Borja Holgado

Borja Holgado
Federal University of Rio de Janeiro | UFRJ · Departamento de Geologia e Paleontologia

BSc in Biology, MSc in Palaeontology

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24
Publications
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146
Citations

Publications

Publications (24)
Article
Full-text available
Faxinalipterus minimus was originally described as a purported pterosaur from the Late Triassic (early Norian) Caturrita Formation of southern Brazil. Its holotype comprises fragmentary postcranial elements, whereas a partial maxilla was referred to the species. The assignment of Faxinalipterus minimus to Pterosauria has been questioned by some stu...
Article
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Amblydectes is a problematic genus proposed more than a century ago for several pterosaur specimens from the Cambridge Greensand. Its problematic nature is due to the fragmentary preservation of the referred specimens, limited to several rostral tips. In the present work is reassessed the validity of Amblydectes crassidens based on new anatomical c...
Article
Full-text available
Research on the postcranial skeletal pneumaticity in pterosaurs is common in the literature, but most studies present only qualitative assessments. When quantitative, they are done on isolated bones. Here, we estimate the Air Space Proportion (ASP) obtained from micro-CT scans of the sequence from the sixth cervical to the fourth dorsal vertebra of...
Article
Aerotitan sudamericanus, from the Upper Cretaceous of the Neuquén Basin (Patagonia, Argentina), is known from a partial jaw fragment which has been interpreted as either an azhdarchid upper jaw, azhdarchid lower jaw, or thalassodromine upper jaw (as the sister-group of Alanqa). Here, we compare it in detail to upper and lower jaws of taxa belonging...
Preprint
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A recent publication of fossil bones of titanosaurs assigned to Aeolosaurini from the Morro do Cambambe site (Mato Grosso state, Brazil, Upper Cretaceous) reported anomalous growth in some of them. Here, we present osteohistological sections of elements to understand not only the microstructure and growth of such bones, but also the nature of those...
Article
Full-text available
Despite being known from every continent, the geological record of pterosaurs, the first group of vertebrates to develop powered flight, is very uneven, with only a few deposits accounting for the vast majority of specimens and almost half of the taxonomic diversity. Among the regions that stand out for the greatest gaps of knowledge regarding thes...
Article
Full-text available
Ornithocheirus wiedenrothi, from the Hauterivian (Early Cretaceous of Germany), is a taxon represented by three-dimensional remains of the lower jaw and wing elements. Its phylogenetic affinities have for long been elusive, though several works had already pointed out that it probably did not belong within the wastebasket genus Ornithocheirus. In t...
Article
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The first pterosaur bone bed from Brazil was reported in 2014 at the outskirts of the town Cruzeiro do Oeste, Paraná State, in the Southern region of the country. Here named 'cemitério dos pterossauros' site, these outcrops were referred to the Goio-Erê Formation (Turonian-Campanian) of the Caiuá Group (Bauru Basin) and revealed the presence of hun...
Article
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The pterosaur record from the Iberian Peninsula is mostly scarce and undefined, but in the last few years some new taxa have been described from different Lower Cretaceous sites of Spain. Here we describe a new genus and species of toothed pterodactyloid pterosaur from the Barremian of the Iberian Peninsula, Iberodactylus andreui gen. et sp. nov.,...
Article
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PurposeThe Bauru Group is worldwide known due to its high diversity of archosaurs, especially that of Crocodyliformes. Recently, it has been suggested that the Crocodyliformes, especially the Baurusuchidae, were the top predators of the Bauru Group, based on their anatomical convergence with theropods and the dearth of those last ones in the fossil...
Article
Full-text available
Purpose: New sauropod remains (MCF-PVPH-889, MCF-PVPH-899, and MCF-PVPH-900) collected from the Plottier Fm (Coniacian–Santonian) in the south-west of the Neuquén Basin, are here reported. The materials proceed from a fluvial outcrop composed by siltstone and fine sandstone, whose fossil record is known for large-sized sauropod taxa. Methods: Due t...
Article
Maximum sizes attained by living actinopterygians are much smaller than those reached by chondrichthyans. Several factors, including the high metabolic requirements of bony fishes, have been proposed as possible body‐size constraints but no empirical approaches exist. Remarkably, fossil evidence has rarely been considered despite some extinct actin...
Conference Paper
Estimates on the body mass and wingspan of Quetzal-coatlus northropi have sparked some controversy, some researchers concluding that it weighted over 500 kg and lacked power to fly. More recent estimates have agreed that the largest individuals (11 m wingspan) weighed up to 250 kg. Mathematical modelling of available metabolic power (Pav) and mecha...
Conference Paper
Full-text available
The ability of insects to produce damage in bones has been preserved on fossil bones as various kinds of traces such as tracks, burrows and borings, which are due to feeding, reproduction and/or shelter behaviour (Roberts et al., 2007). In this contribution, we report some evidences of bioerosion by insects on dinosaur bones. The two specimens stud...
Article
Full-text available
A small accumulation of bones from the Norian (Upper Triassic) of the Seazza Brook Valley (Carnic Prealps, Northern Italy) was originally (1989) identified as a gastric pellet made of pterosaur skeletal elements. The specimen has been reported in literature as one of the very few cases of gastric ejecta containing pterosaur bones since then. Howeve...
Chapter
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The present paper carries out a bibliographical review of the direct and indirect (ichnites) fossil remains of Iberian pterosaurs by compiling the fossil sites and kind of remains described since the mid 20th century to the present. Thus, we try to give an overview of current knowledge on this important group of archosaurs in the Iberian Peninsula,...

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