Clint Boyd

Clint Boyd
North Dakota Industrial Commission Department of Mineral Resources · Fossil Resource Protection Program

PhD

About

53
Publications
13,008
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571
Citations
Citations since 2017
25 Research Items
308 Citations
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20172018201920202021202220230102030405060
20172018201920202021202220230102030405060
Introduction
I currently serve as the Curator of the State Fossil Collection and manager of the paleontology program for the North Dakota Geological Survey.
Additional affiliations
January 2015 - present
North Dakota Geological Survey
Position
  • Senior Paleontologist and Curator
Description
  • Manage the NDGS paleontology program and serve as curator of the North Dakota state fossil collection.
September 2012 - January 2015
South Dakota School of Mines and Technology
Position
  • Haslem Postdoctoral Fellow
January 2003 - July 2005
University of Wisconsin–Madison
Position
  • Fossil Preparation Lab Supervisor

Publications

Publications (53)
Article
Full-text available
The systematic relationships of taxa traditionally referred to as ‘basal ornithopods’ or ‘hypsilophodontids’ remain poorly resolved since it was discovered that these taxa are not a monophyletic group, but rather a paraphyletic set of neornithischian taxa. Thus, even as the known diversity of these taxa has dramatically increased over the past two...
Article
Full-text available
Though the dinosaur Thescelosaurus neglectus was first described in 1913 and is known from the relatively fossiliferous Lance and Hell Creek formations in the Western Interior Basin of North America, the cranial anatomy of this species remains poorly understood. The only cranial material confidently referred to this species are three fragmentary bo...
Article
Full-text available
Improvements in the perceived completeness of the fossil record may be driven both by new discoveries and by reinterpretation of known fossils, but disentangling the relative effects of these processes can be difficult. Here, we propose a new methodology for evaluating historical trends in the perceived completeness of the fossil record, demonstrat...
Article
Full-text available
Three new vertebrate localities are reported from within the Bloom Basin of the North Unit of Badlands National Park, Interior, South Dakota. These sites were discovered during paleontological surveys and monitoring of the park’s boundary fence construction activities. This report focuses on a new fauna recovered from one of these localities (BADL-...
Article
Full-text available
Approaches quantifying the relative congruence, or incongruence, of molecular divergence estimates and the fossil record have been limited. Previously proposed methods are largely node specific, assessing incongruence at particular nodes for which both fossil data and molecular divergence estimates are available. These existing metrics, and other m...
Article
Full-text available
Removal or protection from biostratinomic agents of decomposition, such as predators and scavengers, is widely seen as a requirement for high-quality preservation of soft tissues in the fossil record. In this context, extremely rapid burial is an oft-cited mechanism for shielding remains from degradation, but not all fossils fit nicely into this pa...
Technical Report
Full-text available
A report on the paleontological resources found in Theodore Roosevelt National Park in western North Dakota including local geology, taxonomy of local fossils, fossil localities within the park, museum collections and paleontological archives of materials found in the park, a review of park paleontological research, paleontological interpretation,...
Article
Full-text available
For over a century the mounds of harvester ants (Pogonomyrmex spp.) have been targeted by paleontological field crews as sources of concentrated microvertebrate fossils that facilitate the collection of large numbers of specimens with a small investment of time. This study describes a collection of over 6,000 identifiable micromammal teeth and jaws...
Article
Full-text available
Ornithischians form a large clade of globally distributed Mesozoic dinosaurs, and represent one of their three major radiations. Throughout their evolutionary history, exceeding 134 million years, ornithischians evolved considerable morphological disparity, expressed especially through the cranial and osteodermal features of their most distinguisha...
Article
Full-text available
The upper Maastrichtian Breien Member situated within the lower portion of the Hell Creek Formation in south-central North Dakota records one of the last transgressions of the Western Interior Seaway (WIS) during the terminal Cretaceous. A fragmentary articular-prearticular complex and isolated vertebra belonging to a mosasauroid were recovered in...
Article
Full-text available
An anomalous specimen of Dinictis felina (Carnivora: Nimravidae) from the Scenic Member of the Brule Formation in South Dakota was recently identified possessing bilaterally symmetrically split carinae on the anterior surfaces of the upper canines. The anterior carinae display normal serration patterns from the apex until they bifurcate towards the...
Article
Full-text available
Five marsupial species are recognized from the Brule Formation at two localities in southwestern North Dakota: Fitterer Ranch and Obritsch Ranch (middle Oligocene; Whitneyan North American Land Mammal Age [NALMA]). The herpetotheriids Herpetotherium fugax Cope, 1873a, Copedelphys superstes new species, and the peradectid Nanodelphys hunti (Cope, 18...
Article
Full-text available
In a recent review of the North American rodent fossil record during the Whitneyan biozone (32.0 to 30.0 Ma) a publication error resulted in the table containing the primary data on reported occurrences of rodent taxa not being published in its entirety. This report publishes the full Whitneyan data table used in that study, along with additional t...
Article
Full-text available
Until recently, few well-described rodent faunae from the Whitneyan North American Land Mammal Age [NALMA] were known, hindering studies of rodent diversity, biogeography, and evolutionary patterns during the Oligocene. This study describes a new Whitneyan rodent assemblage from the Obritsch Ranch paleontological locality in the Little Badlands reg...
Article
Full-text available
Eighteen rodent species are recognized from the Fitterer Ranch fauna of North Dakota on the basis of more than a thousand collected specimens. Of the species recognized, four are new: the prosciurine aplodontiids Prosciurus hogansoni and Altasciurus leonardi, the heliscomyid Heliscomys borealis, and the cricetid Eumys lammersi. A previously describ...
Article
Full-text available
This article describes an astragalus from an amynodontid, representing the tribe Metamynodontini from the Arikaree Group.
Article
Full-text available
Specimen F:AM 99259, a well-preserved skull with articulated dentaries representing the earliest ontogenetic stage of development yet observed for a North American nimravid, is occasionally mentioned in the scientific literature. Conflicting reports refer that specimen to either Eusmilus or Nimravus, with no evidence provided to support those refer...
Conference Paper
Full-text available
Three new vertebrate localities are reported from within the Bloom Basin of the North Unit of Badlands National Park, Interior, South Dakota. These sites were discovered during paleontological surveys and monitoring of the park’s boundary fence construction activities. This report focuses on a new fauna recovered from one of these localities (BADL-...
Article
Avian remains are rare from Paleogene sediments in the Great Plains region of North America, especially those that can be identified taxonomically. Here, we describe avian eggshell fragments from the Fitterer Ranch locality within the Oligocene Brule Formation of North Dakota and assign them to ootaxonomy. The specimen examined (NDGS 1934) exhibits...
Preprint
Full-text available
The amphicyonid, or “bear-dog,” Paradaphoenus was a small-bodied, hypercarnivorous carnivoran that lived in North America during the Oligocene. Specimens of Paradaphoenus are rare, in part owing to the common inaccurate referral of specimens to the abundant canid Hesperocyon , which was similar in size and morphology. Four new specimens of Paradaph...
Preprint
Full-text available
The amphicyonid, or “bear-dog,” Paradaphoenus was a small-bodied, hypercarnivorous carnivoran that lived in North America during the Oligocene. Specimens of Paradaphoenus are rare, in part owing to the common inaccurate referral of specimens to the abundant canid Hesperocyon , which was similar in size and morphology. Four new specimens of Paradaph...
Article
Full-text available
This report re-evaluates the depositional history, biostratigraphic “age,” and regional correlates of Chadron Formation rocks in North Dakota using a variety of data sources. A series of well-developed paleosols are herein recognized for the first time at the top of the Chalky Buttes Member across much of southwestern North Dakota. Those paleosols...
Conference Paper
Full-text available
Neornithischians were a major clade of herbivorous dinosaurs that existed from the earliest Jurassic to the latest Maastrichtian. During their long evolutionary history, neornithischians diverged into several highly distinctive clades, including ceratopsians, pachycephalosaurs, and hadrosaurids. However, the origin and the early evolution of Neorni...
Article
During their long evolutionary history, neornithischian dinosaurs diverged into several clades with distinctive adaptations. However, the early evolution within Neornithischia and the resolution of the phylogenetic relationships of taxa situated near the base of the clade remain problematic. This is especially true for those taxa traditionally plac...
Preprint
Full-text available
Crocodyliforms serve as important taphonomic agents, accumulating and modifying vertebrate remains. Previousdiscussions of Mesozoic crocodyliform feeding in terrestrial and riverine ecosystems have often focused on larger taxa and their interactions with equally large dinosaurian prey. However, recent evidence suggests that the impact of smaller cr...
Article
Full-text available
Gaining a detailed understanding of the geographic and temporal variations between Lancian mammalian faunas within the Western Interior Basin requires specimens to be associated with precise stratigraphic data, but only a few mammal-producing localities are documented in such detail from the Hell Creek Formation of southwestern North Dakota. Recent...
Conference Paper
Full-text available
The reported vertebrate fauna of the Brule Formation within North Dakota is largely based on preliminary faunal lists presented without detailed discussion. An effort is underway to refine our knowledge of that fauna, including a thorough review of the microvertebrates collected in the Little Badlands area (Stark County, North Dakota) and held with...
Article
Full-text available
Knowledge regarding the early evolution within the dinosaurian clade Ankylopollexia drastically increased over the past two decades, in part because of an increase in described taxa from the Early Cretaceous of North America. These advances motivated the recent completion of extensive preparation and conservation work on the holotype and only known...
Preprint
Full-text available
Knowledge regarding the early evolution within the dinosaurian clade Ankylopollexia drastically increased over the past two decades, in part because of an increase in described taxa from the Early Cretaceous of North America. These advances motivated the recent completion of extensive preparation and conservation work on the holotype and only known...
Preprint
Full-text available
Knowledge regarding the early evolution within the dinosaurian clade Ankylopollexia drastically increased over the past two decades, in part because of an increase in described taxa from the Early Cretaceous of North America. These advances motivated the recent completion of extensive preparation and conservation work on the holotype and only known...
Preprint
Full-text available
Knowledge regarding the early evolution within the dinosaurian clade Ankylopollexia drastically increased over the past two decades, in part because of an increase in described taxa from the Early Cretaceous of North America. These advances motivated the recent completion of extensive preparation and conservation work on the holotype and only known...
Article
Full-text available
The synonymization of the cimolestan taxa Cymaprimadon and Chadronia from the Late Eocene Chadron Formation is consistently upheld, despite a lack of supporting evidence. Here we show that the synonymization is unjustified, owing to distinct differences between these taxa in the mandibular tooth count (1-1-3-3 vs ?-1-4-3), the identity of the enlar...
Chapter
Full-text available
Ornithopod dinosaurs were bipedal, herbivorous dinosaurs represented in the Late Cretaceous of North Amer-ica by basal ornithopods ("hypsilophodontids") and a clade of derived iguanodontians containing, in part, hadrosaurids. Recent research focused on the Cretaceous macrovertebrates of Grand Staircase–Escalante National Monument and sur-rounding a...
Article
Full-text available
Crocodyliforms serve as important taphonomic agents, accumulating and modifying vertebrate remains. Previous discussions of Mesozoic crocodyliform feeding in terrestrial and riverine ecosystems have often focused on larger taxa and their interactions with equally large dinosaurian prey. However, recent evidence suggests that the impact of smaller c...
Article
Full-text available
Intercostal plates are bony structures positioned lateral to the anterior dorsal ribs in some ornithischian dinosaurs. Some propose these plates are homologous, or functionally analogous, with the uncinate processes of extant avian dinosaurs that assist in breathing, while others suggest they served a defensive function. To elucidate their osteogen...
Article
A small, articulated basal ornithopod skeleton from the Frenchman Formation (late Maastrichtian) of Saskatchewan (RSM P 1225.1), previously referred to the taxon Thescelosaurus, differs from both recognized species of this taxon (Thescelosaurus neglectus and Thescelosaurus garbanii). The differences are taxonomically informative and we recognize th...
Article
The stratigraphic record of first appearances provides an independent source of data for evaluating and comparing phylogenetic hypotheses that include taxa with fossil histories. However, no standardized method exists for calculating these metrics for polytomous phylogenies, restricting their applicability. Previously proposed methods insufficientl...
Article
Full-text available
Considerable controversy surrounds the taxonomy and phylogenetic relationships of Maastrichtian basal neornithischian taxa from North America. Discovery of previously unrecognized cranial material from the paratype specimen of Thescelosaurus neglectus (USNM 7758), along with the examination of two new specimens that preserve nearly complete skulls...
Article
Full-text available
New penguin fossils from the Eocene of Peru force a reevaluation of previous hypotheses regarding the causal role of climate change in penguin evolution. Repeatedly it has been proposed that penguins originated in high southern latitudes and arrived at equatorial regions relatively recently (e.g., 4–8 million years ago), well after the onset of lat...

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