Axel Schönhofer

Johannes Gutenberg-Universität Mainz | JGU · Faculty of Biology

The scientific life of Prof. Dr Jochen Martens (Germany: Mainz) is illuminated on occasion of his 80th birthday. Facts and impressions are given as well as lists of his publications (329), taxa he has described (2 families, 29 genera, 296 species) and that have been dedicated to him (11 genera, 219 species, 1 subspecies). Jochen Martens is a renown...

Weberknecht-Papst, Himalaya-Forscher, Vogelstimmen-Experte, Professor, Mentor, Kollege, Freund... Es gibt viele Beschreibungen für Jochen Martens. Er wird am 10. Juni 2021 80 Jahre alt. Eine gute Gelegenheit, seine Person, seine Leistungen und sein Wirken zu würdigen. Aber wie? Wir drei Schüler, allesamt jünger und unerfahrener als er, wollen es hi...

Biogeographic studies often underline the role of glacial dynamism during Pleistocene (1.806–0.011 Mya) in shaping the distribution of subterranean species. Accordingly, it is presumed that present-day distribution of most specialized cold-adapted (cryophilic) cave-dwelling species should bear the signatures of past climatic events. To test this id...

By the possession of unique exocrine scent glands, Opiliones (harvestmen) arise as a perfect model for studies on the evolutionary history of secretion chemistry. Among gland compounds of harvestmen, it is the quinones that represent recurring elements across the secretions of all suborders. Reliable data on quinone-distribution, however, is only k...

We present new faunistic records of mites and harvestmen from the Maltese Archipelago and reviewed available data on the faunistics of the class Arachnida of the Archipelago. Literature records of Arachnids are rather scarce and uncomprehensive and up to date, checklists dealing with them have not been published except for spiders and gall mites. A...

The updated checklist contains 52 taxa. Four species have been newly recorded since 2004, one species had to be removed. The red data list is prepared according to the present requirements of the German Federal Agency for Nature Conservation (BfN). For each taxon the updated frequency in Germany is classified into one of six criteria classes, from...

With the nomination of the ‘Cave Animal of the Year’ the Society of German Cave and Karst Explorers calls public and authorities’ attention to the understudied biodiversity of subterranean ecosystems. Here the Cave Animal of the Year 2016, Amilenus aurantiacus (Simon, 1881), is presented. It is the first time that a harvestman has been chosen. Its...

Within the central European opilionid fauna the widely used species names Leiobunum rupestre Herbst, 1799 and Leiobunum tisciae Avram, 1968 pose taxonomic and distributional problems. In addition, Nelima apenninica Martens, 1969 is close to L. tisciae in terms of external and genital morphology, but is specifically distinct. While coxal denticulati...

The rigid leg segments of arthropods are flexibly connected by joints, which usually consist of two ball-and-bowl hinges, permitting a uniaxial pivoting up to 140°. Here, we report the occurrence of hyperflexible joints (range of movements = 160–200°) in the pedipalps (second pair of appendages) of some harvestmen (Sabaconidae and Nemastomatidae),...

Background Urban green spaces can harbor a considerable species richness of plants and animals. A few studies on single species groups indicate important habitat functions of cemeteries, but this land use type is clearly understudied compared to parks. Such data are important as they (i) illustrate habitat functions of a specific, but ubiquitous ur...

Location of sampled cemetery sections (grey) and sampled plots (small squares).

Pedipalps are the most versatile appendages of arachnids. They can be equipped with spines (Amblypygi), chelae (Scorpiones), or adhesive pads (Solifugae), all of which are modifications to grasp and handle fast-moving prey. Harvestmen (Opiliones) show a high diversity of pedipalpal morphologies. Some are obviously related to prey capture, like the...

Large prosomal scent glands constitute a major synapomorphic character of the arachnid order Opiliones. These glands produce a variety of chemicals very specific to opilionid taxa of different taxonomic levels, and thus represent a model system to investigate the evolutionary traits in exocrine secretion chemistry across a phylogenetically old grou...

Large prosomal scent glands constitute a major synapomorphic character of the arachnid order Opiliones. These glands produce a variety of chemicals very specific to opilionid taxa of different taxonomic levels, and thus represent a model system to investigate the evolutionary traits in exocrine secretion chemistry across a phylogenetically old grou...

We estimated a multigenic molecular phylogeny and reconstructed biogeographic history for the European harvestman genus Ischyropsalis C.L. Koch 1839 (Dyspnoi). To reconstruct historical biogeographic patterns we conducted an algorithmic VIP analysis which revealed patterns consistent with a vicariance-dominated history. The existing morphology-base...

Gluing can be a highly efficient mechanism of prey capture, as it should require less complex sensory-muscular feedback. Whereas it is well known in insects, this mechanism is much less studied in arachnids, except spiders. Soil-dwelling harvestmen (Opiliones, Nemastomatidae) bear drumstick-like glandular hairs (clavate setae) at their pedipalps, w...

The phylogeny of the spider family Sparassidae is comprehensively investigated for the first time using four molecular markers (mitochondrial COI and 16S; nuclear H3 and 28S). Sparassidae was recovered as monophyletic and as most basal group within the RTA-clade. The higher-level clade Dionycha was not but monophyly of RTA-clade was supported. No a...

An update of the systematics and determination key of the Opiliones suborder Dyspnoi is provided. The included catalogue represents the first comprehensive species and synonymy listing since Roewer (1923). It summarises all taxonomic changes to date and attempts to be a sound basis against the exponential growing number of online errors, for which...

Confirmation of Homalenotus quadridentatus (Opiliones: Sclerosomatidae) for the fauna of Germany. The sclerosomatid harvestman Homalenotus quadridentatus (Cuvier, 1795), long known close to the border of Germany, is hereby confirmed for the country. Several specimens were observed and collected in a small urban garden area in the town Aachen, sugge...

Within the species-rich European harvestman genus Trogulus Latreille, 1802, the Balkan Trogulus torosus species-group as defined by Schönhofer and Martens is revised. The group is remarkable because it includes the world’s largest Opiliones species, Trogulus torosus Simon, 1885, and Trogulus ozimeci sp. nov. is the first member of the family showin...

Bei der Inventarisierung subterraner Lebensräume in Luxemburg wurden im Zeitraum 2007 bis 2009 370 Weberknechte aus 15 Arten erfasst. Ein Großteil der Arten (73%) wird als eutrogloxen gewertet; sie wurden zufällig im Eingangsbereich der Höhlen gefunden. Echte Höhlentiere (Eutroglobionten) fehlen nördlich der Alpen. Zwei Arten wurden regelmäßig in d...

We investigated the relative phylogenetic position of the spider genera Psechrus Thorell, 1878 and Fecenia Simon, 1887 comprising the family Psechridae Simon, 1890 within the order Araneae (plus 50 outgroup taxa) using molecular data of the nuclear 28S rRNA gene and the mitochondrial cytochrome c oxidase subunit I (COI) gene. We further revised the...

We investigated the phylogeny and biogeographic history of the Holarctic harvestmen genus Sabacon, which shows an intercontinental disjunct distribution and is presumed to be a relatively old taxon. Molecular phylogenetic relationships of Sabacon were estimated using multiple gene regions and Bayesian inference for a comprehensive Sabacon sample. M...

Next-generation sequencing technologies are rapidly transforming molecular systematic studies of non-model animal taxa. The arachnid order Opiliones (commonly known as "harvestmen") includes more than 6,400 described species placed into four well-supported lineages (suborders). Fossil plus molecular clock evidence indicates that these lineages were...

Individual gene matrices. (DOCX)

Results of analyses of unpartitioned nucleotide matrices with third position sites removed. (TIF)

Gene ontologies. (DOCX)

Results from gene ontology analysis. (DOCX)

Results of alternative clock analyses. (DOCX)

Summary information for individual gene matrices (e.g., gene identities, nucleotide alignment lengths, PI values, average pairwise divergence values, taxon composition, etc.). (XLSX)

Next-generation sequencing technologies are rapidly transforming molecular systematic studies of non-model animal taxa. The arachnid order Opiliones (commonly known as “harvestmen”) includes more than 6,400 described species placed into four well-supported lineages (suborders). Fossil plus molecular clock evidence indicates that these lineages were...

In Opiliones, male genital morphology plays a key role in systematics and understanding the evolution of the group because functional modifications often outline higher taxonomic levels. In Nemastomatidae, different lateral stabilizing structures on the penial truncus cane developed independently. These wing-like protuberances serve as anchor point...

Trogulus banaticus Avram, 1971 is characterised and recorded as new for Slovenia. This species was previously mistaken for T. coriziformis C. L. Koch, 1839 and T. graecus Dahl, 1903 which were later rejected from the Slovenian fauna. T. banaticus is compared with the similar, and partly sympatric, T. tingiformis C. L. Koch, 1847 with which it has o...

Ischyropsalis dentipalpis Canestrini, 1872 is re-defined, and a neotype from the type locality in the Aosta Valley, Italy, is assigned. I. helvetica Roewer, 1916 (sensu Martens 1978) is shown to be conspecific and therefore is synonymised with I. dentipalpis. A population from Bergamo Province, Italy, formerly assigned to I. entipalpis (sensu Marte...

The development of winter sport resorts above the timberline may affect every ecosystem component. We analyzed the effect of ski-pistes on the abundance and species richness of arthropods (namely carabids, spiders, opilionids, and grasshoppers) trapped in grasslands adjacent to the ski-run, on ski-pistes and at the edge between these two habitat ty...

This is the first comprehensive study to evaluate the relationships between the western palearctic harvestman families Dicranolasmatidae, Trogulidae and Nemastomatidae with focus on the phylogeny and systematics of Trogulidae, using combined sequence data of the nuclear 28S rRNA and the mitochondrial cytochrome b gene. Bayesian analysis and Maximum...

A molecular genetic analysis of the cytochrome b gene (cyt b) revealed a monophyletic group of six widely separated but closely related haplotype clusters (4.8– 21.4% distance values) among minute representatives of Trogulus Latreille, 1802 from the Balkan Peninsula. All representatives have in common a large penial stylus that is sickle-like and s...

The harvestman fauna of the Western Soutpansberg was investigated during two collecting trips. Comments are made on the variability, ecology and remarkable morphological features of the collected material. Characters are displayed and differences to type series are highlighted. Monomontia neglecta sp. n. is described from material housed at the Nat...

Leiobunum religiosum (Arachnida: Opiliones): first record for Germany. L. religiosum Simon, 1879 was found at Mayen near Koblenz (Rhineland-Palatinate), Germany. This is the first record for Germany and is about 500 km from the known distribution area of the species in the southwestern Alps. The German population seems to be restricted to the ancie...

Within the well researched European fauna of harvestmen, the genus Trogulus Latreille exhibits unexpectedly high cryptic diversity. The species’ uniform morphology hinders an exclusively morphological approach to their systematics and taxonomy, and a preliminary molecular study estimated the number of species to be three times higher than currently...

Since about the year 2000 a hitherto unidentified species of the genus Leiobunum C. L. Koch, 1839, has rapidly invaded central and western Europe. Records are known from The Netherlands (probably the country of first occurrence in Europe), Germany, Austria and Switzerland. This introduced species, until now, mainly inhabits walls of buildings and r...

N. bidentatum Roewer, 1914 was found at two places in Germany: first on the island “Harriersand” in the Weser river (Lower Saxony), second on the banks of the river Elbe in the Elbsandsteingebirge (Saxony). Adjacent to the latter locality an occurrence in the Czech Republic could be located close to the German/Czech border in the floodplain of the...

From the beginning of November 2003 to January 2004 the opilionid fauna was collected in a floodplain forest of the Rhine valley west of Mainz. Altogether 1643 individuals were captured in pitfall traps; these belong to six species representing three families. The thermophilous species Astrobunus laevipes, Nemastoma dentigerum and Trogulus martensi...

Central and North European harvestmen from the Senckenberg Museum Frankfurt (Arachnida: Opiliones). About 1100 individuals belonging to 32 opilionid species (300 series), mainly of German origin, were identified based on material provided by, and stored at, the Senckenberg Museum, Frankfurt (Main). A list of all species with habitat information is...