Torrey earned B.S. and M.S. degrees in zoology from the University of California, Berkeley, in 1895 and 1898 respectively, a Ph.D. in zoology from Columbia University in 1903, and an M.D. from the Medical College of Cornell U...
Sertularella maureenae, n. sp. (Hydrozoa: Sertulariidae), is described from the Pacific coast of Canada. The species is characterized by its predominantly stolonal colony form with pedicellate, annulated hydrothecae, together with the presence of large, ovate gonothecae having distinct annulations, a prominent neck, and 4–6 well-developed cusps sur...
An account is given of 48 species identified in a small collection of hydroids from the west coast of Sweden. Assigned to 39 genera and 18 families, most are well-known from boreal waters of the North Atlantic Ocean. Campanulina panicula G.O. Sars, 1874 is referred to Racemoramus, a new genus of leptothecates distinguished in part by having monopod...
Based largely on collections from the Calanus–Salvelinus expeditions, 54 species of thecate hydroids were identified from the shelf waters of northern Canada between northeastern Newfoundland and the Alaska–Yukon border. Common species included Halecium muricatum, Calycella syringa, Campanularia integra, C. speciosa, C. volubilis, Gonothyraea loven...
Sixteen species of athecate hydroids were identified in collections from the shelf waters of northern Canada between the Strait of Belle Isle and the Alaska–Yukon border. This brings the number of athecate species reported from Canada's north coast to 21, and the total number of hydroid species known from the region to 92. One species, Eudendrium c...
Boreohydra simplex was collected at a depth of 400 m in Cabot Strait, eastern Canada; this solitary, mud-dwelling hydroid is previously unreported from the western North Atlantic. Elsewhere, it has been found along the coasts of Scandinavia, Britain, and Iceland in the North Atlantic, and from South Georgia in the South Atlantic.
Shallow-water (0–100 m depth) hydroid faunas reported from 26 locations along the western North Atlantic coast between the high Canadian Arctic archipelago and the Caribbean Sea were compared. Species numbers varied widely between locations, but were highest in the tropics and subtropics, lowest in arctic and subarctic waters, and intermediate in m...
Bougainvillia aberrans n.sp. is described from Bermuda in the western North Atlantic Ocean. Specimens were collected at a depth of 150 fathoms (274 m) from the polypropylene buoy line of a crab trap. The hydroid colony of B. aberrans is erect, with a polysiphonic hydrocaulus, a smooth to somewhat wrinkled perisarc, hydranths having a maximum of abo...
Hydroid diversity and abundance appear to be low in the mid-abyssal zone of the western North Atlantic Ocean. Only two species (Acryptolaria longitheca,?Opercularella sp.) were collected during investigations by submersible (Deep Submergence Vehicle Alvin) at depths between 3011 and 3550 m along the northwest slope of the Bermuda Pedestal in March...
Plicatotheca anitae, n.gen., n.sp. (Hydrozoa: Phialellidae) is described from material collected on coral rubble at a depth of 60–90 m off Bermuda. The species has previously been reported from a depth of 430 m off Durban, South Africa, as Opercularella sp. Plicatotheca is distinguishable from the related genera Opercularella, Dipleuron, and Phiale...
Gustav Heinrich Kirchenpauer was born 2 February 1808 in Hamburg, and he died in that city on 3 March 1887. In 1810 he was taken with his family to St. Petersburg, Russia, to escape economic hardships during the occupation of Hamburg and continental blockade of the United Kingdom by Napoleon. He attended secondary school and university in Dorpat (T...
This article examines the life and zoological research of Jesse Walter Fewkes of the United States, whose later career as a pioneer in cultural anthropology overshadowed his significant earlier contributions to zoology. The primary focus of his zoological work was on planktonic Cnidaria, especially the Siphonophora. He also carried out detailed inv...
A systematic account is given of 17 families, 25 genera, and 32 species of anthoathecate hydroids and limnopolyps reported from Hawaii. Applying Reversal of Precedence provisions in zoological nomenclature, the familiar hydrozoan genus names Hydractinia Van Beneden, 1844a, Bimeria Wright, 1859a, and Porpita Lamarck, 1801 are designated as valid and...
Gustav Heinrich Kirchenpauer was born 2 February 1808 in Hamburg, and he died in that city on 3 March 1887. In 1810 he was taken with his family to St. Petersburg, Russia, to escape economic hardships during the occupation of Hamburg and continental blockade of the United Kingdom by Napoleon. He attended secondary school and university in Dorpat (T...
Thomas Hincks was born 15 July 1818 in Exeter, England. He attended Manchester New College, York, from 1833 to 1839, and received a B.A. from the University of London in 1840. In 1839 he commenced a 30-year career as a cleric, and served with distinction at Unitarian chapels in Ireland and England. Meanwhile, he enthusiastically pursued interests i...
Charles Wesley Hargitt was born near Lawrenceburg, Indiana, USA, and died at Syracuse, New York. After a brief career as a Methodist Episcopal Minister, he carried out graduate studies in biology at Illinois Wesleyan University and Ohio University. He served briefly on the faculty at Moores Hill College and later at Miami University of Ohio before...
Hydroids of the Allan Hancock Pacific Expeditions, and those of the Allan Hancock Caribbean Sea Expedition, were examined by Charles McLean Fraser in a series of reports published between 1938 and 1948. A total of 159 new nominal species was described from material collected in the eastern Pacific between Peru and southern British Columbia. Account...
Hydroids were examined in three different collections of specimens, acquired in 1969, 2002, and 2004, from the Caribbean coast of Panama. Eighteen stations were sampled in the Bocas del Toro area, western Panama. Nine others were situated in Colón and vicinity, and a single station was made at Portobelo in the east. Seventy-nine species were identi...
Charles Cleveland Nutting (1858-1927) began his academic career at the University of Iowa. and at the Natural History Museum of the university, in 1886. A naturalist with a bent for ornithology. he became interested in cnidarians, and especially hydroids and alcyonarians. as an outcome of expeditions to the Bahamas in 1888 and 1893. Most noteworthy...
Charles Cleveland Nutting (1858–1927) began his academic career at the University of Iowa, and at the Natural History Museum
of the university, in 1886. A naturalist with a bent for ornithology, he became interested in cnidarians, and especially hydroids
and alcyonarians, as an outcome of expeditions to the Bahamas in 1888 and 1893. Most noteworthy...
Hydroids were examined in collections from a biological survey of Northumberland Strait undertaken by Fisheries and Oceans Canada from June to August 1975. No investigations have been undertaken previously on hydroids of the study area. Forty-eight species referable to 12 families were present in the samples, with Sertulariidae dominating in both n...
The hydroid fauna of the Galápagos Islands, Ecuador, is known largely from reports of collections made between 1932 and 1938 during several Allan Hancock Pacific Expeditions. Of some 100 nominal species (excluding Stylasteridae) reported from the archipelago overall, including species obtained during other expeditions, 81 are recognized as valid. A...
This report provides the first general account of the marine hydroids (Leptolida), excluding Milleporidae and Stylasteridae, of Guam and the Commonwealth of the Northern Marianas Islands (CNMI). The siphonophore Physalia physalis is also recorded from the region. Seventeen families and 43 species, a majority of them (80%) leptothecates, are listed...
Three seamounts flanking the oceanic island of Bermuda were sampled for hydroids. Collecting was undertaken by submersible (SDL-1) and by dredge at depths between 48 and 107 m on the summits of Argus and Challenger banks. A shallower collection (<20 m) from the pilings of a tower on Argus Bank was made using SCUBA. Major bottom types on both banks...
Eudendrium bathyalis, a new species, is described from relatively deep-water (283 m) on the offshore slope of the oceanic island of Bermuda. Its distinctive characters are the small size of the colony, the cnidome, comprising two sizes of microbasic euryteles and the unreduced female blastostyle, bearing eggs supported by long peduncles.
The purpose of this application is to conserve the name Clytia noliformis (McCrady, 1859) for a well-known marine hydroid (family campanulariidae). McCrady's (1859) type material of C noliformis is lost but the hydroid he described is now believed to have been adifferent species from C noliformis auct. and perhaps conspecific with C hemisphaerica (...
Hydroids were examined in 88 qualitative samples from a depth range of 0—4550 m in the Bermuda area. Of 89 species represented overall, 54 were present in samples from 0 to 25 m. Thirty-one of these 54 taxa were found only within that bathymetric zone. Hydroid species numbers remained relatively high (36) in collections from shallow neritic bottoms...
An account is given of Sertularia vervoorti spec. nov., a small leptothecate species collected in coastal waters of southeastern Brazil. Specimens were found on brown algae, collected at depths of 15-20 m on the inner continental shelf off Espírito Santo state. This hydroid resembles Sertularia tumida All-man, 1877, S. linealis Warren, 1908, and S....
Dry season distribution of hydroids in a small tropical estuary, Pernam-buco, Brazil. Zool. Verh. Leiden 323, 31.xii.1998: 69-78, figs 1-3.— Hydroid distribution patterns along a horizontal ecocline in the Rio Formoso/Rio Ariquindá/Rio Porto Alegre system, a small and seasonally poikilohaline estuary on the tropical northeast coast of Brazil, were...