Attila Lajos Borhidi

University of Pécs | PTE · Department of Systematic and Ecological Botany and Botanical Garden

The last synthetic treatment of the Rogiera genus was published by D. Lorence in the Flora MesoAmericana (Vol. 4, part 2) based on a polyphyletic concept including two genera of different tribes and leaving the results of the molecular studies out of consideration or misinterpreting them (Lorence 2012). Rovaeanthus has been distinguished at first b...

A new genus was described based on Portlandia guatemalensis Standl. of uncertain taxonomic position. The proposal of D. H. Lorence to place it into the genus Coutaportla Urb. as C. guatemalensis (Standl.) Lorence has not been supported by the discovery of a new species of Coutaportla (C. pailensis) nor by the molecular taxonomic studies of the grou...

The last synthetic treatment of the Arachnothryx genus was published by D. Lorence in the Flora Mesoamericana (Vol. 4, part 2, 2012) based on a polyphyletic concept including several genera of different tribes and leaving the results of the molecular studies out of consideration. The analytic key offered by the author is based firstly on the highly...

The new genus is based on Palicourea locellata C. M. Taylor described from the Colombian Andes and placed into the Nonatelia section reconsidered and strongly enlarged by C. M. Taylor including 26 species into the originally monotypic section. According the new protologue of the section it is a highly varied and rather heterogeneous group in morpho...

Morphological and molecular studies show that many species classified in Psychotria L. subgen. Heteropsychotria Steyerm. belong to other genera, as Margaritopsis C. Wright, Carapichea Aubl., Notopleura (Oerst.) Bremekamp, and to Palicourea Aubl. (Andersson 2001, 2002a, b, Nepokroeff 1997, Nepokroeff et al. 1999, Razafimandimbison et al. 2014, Taylo...

The subgenus Heteropsychotria was detected and described by Julian Steyermark in the frame of the monographic series of the Botany of the Guyana Highland (Steyermark 1972) and repeatedly in a more illustrated version in the Flora of Venezuela (Steyermark 1974). Within the subgenus he distinguished 13 sections and 31 series. Three sections (Nrs 4, 7...

The authors carried out a taxonomic study and revised and identified about 7,000 herbarium sheets of the genera Palicourea and Psychotria preserved in the collections of the Academy of Sciences of Cuba (HAC) and of the National Botanical Garden of Cuba (HAJB) with the purpose to complete the manuscript in preparation for the volume of the Rubiaceae...

Spontaneous dynamics of unmanaged fl oodplain oak and submontaneous Illyrian beech forests were compared in the Pannonian ecoregion (southern Hungary), with special respect to changes of abundance of species in the ground layer. Due to gap forming, light availability, estimated by Ellenberg indicator values, improved signifi - cantly in oak stands,...

Spontaneous dynamics of unmanaged floodplain oak and submontaneous Illyrian beech forests were compared in the Pannonian ecoregion (southern Hungary), with special respect to changes of abundance of species in the ground layer. Due to gap forming, light availability, estimated by Ellenberg indicator values, improved significantly in oak stands, in...

A new pyrene type is reported, it is morphologically different from that found in the genera Palicourea and Psychotria. This new type is characterised by being triangulate in transverse section with an elevated central-dorsal crest. It was observed in two species of Palicourea with dissimilar floral characteristics. Based on this finding, we propos...

The reorganisation and new circumscription of the gigantic genus Psychotria started with the classic article of Nepokroeff and co-authors in 1999, based on molecular phylogenetic studies. Its statements and suggestions were followed by the separations and modifications of different genera, as the extension of Margaritopsis by Andersson in 2001 and...

The validation of two Bouvardia (Rubiaceae) species of Mexico is carried out recommended by Govaerts because the placing of their type materials was not included into the proto-logues. New names are proposed for a Mexican Bouvardia and a neotropical Uragoga (both Rubiaceae) species having illegitimate names. A correction of author's name is propose...

The new Arachnothryx species collected in Veracruz is a vicarious one of the next related Arachnothryx secundiflora (B. L. Rob.) Borhidi of Oaxaca, Tabasco and Chiapas, which differs in having sessile bractless cymes. Another closely related species A. linguiloba Borhidi et Diego Pérez of Guerrero differs in size and pubescence of stipules and leav...

The new Guettarda species of Oaxaca (Mexico) is related to G. foliacea Standl. of a distribution from Tabasco and Chiapas to Panama, but differs by having long acuminate and falcate leaves with lineolate quaternary nerves on the abaxial surface, inflorescences with very short ovate and glabrous bracteoles and much shorter flowers with 8–9 mm long c...

The Randia sepium is an extremely interesting species with its coriaceous medium-sized leaves glabrous and brilliant on both sides with 4–7 pairs of secondary nerves and terminal inflorescences, with medium-sized hirsute flowers and pubescent corolla lobes on both sides not having closely related taxon in the Mexican and Mesoamerican flora. All Ran...

The new Psychotria species belongs to the panamensis-mexiae complex with its large long deciduous ferrugineous terminal stipules, but differs from them by having shorter inflo-rescence, 2–3 times larger flowers, smaller leaves and less numerous secondary nerves. It is of allopatric distribution being endemic to Jalisco state, while the area of P. m...

The new species of Mitracarpus of Oaxaca y Tabasco is prominent of its extremely small flowers almost completely hidden among the longer calyx lobes. It differs from Mitracarpus breviflorus A. Gray, the other small-flowered species of the genus, by being a perennial herb with 4-angular hairy stems lignescent at the base and the larger, elliptical,...

The Randia sepium is an extremely interesting species with its coriaceous medium-sized leaves glabrous and brilliant on both sides with 4-7 pairs of secondary nerves and terminal inflorescences, with medium-sized hirsute flowers and pubescent corolla lobes on both sides not having closely related taxon in the Mexican and Mesoamerican flora. All Ran...

The new Psychotria species belongs to the panamensis-mexiae complex with its large long deciduous ferrugineous terminal stipules, but differs from them by having shorter inflorescence, 2-3 times larger flowers, smaller leaves and less numerous secondary nerves. It is of allopatric distribution being endemic to Jalisco state, while the area of P. me...

The new Arachnothryx species collected in Veracruz is a vicarious one of the next related Arachnothryx secundiflora (B. L. Rob.) Borhidi of Oaxaca, Tabasco and Chiapas, which differs in having sessile bractless cymes. Another closely related species A. linguiloba Borhidi et Diego Pérez of Guerrero differs in size and pubescence of stipules and leav...

A new species of Chomelia is described; at this time it has been collected only at the type locality. It is similar to the Mexican-Mesoamerican species Chomelia longituba (Borhidi) Borhidi, but can be easily distinguished by the position of the inflorescence, by the form of the stigma lobes and the size of the fruit.

The new Guettarda species of Oaxaca (Mexico) is related to G. foliacea Standl. of a distribution from Tabasco and Chiapas to Panama, but differs by having long acuminate and falcate leaves with lineolate quaternary nerves on the abaxial surface, inflorescences with very short ovate and glabrous bracteoles and much shorter flowers with 8-9 mm long c...

The new Arachnothryx species collected in Veracruz is a vicarious one of the next related Arachnothryx secundiflora (B. L. Rob.) Borhidi of Oaxaca, Tabasco and Chiapas, which differs in having sessile bractless cymes. Another closely related species A. linguiloba Borhidi et Diego Pérez of Guerrero differs in size and pubescence of stipules and leav...

A new species of Guettarda is described from a collection made by José C. Soto Nuñez in Guerrero. It is similar to G. guerrerensis Borhidi of Guerrero and G. quadrifida Borhidi et Reyes-García of Chiapas, and grows in tropical deciduous forests on igneous rocks.

A new species named as Psychotria zapotecana (Spermacoceae tribe) was collected in the State Oaxaca (South Mexico) in the framework of the collecting program of SERBO. It is related to P. mirandae C. W. Hamilt. of Chiapas Veracruz and Guatemala, but differs from it by having shorter chartaceous leaves, green when dry with abaxially domatiate and hi...

A new species named as Randia keniae (Gardenieae tribe) was collected in the State Oaxaca (Mexico), by Kenia Velasco Gutierrez in the framework of the collecting program of SERBO. It is related to R. hypoleuca Borhidi et E. Martínez of Michoacan and Oaxaca, but differs from it by having sessile to subsessile obovate leaves, with rounded apex and cu...

A new species of Randia is described from a collection made by José C. Soto Nuñez in Michoacán. It is similar to R. laevigatoides Borhidi spreading from Guerrero to Oaxaca and Chiapas, but differs from it in having bisexual flowers, leaves with obtuse to rounded base, calyx with 2 to 5 mm long linear to linear-obovate lobes, shorter corolla with gr...

A new endemic species of the genus Arachnothryx was collected in the State Jalisco, (Mexico Central) related to A. jurgensenii (Hemsl.) Borhidi. With the new taxon the genus is represented by 60 species, 44 of them endemic to the Flora of Mexico.

A new species named as Borreria myrtilloides (Spermacoceae tribe) was collected in the State Oaxaca (South Mexico) in the framework of the collecting program of SERBO. It is related to B. nesiotica B. L. Rob. of Baja California and the Socorro Island, but differs from all similar species by having perennial to subfrutescent habit, very small shiny...

Three new species belonging to the Bouvardia genus are recognised and described by the authors of the paper. B. lancifolia Borhidi et Salas-Morales of Oaxaca from the relationship of B. rosea Schltdl. of Central Mexico, the B. sinaloae Borhidi et E. Martinez Salas of Sinaloa, related to the B. sancaroli Borhidi et M. Martinez-Diaz of Tamaulipas and...

Three new species belonging to the genus of Randia are recognised and described by the authors of the paper. The R. colimensis is an endemic of the Volcano Colima and surroundings , vicarious of R. xalapensis Mart. et Gal. largely distributed in the Sierra Madre Oriental from Tamaulipas and San Luis Potosí to Tabasco and Chiapas. R. dionisi and R....

A new species of Randia is described from a collection made by José C. Soto Nuñez in Michoacán. It is similar to R. laevigatoides Borhidi spreading from Guerrero to Oaxaca and Chiapas, but differs from it in having bisexual flowers, leaves with obtuse to rounded base, calyx with 2 to 5 mm long linear to linear-obovate lobes, shorter corolla with gr...

A new species of the genus Bouvardia belonging to the subgen. Bouvardiastrum, sect. Gymnosiphon was collected by José Pascual in the southern part of the state Oaxaca in Mexico. At first glance it is similar to the polymorphic B. multiflora (Cav.) Schult. et Schult. f. widely distributed in Mexico to El Salvador, but differs from it basically in ha...

In the collections accumulated within the framework of the Flora of Guerrero Project appeared a remarkable Psychotria taxon belonging to the large and rather polymorphic species of P. horizontalis Sw. but differs from both known subspecies of it, which are the type and the subsp. glaucescens (Kunth) Borhidi, with discolourous leaves and completely...

Two new Rogiera species have been detected in the collections of the Andayu group of Oaxaca (R. oaxacensis) and of the State University of Tabasco (R. tabascensis). Both species are notable because of their pending, creeping, liana-like habit growing on calcareous rocks, which is a rare geological formation in the mountainous systems of México form...

Two new species of the genus Didymaea from Oaxaca and a new species of the genus Mitracarpus from Campeche — both belonging to the tribe Spermacoceae — has been detected and are described by the authors. D. ixtepejiensis is a dense, creeping herb of a high-montane pine woodland, with 1-nerved leaves. The next related species is D. microphylla L. O....

For the second and enlarged edition of the Rubiaceas de Mexico (2012) several new combinations, 5 to Edithea (E. guerrerensis, E. oaxacana, E. rupicola, E. serboi and E. sousae all transferred from Deppea) 2 to Houstonia (H. mcvaughii, and H. acerosa subsp. tamaulipana, both from Hedyotis) one Palicourea (P. buchtienii transferred from Psychotria)...

Within the frame of the Deppea two types of floral organisation have been recognised: the classic genus based on the type D. erythrorhiza Cham. et Schltdl., characterised by rotate corolla with very short tube, longer filaments and long linear anthers both longer than the corolla tube. The other floral type is characterised by funnel-form corolla,...

Two new Randia species are described from the aculeata-complex, characterised by small leaves, flowers and fruits, the later with soft exocarpium. This group apparently has its diversification centre in Mexico represented actually by 19 species and 3 varieties. Randia crucis is a deciduous tree up to 6mhigh in Oaxaca, characterised by terminal pair...

Terrellianthus is a monotypic genus of Mexico and Guatemala, created for finding a correct position for the enigmatic Hedyotis serpyllacea Schltdl. described in 1835, and discussed at great length up to now.

Terrellianthus is a monotypic genus of Mexico and Guatemala, created for finding a correct position for the enigmatic Hedyotis serpyllacea Schltdl. described in 1835, and discussed at great length up to now.

A new species of the genus Deppea was collected in the zone of Chimalapas (Oaxaca), area especially rich in rare and endemic species. It is named as D. chimalaparum and belongs to the section Deppea and to the group of small-flowered species with unequal calyx lobes. Within this group it differs from D. obtusiflora (Benth.) Benth. by having deciduo...

Based on molecular and morphological studies the circumscription and reshaping of the genera Palicourea and Psychotria have been proposed (Andersson and Rova 1999, Nepokroeff et al. 1999 and several papers of C. M. Taylor). The most important change is the new placement of the Psychotria subgen. Heteropsychotria Steyerm. (1972) in the Palicourea ge...

Two new species were found in the inexhaustible treasury of the collections made by Erik Leonard Ekman in Hispaniola during the 1920s. One is Varronia gonavensis (Boraginaceae) from the Gonave Island of Haiti, the other one is a new Guettarda (G. vulpina) from the Samana Peninsula of Santo Domingo. Both are endemics to the Hispaniolan flora.

Three new species of the genus Deppea s. str. are discovered and described collected in the federal states Puebla, Veracruz and Chiapas, respectively. Deppea amaranthoides of Veracruz is closely related to the amaranthina-nitida group, but differs in having chartaceous leaves and much larger flowers. D. hoffmannioides of Puebla can be distinguished...

Chiococca motleyana is a new name to replace the later homonym Chiococca pubescens arising after the transfer of Asemnantha pubescens Hook. f. into the genus Chiococca against the earlier hom-onyms described by Humboldt and Bonpland ex Roemer and Schultes 1819, and by Standley in 1919. Asemnantha was separated from Chiococca by Hooker f. in 1873, b...

A new Rondeletia species (Rondeletieae, Rubiaceae) was collected by the combined German- Cuban research group of the National Botanical Garden of Cuba in the dry southern region of the historical province Oriente, actually Prov. Guantanamo, in the valley of Rio Seco between Imias and Cajobabo. The region is rich in small leaved sclerophyllous speci...

Donnellyanthus is a new genus for the correct accommodation of the enigmatic species Bouvardia deamii Donnell-Smith, ranging from Mexico (Oaxaca) to Honduras and Salvador. Its own special character combination differs from Bouvardia in having imbricate corolla lobes and entire stipules. Therefore it was transferred to Rondeletia by Standley, but it...

The paper revises and re-discusses the literature dealing with the phytosociology of the Spiraea rock heath communities based on relevés collected in North Hungary (Northern Hungarian Mountain Range) and South Hungary (Mecsek and Villány Mts). The comparative analyses confirmed the separation of the formerly described two associations and added one...

A new species and a new variety of the genus Hoffmannia Sw. were collected in the rich flora of Oaxaca. Hoffmannia silviarum is related to H. psychotriifolia (Benth. in Oerst.) Griseb. and dedicated to the honour of two significant persons in the exploration and understanding of the Rubiaceae flora of Mexico.

Two new Bouvardia species are described in this paper belonging to the subgenus Bouvardioides W. H. Blackwell, a large-flowered one from the Federal State of México, with short sessile leaves names as B. amplexicaulis, the other is a xeromophic but unarmed one originated from the dry areas of Oaxaca, closely related to B. erecta, with 4-angled shin...

The existence of an earlier homonym of the generic name Neomartensia Yoshida et Mikami in the family Delesseriaceae of the red algae made necessary to give a new name: Martensianthus for six Rubiaceae species of the Mexican flora to replace the later homonym Neomartensia Borhidi et Lozada-Pérez 2010.

Three new spiny species of the genus Randia (Gardenieae, Rubiaceae) were collected in the dry areas of the states Oaxaca and Michoacán, named R. hypoleuca, in Puebla (R. pueblensis) and in Veracruz (R. veracruzana) living in dry thickets and dry deciduous forests. Randia alvarocamposii is a new name for replacing the older homonym R. horrida 2007....

Six new species of Bouvardia Salisb. have been discovered from the states Oaxaca and Guerrero. All are belonging to the subgenus Bouvardiastrum W. H. Blackwell, actually represented by 30 species in the flora of Mexico. Ten of them have corolla tube naked in the inside a character uncommon in the genus. Based on this important feature, two new sect...