
Ángel ViloriaInstituto Venezolano de Investigaciones Científicas | IVIC · Centro de Ecologia
Ángel Viloria
Doctor of Philosophy
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252
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Introduction
Publications
Publications (252)
A new species of Lepidoptera (Papilionoidea) is described on the basis of specimens collected at upper elevations in the Guiana Shield: Thaeides hyperion Bálint, Costa & Grishin, n. sp. (Lycaenidae: Theclinae: Eumaeini). Due to its adaptation to mountainous areas, it is probably a taxon endemic to the biogeographic Pantepui Province. With 16 figure...
Obituario. Nota sobre la vida y obra de la bióloga y ornitóloga uruguayo-venezolana, Virginia Sanz D’Angelo (1964-2024).
Defective taxonomic descriptions and the electronic publication fashion. A comment on Braby et al. (2024) and a rectification Descripciones taxonómicas defectuosas y la moda de la publicación electrónica. Comentario sobre Braby et al. (2024) y una rectificación ABSTRACT Some reflections are presented on the global practice of publishing scientific...
Noticias sobre el estudio de la coleccion Macleay de mariposas en el Chau Chak Wing Museum de la Universidad de Sydney, Australia, y la localizacion de material tipo de especies descritas por Dru Drury, Johann C. Fabricius y Edward Donovan.
Noticia sobre el retorno a Venezuela de material tipo de especies fosiles de peces devonicos de la Sierra de...
Ensayo sobre la vida y obra de Alfred Russel Wallace (1823-1913) en conmemoracion del centenario de su nacimiento.
The Llanos of Colombia and Venezuela are an ecoregion composed of savannas, forests, and wetlands, with a high biodiversity and once home to a high-density jaguar (Panthera onca) population. We used published and new jaguar presence–absence data from 2001 to 2020 and combined logistic regression with kriging interpolation to model jaguar occurrence...
El espacio insular venezolano comprende gran parte del espacio marítimo del país, denominado mar territorial (Mar Caribe y Atlántico); su presencia expande la frontera política venezolana con Estados Unidos de América, Francia, Países Bajos, República Dominicana y Trinidad & Tobago. Agrupa las islas mayores de Margarita, Coche y Cubagua (estado Nue...
A new subspecies of Lepidoptera (Papilionoidea) is described from the Pantepui: Heliconius elevatus jigginsi Costa & Neild, n. ssp. (Nymphalidae, Heliconiinae, Heliconiini) from Auyán Tepui (Venezuela), probably strictly endemic to the Auyán Massif. A new synonymy is proposed: Heliconius elevatus sonjae Neukirchen, 1997, n. syn. of Heliconius eleva...
Zegara fernandezi (González, 1992) (Castniidae, Castniinae) is a little-known butterfly-moth due to its rarity. New evidence provides information about its habitat, linked to the summits of some table mountains of the Pantepui; several specimens collected to date are illustrated showing remarkable polymorphism, as well as its southern Venezuelan di...
The diverse, largely Neotropical subtribe Euptychiina is widely regarded as one of the most taxonomically challenging groups among all butterflies. Over the last two decades, morphological and molecular studies have revealed widespread paraphyly and polyphyly among genera, and a comprehensive, robust phylogenetic hypothesis is needed to build a fir...
Report of the rescue operation of a male manatee (Trichechus manatus) in La Blanquilla island in the Caribbean Sea (Venezuela). This captive bred individual had been released with a satellite tracker in the coast of Ceará, Brazil, on July 6, 2022, and was recovered in Venezuela, September 5, 2022, after swimming 4,000+ km.
Two new satyrine butterflies known so far only from the Venezuelan territory are diagnosed, described, illustrated and compared: Rudyphthimoides iseai gen. nov. et sp. nov. (Cordillera de la Costa, northern Venezuela) and Stephenympha pauliana gen. nov. et sp. nov. (Serranía del Cuao, Amazonas, southern Venezuela). Seven new combinations, two new s...
Oxeoschistus puerta sabinoi ssp. nov., a pronophiline butterfly (Lepidoptera: Nymphalidae: Satyrinae) from the montane and upper montane forests (1,100-1,900 m a.s.l.) of the Venezuelan slope of the Sierra de Perijá, Zulia state, Venezuela, is described, illustrated and compared with its close congeners. The taxon O. puerta magnus Pyrcz & Viloria,...
We revised the Odonata collection of the Museo del Instituto de Zoología Agrícola (MIZA), Facultad de Agronomía, Universidad Central de Venezuela (UCV), creating a database for nine selected genera and 35 species. The data were submitted to a Parsimony Analysis of Endemicity (PAE) using the WINCLADA/NONA program. Our results show a total of 25 area...
Two new species of the diving beetle genus Liodessus Guignot, 1939 are described from high mountain regions in Venezuela: Liodessus meridensis sp. nov. from Laguna de Mucabají, Mérida and L. venezuelensis sp. nov. from Laguna de Mucabají and below Pico Bolívar, Mérida. We delineate the species using morphological characters such as male genital str...
A new subspecies of Lepidoptera (Papilionoidea) is described from the Pantepui: Pterourus menatius karljordani Neild, Costa, Jost & Grishin, n. ssp. (Papilioninae, Papilionini) from the Gran sabana (Bolivar state, south-eastern Venezuela).
Book review
Contenido: Resumen y Abstract; Presentación: un tesoro para compartir y cuidar (por José Manuel Restrepo Abon-dano); Introducción (por Patricia Londoño Vega); 1. ¿Qué es y cómo se desarrolla una mariposa? (por Nicolás Peláez Restrepo); 2. ¿Por qué en Colombia hay tantas y tan varia-das mariposas?; Riqueza de Colombia en mariposas: pers-pectiva comp...
Investigations on the nomenclature of the American butterfly genus Hermeuptychia Forster, 1964 (Lepidoptera: Nymphalidae, Satyrinae) yielded the reinstatement of three species, the proposal of four new combinations, and the recognition of eight new synonymies; mainly through the study and scrutiny of information published in scientific papers, book...
Some personal reminiscences of British Paleontologist Brian G. Gardiner
Book Review
Book Review
Se describe una nueva especie y dos nuevas subespecies de Lepidoptera (Papilionoidea, Riodinidae, Pieridae y Nymphalidae) del Pantepui: Emesis malik Callaghan, Costa, Trujano-Ortega & Benmesbah, n. sp. (Riodininae, Emesidini), Lieinix christa bousquetsi Costa, Neild, Grishin & Viloria, n. ssp. (Dismorphiinae, Dismorphiini) y Euptychia marceli divis...
All animals, including carnivores, adapt their daily activity duration and distribution to satisfy food demands, breed, or avoid mortality risk. We used the kernel density method to estimate daily movement activity levels and movement activity patterns of jaguars in Hato Piñero, in Venezuelan Western Llanos, based on 3,656 jaguar detection time rec...
The Andean Region presents a great variety of natural landscapes in which there
are mountainous areas with a high richness of butterflies of the Pronophilina subtribe. The
diversity and structure of the community of pronophiline butterflies (Nymphalidae, Satyrinae)
was determined in this study in an altitudinal gradient of the Cerro de Tierra Negra...
To characterize the diversity, structure and composition of the assemblage of diurnal butterfly species (Lepidoptera: Papilionoidea), standardized captures were made for twelve months (June 2012-May 2013) in two sites of a cloud forest landscape in the Cordillera de la Costa, northern Venezuela (Altos de Pipe, 1,450 and 1,650 m asl, Los Salias Muni...
The identity, status and taxonomic position of seven species of Neotropical satyrine butterflies are discussed and defined: Euptychia modesta Butler, 1867, Euptychia nebulosa Butler, 1867, Neonympha alcinoe C. Felder & R. Felder, 1867, Eup¬tychia mimas Godman, 1905, Euptychia mimula Hayward, 1954, Euptychia pamela Hayward, 1957 and Cissia gomezi Si...
Se describen cuatro nuevas subespecies y una nueva especie de Lepidoptera (Papilionoidea) del Pantepui: Pseudopieris viridula autana Costa & Viloria, n. ssp., Dismorphia zathoe hernandi Costa & Neild, n. ssp., Dismorphia zathoe blanca Costa & Attal, n. ssp. y Dismorphia crisia eburnea Benmesbah & Costa, n. ssp. (dismorphiinae, dismorphiini); Pyrrho...
Plutella xylostella (Linnaeus, 1758) has been claimed to be the most widely distributed Lepidoptera species. However, it is a highly polymorphic species, and there is evidence that what is known as P. xylostella is a species group, so it is possible that all records do not correspond to this species. Aim of the work was to evaluate the variability...
We present the observations and results of the life cycle of Eretris catargyrea (Staudinger, 1887), stat. rev., a nymphalid butterfly of the subfamily Satyrinae, endemic to the Colombian Andes. This study was carried out in the Municipality of Choachí, Cundinamarca, Colombia, at 2,300 m asl. The life cycle of this species was complete in 74 days fr...
Lepidoptera from the Pantepui. Part IX. New Nymphalidae (Satyrinae) and Riodinidae (Riodininae two new species and a subspecies of Lepidoptera (Papilionoidea) are described from the Pantepui: Opsiphanes tiberius attal & Blandin, n. sp. (satyrinae, Brassolini), Euptychia woroina Viloria & Benmesbah, n. sp. (satyrinae, satyrini) and Symmachia virgaur...
The watercolors of the species of the subfamily Satyrinae (Nymphalidae, Lepidoptera) illustrating Théophile Raymond’s “Butterflies of Venezuela” (1982) are identified. Comments are made on the author, his work and the editions of Raymond’s book, as well as some of the taxa mentioned, including the proposal of one change of status.
Brief details on the biography of the German Biologist, Naturalist, Cytogeneticist Michael Schmid is presented here in. Includes some details on his connection with Venezuela, a country where he grew up and returned to do research. Comments on the origin and relevance of his first published book "Mariposas de Venezuela," which is the first Venezuel...
Se hace una revisión comentada de los contenidos del libro The ecology and evolution of Heliconius butterflies, por Chris D. Jiggins (2017, Oxford University Press)
Two new subspecies and two species of Lepidoptera (Papilionoidea) are described from the Venezuelan portion of the Pantepui: Memphis paulus clarae Attal & Costa, ssp. nov. (Charaxinae, Anaeini), Stalachtis halloweeni miranda Costa & Fratello, ssp. nov. (Riodininae, Stalachtini), Forsterinaria chaniorum Viloria & Costa, sp. nov. and Euptychia sarah...
Abstract. Lamprospilus ayawi sp. nov. (Lycaenidae, Theclinae), whose male possesses distinctive transparent wing patches,
is described from the Guiana Highlands of South America, on the basis of six males and five females from Venezuela (Auyán
and Sororopán Tepuis) and one male from Guyana (Acarai Mts.).
Resumen. Lamprospilus ayawi sp. nov. (Lycae...
The genus Llorenteana, gen. nov., is diagnosed and described to allocate the enigmatic, relictual, satyrine butterfly, Euptychia pellonia Godman, 1901. The taxonomy of this species had been superficial, variable and unstable for more than a century. The new genus is monobasic and endemic to the montane pine-oak forests of the Northern Mexican Altip...
Four genera of butterflies belonging to the Nymphalidae: Satyrinae are erected, diagnosized, described and discussed in or-
der to reassign ten species recently classified within the genus Magneuptychia Forster, several of which occur in Colombia and Venezuela: Nubila Viloria, Andrade & Henao, gen. nov. (to include Euptychia gera Hewitson, E. metag...
Four genera of butterflies belonging to the Nymphalidae: Satyrinae are erected, diagnosized, described and discussed in or-
der to reassign ten species recently classified within the genus Magneuptychia Forster, several of which occur in Colombia and Venezuela: Nubila Viloria, Andrade & Henao, gen. nov. (to include Euptychia gera Hewitson, E. metag...
A basic description of the mountain landscape in the Guiana Shield leads to the primordial concept of Pantepui as a biogeographic entity. A historical account of the discovery and study of the butterflies in the montane zones of Venezuela, Brazil and Guyana within this region (tepuis) introduces a checklist of its endemic butterfly taxa and data on...
Resumen. – Se describen tres nuevas especies y una subespecie de Lepidoptera Papilionoidea del Pantepui venezolano: Megaleas cervelina Orellana & Costa, n. sp. (Hesperiidae, Hesperiinae), Adelpha irmina arcana Benmesbah & Costa, n. ssp. (Nymphalidae, Limenitidinae), Pedaliodes parakana Viloria & Costa, n. sp., y Eretris cuaoensis Viloria & Attal, n...
Perfil biográfico de Indiana Cristóbal Ríos-Málaver (Manizales, 1983-), biólogo y ecólogo colombiano, especialista en mariposas neotropicales, coautor del libro Colombia, País de mariposas (2018).
Little is known about the diversity and biology of Calliphoridae in northern South America, especially in Venezuela. In particular, the effects of deforestation/ urbanisation (and their associated synanthropic processes) on Calliphor idae species remains poorly known. In this context, the goal was to evaluate the relationship between Calliphoridae...
Abstract. Two new species of butterfly (Lepidoptera, Papilionoidea, Lycaenidae) from the Pantepui (Venezuela) are described; Ocaria faurei Bálint, Attal & Costa, sp. nov. and Strymon auyana Bálint, Benmesbah & Viloria, sp. nov. (Lycaenidae: Theclinae: Eumaeini). Notes and data are provided on the representatives of the genera Ocaria Clench, 1970 an...
The complete life cycle of Oressinoma typhla Doubleday, [1849] and the morphology of its immature stages are described. Biological and ecological information is provided for each stage, and its host plant is identified as Cyperus involucratus (Cyperaceae) in the mountains of northern Venezuela.
Editorial El basilisco, Harold Molero y la región carbonífera del Guasare
Book review
Broad scale population estimates of declining species are desired for conservation efforts. However, for many secretive species including large carnivores, such estimates are often difficult. Based on published density estimates obtained through camera trapping , presence/absence data, and globally available predictive variables derived from satell...
Pearson correlation matrix for jaguar density estimates (dependent variable) and a set of predictive variables.
(DOCX)
Spatial distribution of the posterior standard deviations of adjusted jaguar density estimates.
(DOCX)
Estimates of jaguar populations within protected areas by current range countries.
(DOCX)
Spatial distribution of the posterior standard deviations of estimated probabilities of jaguar occurrence.
(DOCX)
Density estimates and other relevant data used for the analysis of spatial variation in jaguar population densities.
(DOCX)