Ceri Shipton

• PhD
• Lecturer at University College London

This paper assesses the evidence for cumulative culture in the Palaeolithic through the lens of the most widely available line of evidence: knapped stone. Two types of cumulative culture are defined: additive traits in an individual’s repertoire, versus a population wide stock of skills. Complexity may both cumulate within a single realm of experti...

The handaxe is an iconic stone tool form used to define and symbolise both the Acheulean and the wider Palaeolithic. There has long been debate around the extent of its morphological variability between sites, and the role that extrinsic factors (especially raw material, blank type, and the extent of resharpening) have played in driving this variab...

Archaeological evidence attests multiple early dispersals of Homo sapiens out of Africa, but genetic evidence points to the primacy of a single dispersal 70-40 ka. Laili in Timor-Leste is on the southern dispersal route between Eurasia and Australasia and has the earliest record of human occupation in the eastern Wallacean archipelago. New evidence...

This paper re-examines earlier Palaeolithic core technology from British sites assigned to MIS 11, 9, and 7 using primarily a châine opératoire approach, with the objective of better understanding the earliest occurrence and distribution of Levallois and other prepared-core technologies across the Old World. Contrary to previous interpretations (Wh...

The presence or absence of handaxes endures as the major criterion of Lower Palaeolithic classification, with contemporaneous core-and-flake industries modelled as simpler counterparts to Acheulean technology. This is based on the supposed absence of formal tools, particularly of large cutting tools (LCTs) which are understood to be important withi...

In this chapter we examine the evidence for modern human dispersal, early settlement and later adaptations to the southern islands of the Wallacean Archipelago. We discuss the features that distinguish modern human occupation in southern Wallacea during the Pleistocene from those in the northern islands. In this context we examine the location of s...

Asitau Kuru provides a unique record of human behaviour from the first arrival of Homo sapiens onto Timor Island around 44,000 years ago through to the near present. In particular, this site has produced a large number of marine shell artefacts which have been central to rewriting our understanding of the sophistication of cultural behaviours enact...

This handbook is currently in development, with individual articles publishing online in advance of print publication. At this time, we cannot add information about unpublished articles in this handbook, however the table of contents will continue to grow as additional articles pass through the review process and are added to the site. Please note...

Archaeological sites with surface hearths are a ubiquitous feature across the arid zones of the Arabian interior. At Jebel Oraf, in the Jubbah basin of the Nefud Desert of northern Arabia, numerous grinding stone fragments were found in association with hearths, though the original purpose of these stones was unclear owing to the poor preservation...

The Neolithic of northern Arabia is characterised by monumental stone structures, ephemeral ‘hearth sites’ indicative of a highly mobile lifestyle, and a rich rock art heritage with iconic representations of domesticated livestock. However, the character and timing of occupation prior to the spread of pastoralism (ca. 6000 BCE) remains elusive, wit...

Panga ya Saidi is a large, open-roofed, solutional cave located in the Dzitsoni limestone hills that run parallel to the Kenyan coast, approximately halfway between Mombasa and Malindi (3.67° S, 39.74° E) (Fig. 1). The location of the site is of significance since Panga ya Saidi is a rare near-coastal Stone Age site, with most archaeological invest...

Homo sapiens were adept at fishing in a range of aquatic habitats by the time they left Africa and reached Southeast Asia ca. 73 kya. In the insular region of Wallacea, humans adapted to a significant maritime environment with sophisticated marine fishing methods and technology by at least 42 kya. However, despite a growing array of evidence sugges...

Reconstructing the technical and cognitive abilities of past hominins requires an understanding of how skills like stone toolmaking were learned and transmitted. We ask how much of the variability in the uptake of knapping skill is due to the characteristics of the knapping sequences themselves? Fundamental to skill acquisition is proceduralization...

The crossing of the Wallacean islands and settlement of Sahul by modern humans over 50,000 years ago, represents the earliest successful seafaring of our species anywhere in the world. Archaeological research throughout this vast island archipelago has recovered evidence for varied patterns in island occupation, with accumulating evidence suggestin...

This handbook is currently in development, with individual articles publishing online in advance of print publication. At this time, we cannot add information about unpublished articles in this handbook, however the table of contents will continue to grow as additional articles pass through the review process and are added to the site. Please note...

This article offers some hypotheses to explain Later Stone Age lithic miniaturization: the systematic creation of small stone flakes on the finest-grained materials. Fundamentally, this phenomenon appears to represent the prioritization of stone tool sharpness over longevity, and a disposable mode of using stone tools. Ethnographic evidence from Au...

The peopling of Sahul (the combined landmass of New Guinea and Australia) is a topic of much debate. The Kimberley region of Western Australia holds many of Australia's oldest known archaeological sites. Here, we review the chronological and archaeological data available for the Kimberley from early Marine Isotope Stage 3 to the present, linking ep...

The appearance of the Acheulean and the production of new bifacial tools marked a revolution in human behavior. The use of longer and complex operative chains, with centripetal and recurrent knapping, adapted to different raw materials, created long useful edges, converging in a functional distal end. How and why these handaxes vary has been the su...

Stone tools are a manifestation of the complex cognitive and dexterous skills of our hominin ancestors. As such, much research has been devoted to understanding the skill requirements of individual lithic technologies. Yet, comparing skill across different technologies, and thus across the vast timespan of the Palaeolithic, is an elusive goal. We s...

The Holocene of eastern Africa saw extreme climatic fluctuations between hyper-humid and arid conditions, which manifested differently across the region's lake basins, coastal ecotones, and terrestrial biomes. Changes to resource availability, distribution, and predictability presented different constraints and opportunities to diverse hunter-gathe...

Archaeologists have long emphasized the importance of large-scale excavations and multi-year or even decades-long projects at a single site or site complex. Here, we highlight archaeological field strategies, termed coring, profiling, and trenching (CPT), that rely on relatively small-scale excavations or the collection of new samples from intact d...

Levallois technology characterizes the Middle Stone Age/Middle Palaeolithic, but one of its earliest manifestations is from the preceding Acheulean of the Kapthurin Formation, in the Rift Valley of east Africa. Here, ~ 400 ka, hominins were creating large flake blank handaxes and cleavers through Levallois knapping. Comparing these tools with other...

Mainland Australia was connected to New Guinea and Tasmania at various times throughout the Pleistocene and formed the supercontinent of Sahul. Sahul contains some of the earliest known archaeological evidence for Homo sapiens outside of Africa, with a growing record of early complex social, technological, and artistic life. Here we present an over...

Pleistocene hominin dispersals out of, and back into, Africa necessarily involved traversing the diverse and often challenging environments of Southwest Asia 1–4 . Archaeological and palaeontological records from the Levantine woodland zone document major biological and cultural shifts, such as alternating occupations by Homo sapiens and Neandertha...

The antiquity and nature of coastal resource procurement is central to understanding human evolution and adaptations to complex environments. It has become increasingly apparent in global archaeological studies that the timing, characteristics, and trajectories of coastal resource use are highly variable. Within Africa, discussions of these issues...

Open Access Check for updates on crossmark Research articles Collagen fingerprinting traces the introduction of caprines to island Eastern Africa Courtney Culley, Anneke Janzen, Samantha Brown, Mary E. Prendergast, Jesse Wolfhagen, Bourhane Abderemane, Abdallah K. Ali, Othman Haji, Mark C. Horton, Ceri Shipton, Jillian Swift, Tabibou A. Tabibou, He...

Lake Woods is a large freshwater body on the northern edge of Australia’s arid zone where research since the late 1960s has suggested late Pleistocene, later Holocene, and recent human occupations. We report on surface collections and test excavations undertaken at four sites on the western side of the lake basin in 2019. Each site produced a disti...

Hearth sites are characteristic of Holocene occupation in the Arabian sand seas but remain mostly unstudied. Excavations of two multi-period hearth sites in the Jebel Oraf palaeolake basin, in the oasis of Jubbah, now substantially increase our knowledge of these sites. In total, 17 of 170 identified hearths were excavated at Jebel Oraf 2 (ORF2), a...

The origin and evolution of hominin mortuary practices are topics of intense interest and debate1–3. Human burials dated to the Middle Stone Age (MSA) are exceedingly rare in Africa and unknown in East Africa1–6. Here we describe the partial skeleton of a roughly 2.5- to 3.0-year-old child dating to 78.3 ± 4.1 thousand years ago, which was recovere...

‘Normativity’ refers to the human conformity to the behavioral modes of a society, which underpins diverse aspects of our behavior, including symbolism, cooperation, and morality. It has its developmental basis in overimitation, the uniquely human bias towards replicating the intentional actions of a demonstrator, regardless of their causal relevan...

The morphological differentiation of African bovids in highly fragmented zooarchaeological assemblages is a major hindrance to reconstructing the nature and spread of pastoralism in sub-Saharan Africa. Here we employ collagen peptide mass fingerprinting, known as Zooarchaeology by Mass Spectrometry (ZooMS), coupled with recently published African Z...

Stone artifacts from Makpan cave on Alor island date from ~40 ka, filling a gap in the Marine Isotope Stage (MIS) 3-2 record of southern Wallacea between Liang Bua on Flores to the west, and Asitau Kuru on Timor to the east. Since Alor is a largely volcanic island, the Makpan stone artifacts are dominated by igneous materials, distinguishing them f...

The Middle to Later Stone Age transition is a critical period of human behavioral change that has been variously argued to pertain to the emergence of modern cognition, substantial population growth, and major dispersals of Homo sapiens within and beyond Africa. However, there is little consensus about when the transition occurred, the geographic p...

The Wallacean archipelago between the Indian and Pacific Oceans is a critical biogeographic boundary for all kinds of animals, from butterflies to birds. Humans are no exception, and in this paper we offer a three stage model for how our genus overcame this boundary. We review how Lower Palaeolithic hominins were able to colonize the larger islands...

Mainland Australia was connected to New Guinea and Tasmania at various times throughout the Pleistocene and formed the supercontinent of Sahul. Sahul contains some of the earliest known archaeological evidence for Homo sapiens outside of Africa, with a growing record of early complex social, technological, and artistic life. Here we present an over...

We report archaeological findings from a significant new cave site on Alor Island, Indonesia, with an in situ basal date of 40,208–38,454 cal BP. Twenty thousand years older than the earliest Pleistocene site previously known from this island, Makpan retains dense midden deposits of marine shell, fish bone, urchin and crab remains, but few terrestr...

Engraving sites are rare in mainland and Island Southeast Asia (ISEA) where painted art dominates the prehistoric artistic record. Here we report two new engraving sites from the Tutuala region of Timor-Leste comprising mostly humanoid forms carved into speleothem columns in rock-shelters. Engraved face motifs have previously been reported from Len...

The first excavations on Obi Island, north-east Wallacea, reveal three phases of occupation beginning in the terminal Pleistocene. Ground shell artefacts appear at the end of the terminal Pleistocene, the earliest examples in Wallacea. In the subsequent early Holocene occupation phase, ground stone axe flakes appear, which are again the earliest ex...

Between 10 and six thousand years ago the Arabian Peninsula saw the most recent of the ‘Green Arabia’ periods, when increased rainfall transformed this generally arid region. The transition to the Neolithic in Arabia occurred during this period of climatic amelioration. Various forms of stone structures are abundant in northern Arabia, and it has b...

This paper introduces primary data on site contents, chronology and stratigraphy for four subsurface middens, which formed through the late Holocene on the Dampier Peninsula. Data from one surface midden collection are also presented. In this monsoonal coastal locality, variations in dune stability and sand flux are critical to archaeological site...

This chapter examines the issue of whether the Acheulean is a genuine homologous cultural entity, descended via a chain of social reproduction from a common ‘ancestor’, or whether it was a technological phase that was repeatedly independently invented. An anecdotal experiment is used to determine the relative ease of inventing biface knapping from...

Archaeological surveys and excavations in the Jebel Oraf palaeolake basin, north-western Saudi Arabia, have identified a well-preserved early- to mid-Holocene landscape. Two types of occupation site can be distinguished: nine small and ephemeral scatters from single occupation phases on the slopes of sand dunes and three hearth sites indicative of...

There is a large, if disparate, body of archaeological literature discussing specific instantiations of symbolic material culture and the possibility of ritual practices in Neanderthal populations. Despite this attention, however, no single synthesis exists that draws upon cognitive, psychological and cultural evolutionary theories of ritual. Here,...

The handaxes of north-western Europe are some of the most varied in the Acheulean world, with the meanings of that variation debated since the late nineteenth century. To reassess handaxe form in this region, we performed a 3D morphometric analysis of 150 handaxes from five British Acheulean assemblages: Boxgrove, High Lodge, Hitchin, Swanscombe Mi...

The resource-poor, isolated islands of Wallacea have been considered a major adaptive obstacle for hominins expanding into Australasia. Archaeological evidence has hinted that coastal adaptations in Homo sapiens enabled rapid island dispersal and settlement; however, there has been no means to directly test this proposition. Here, we apply stable c...

African Middle Stone Age (MSA) populations used pigments, manufactured and wore personal ornaments, made abstract engravings, and produced fully shaped bone tools. However, ongoing research across Africa reveals variability in the emergence of cultural innovations in the MSA and their subsequent development through the Later Stone Age (LSA). When p...

India is located at a critical geographic crossroads for understanding the dispersal of Homo sapiens out of Africa and into Asia and Oceania. Here we report evidence for long-term human occupation, spanning the last ~80 thousand years, at the site of Dhaba in the Middle Son River Valley of Central India. An unchanging stone tool industry is found a...

In this paper, we look at a situation of long-term continuity to understand the circumstances that mediate against behavioural change. Using newly excavated material from Asitau Kuru, Timor-Leste, we assess continuity in stone tool technology, as well as pigment and bead use over a span of 44,000 years. The sequence is divided into three occupation...

The ecological adaptations that stimulated the dispersal and technological strategies of our species during the Late Pleistocene remain hotly disputed, with some influential theories focusing on grassland biomes or marine resources as key drivers behind the rapid expansion and material culture innovations of Homo sapiens within and beyond Africa. H...

The environmental extremes of the Last Glacial Maximum and the subsequent warming and sea-level rise into the Holocene had profound implications for human behavior across much of the world. In northern New Guinea, the Maluku Islands, and the Philippines, shell adzes appear during this period alongside contact between islands. In this paper we prese...

Human social transmission is unrivalled in its precision and complexity. High-fidelity social transmission ensures each generation does not have to reinvent the wheel, while the sharing of knowledge and skills enables the extraordinary feats of technology and artistry. This chapter explores the evolutionary foundations of our high-fidelity social t...

Abstract The Acheulean is the longest lasting cultural–technological tradition in human evolutionary history. However, considerable gaps remain in understanding the chronology and geographical distribution of Acheulean hominins. We present the first chronometrically dated Acheulean site from the Arabian Peninsula, a vast and poorly known region tha...

The nature and trajectory of coastal and maritime adaptations, and the complex ways foraging economies have been structured to include both marine and terrestrial resources, are becoming key topics of interest in African archaeological research. There is, therefore, an increasing need to understand the longer-term context for more recent shifts in...

The origins of the South Asian microlithic is hotly debated, with introduction by colonizing Homo sapiens or local adaptive response to climate change, contributing the two main competing models. Here we review the lithic evidence from two regions (the Jurreru Valley and the Middle Son Valley) where well-dated sites spanning 85-6 ka provide detaile...

The Acheulean stone tools of Homo erectus and Homo heidelbergensis are the longest enduring of all archaeological cultures, lasting for 1.5 million years. Three competing hypotheses have been proposed to explain this longevity: that Acheulean technology lies in a zone of latent, easy to invent solutions to problems that H. erectus and H. heidelberg...

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Despite occupying a central geographic position, investigations of hominin populations in the Arabian Peninsula during the Lower Palaeolithic period are rare. The colonization of Eurasia below 55 degrees latitude indicates the success of the genus Homo in the Early and Middle Pleistocene, but the extent to which these hominins were capable of innov...

Statistical tests of Whalen excavation artefacts. (XLSX)

Acheulean bifaces dominate the archaeological record for 1.5 million years. The meaning behind the often symmetrical forms of these tools is the topic of considerable debate, with explanations ranging from effectiveness as a cutting tool to sexual display. Some, however, question whether the symmetry seen in many Acheulean bifaces is intentional at...

The originally published version of this Article contained an error in Fig. 3, whereby an additional unrelated graph was overlaid on top of the magnetic susceptibility plot. Furthermore, the Article title contained an error in the capitalisation of 'Stone Age'. Both of these errors have now been corrected in both the PDF and HTML versions of the Ar...

The Middle to Later Stone Age transition in Africa has been debated as a significant shift in human technological, cultural, and cognitive evolution. However, the majority of research on this transition is currently focused on southern Africa due to a lack of long-term, stratified sites across much of the African continent. Here, we report a 78,000...

The question "How common is convergence?" remains unanswered and may be unanswerable. Our examples indicate that even the minimum detectable levels of convergence are often high, and we conclude that at all levels convergence has been greatly underestimated.-Moore and Willmer (1997: 1) Convergence in stone-tool technology, much like in biology, was...

Scholars from a variety of disciplines consider cases of convergence in lithic technology, when functional or developmental constraints result in similar forms in independent lineages. Hominins began using stone tools at least 2.6 million years ago, perhaps even 3.4 million years ago. Given the nearly ubiquitous use of stone tools by humans and the...

Over the 1.5-million-year duration of the Acheulean, there is considerable variation in biface finesse. It is not clear, however, if there is an improvement in biface knapping ability over time, or if variation between sites is largely unrelated to their age. The diversity and duration of the East African Acheulean presents an opportunity to examin...

Aim Our knowledge of the prehistoric distribution of animal species is so far largely dependent on the location of excavated archaeological and palaeontological sites. In the absence of excavated faunal remains, many species that were present in the Levant and North Africa have been assumed to have been absent on the Arabian Peninsula. Here, we exp...

A systematic survey of rock art and associated archaeological features in the Jubbah oasis provides evidence of Holocene occupation from the early Holocene to the present. In total 1249 panels with rock art and inscriptions, and 159 archaeological sites, were recorded on twelve different jebels. Analyses of rock art content and engraving stratigrap...

We assembled genome-wide data from 16 prehistoric Africans. We show that the anciently divergent lineage that comprises the primary ancestry of the southern African San had a wider distribution in the past, contributing approximately two-thirds of the ancestry of Malawi hunter-gatherers ∼8,100–2,500 years ago and approximately one-third of the ance...

Human-mediated biological exchange has had global social and ecological impacts. In sub-Saharan Africa, several domestic and commensal animals were introduced from Asia in the pre-modern period; however, the timing and nature of these introductions remain contentious. One model supports introduction to the eastern African coast after the mid-first...

Human-mediated biological exchange has had global social and ecological impacts. In sub-Saharan Africa, several domestic and commensal animals were introduced from Asia in the pre-modern period; however, the timing and nature of these introductions remain contentious. One model supports introduction to the eastern African coast after the mid-first...

Details of methods used in ancient DNA (aDNA) and Zooarchaeology by Mass Spectrometry (ZooMS) collagen fingerprinting analyses. (DOCX)

Total reads used in the BLAST analysis and results of Burrows-Wheeler Alignments (BWA). (DOCX)

Results of experimental study of false positives. Incorrect genus identifications resulting from 500 test "libraries" obtained from whole mtDNA genomes of the genus Gallus. See text for explanation of experimental method. (DOCX)

Reference specimens for ZooMS collagen fingerprinting. (DOCX)

Decision tree illustrating research protocols. Tree illustrates the selection of faunal samples, the order in which specific analyses were applied to each subsample, and result. (TIF)

Landmarks used in dental analysis. R. exulans tooth in occlusal view with simplified diagram to the right. The fixed landmarks are illustrated by large blue circles, sliding semi-landmarks by small red circles. The boundaries of the cusps and the stylids (small flat or saddle like surfaces joining cusps) are difficult to precisely identify, but hav...