Àngel Hernández Luján

Àngel Hernández Luján
Institut Català de Paleontologia Miquel Crusafont | ICP · Area Neogene and Quaternary Faunas

PhD

About

69
Publications
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277
Citations

Publications

Publications (69)
Article
Archaeological investigations carried out in the cave Zubbio di Cozzo San Pietro, Bagheria, Sicily, revealed the presence of a few skeletal elements of a large-sized tortoise in a funerary area dating to the Copper/Bronze Age. The tortoise has been AMS-dated revealing an age of 12.5 ± 0.5 kyr BP and therefore it pre-dates the funerary activities. T...
Conference Paper
Full-text available
The late Oligocene genus Proailurus is generally accepted as the earliest member of the family Felidae. Later on, during the Miocene four different felid species are traditionally recognized and included in the paraphyletic genus Pseudaelurus: P. quadridentatus, P. romieviensis, P. lorteti, and P. turnauensis. More recently various authors have ass...
Article
Els Casots is one of the richest fossil vertebrate sites of the Vallès-Penedès Basin (Catalonia, Spain). It was discovered in 1989 and excavated briefly during the 1990s, resulting in the recovery of thousands of remains and the erection of several new mammal species. Excavations were resumed in 2018 and continue to date. Here we provide updated re...
Article
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The holotype of the Tenerife (Canary Islands) giant rat, Canariomys bravoi, an almost complete cranium from the Late Pleistocene site of Cueva de las Palomas, is described for the first time. This species is characterized by its large size, robust skull with a short rostrum, dorsal inflation at the level of the infraorbital foramen, and moderately...
Article
This article focuses on the sciurid (Rodentia, Sciuridae) remains from the early Miocene (MN4) Mokrá-Quarry sites, Moravia, Czech Republic. The sciurid assemblage in the different fissures ranges from one genus in MWQ4/2018 to three different genera in MCQ3/2005. Regarding subfamily Sciurinae, the dentognathic remains are referred to Palaeosciurus...
Article
Fossil turtles from Mokrá-Quarry, South Moravia Region, Czech Republic, are described in this paper. Remains come from two already known karstic fissures uncovered in Mokrá-Western Quarry (1/2001 Turtle Joint and 2/2003 Reptile Joint), as well as three new karstic fissures from Mokrá-Western Quarry (TC/2001 and 4/2018) and Mokrá-Central Quarry (3/2...
Article
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Deinotheres (Proboscidea, Deinotheriidae) are a clade of non-elephantiform proboscideans that originated in Africa and dispersed into Eurasia by the early Miocene. In Europe, deinotheres are first recorded in Greece during MN3, although they did not become a common faunal element throughout Europe until MN4. Early Miocene (MN3–MN4) deinothere remai...
Conference Paper
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Several almost complete shells, skulls and postcranial bones of trionychid turtles from the Most Basin (NW Bohemia, MN3 mammal zone) have never been published or revised. Here we present the reappraisal of the trionychid turtles from the Most Basin (Czech Republic) together with the revision of Rafetus bohemicus from Břešťany Clay, including new di...
Article
Czujan's sandpit is an abandoned quarry in the Vienna Basin (Mikulov, Czech Republic) that has yielded an important middle Miocene vertebrate assemblage. Here we re-describe the site from the perspective of sedimentology, taphonomy, and paleoenvironments, and further review the biochronology of the fauna to clarify the age. The updated faunal list...
Article
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Fossil crocodylians from the early Miocene (Eggenburgian, MN3a) sites of Ahníkov (Most Basin, Czech Republic) are described in this paper. The new material presented here includes over 200 remains (bones, teeth and osteoderms), and therefore constitutes the largest crocodylian sample known from the fossil record of the Czech Republic. Assignment of...
Article
Three species of Deinotherium sensu stricto (Proboscidea, Deinotheriidae), i.e., excluding Prodeinotherium, generally considered to have nonoverlapping chronostratigraphic distributions, are currently recognized from the Miocene of Europe: Deinotherium levius (late Astaracian/Aragonian, MN7+8), Deinotherium giganteum (type species; Vallesian, MN9–M...
Article
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Two fossiliferous karstic fissures from the Mokrá-Western Quarry, MWQ (1/2001 Turtle Joint; 2/2003 Reptile Joint) provided a diverse vertebrate fauna from the early Miocene (Burdigalian, MN 4) including squamates. The rather warm climatic conditions during the Miocene Thermal Maximum (17.8-17.7 Ma) enabled dispersal of thermophilic lizards and snak...
Article
The dispersal of Crocodylus from Africa to Europe during the Miocene is not well understood. A small collection of cranial fragments and postcranial elements from the latest Miocene (6.2 Ma) site of Venta del Moro (Valencia, Spain) have previously been referred to Crocodylus cf. C. checchiai Maccagno, 1947 without accompanying descriptions. Here we...
Conference Paper
Full-text available
First alligatoroid Diplocynodon remains from the Ahníkov/Merkur Mine site (A/MM) are described here. A/MM is the richest fossil site of the Most Basin (Eger Graben, Czech Republic), including more than 150 taxa of fossil vertebrates reported to date. Specifically, more than 300 remains of fossil crocodilians have been collected, but they have never...
Article
Full-text available
Phalangeal reduction is a common and widespread phenomenon among tortoises that has been associated with the adaptation to terrestrial life. While reduced manual digit 1 appears characteristic in almost all Testudo species, it is uncertain why the metacarpal I and distal carpal of the same digit are completely missing in some individuals of Hermann...
Article
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Revision on historical finding of the giant turtle from the Brno Sand (middle Miocene, lower Badenian) The collections of Moravian Museum in Brno contain for decade rare find of fossil giant terrestrial tortoise of the genus Titanochelon. There was no more information about the finding with the only exception for the name of the locality "Královo P...
Article
Full-text available
The genus Diplocynodon was a fossil alligatoroid that inhabited Europe from the Paleocene until the Miocene. During the Eocene–Oligocene transition, circa 35 million years ago, a terrestrial faunal turnover occurred in Eurasia during the Grande Coupure. This overall climate shift is linked with the extinction of numerous European Eocene endemic mam...
Article
The early Miocene deposits of Bohemia (Czech Republic) contain numerous fragmentary crocodylian remains. Despite this abundance, a detailed taxonomical assignment of these remains was impossible due to the absence of diagnostic cranial elements. Here, we report two partially preserved skulls together with some osteoderms and a partially preserved v...
Article
Worm lizards, or amphisbaenians, of the genus Blanus are found in various countries around the Mediterranean Sea. In addition to four extinct species, seven extant taxa are currently recognized. Here, we present the first comparative analysis of the cranial osteology of Blanus including all extant species. The results of this analysis show a homoge...
Article
Full-text available
Monitor lizards (genus Varanus) inhabited Europe at least from the early Miocene to the Pleistocene. Their fossil record is limited to about 40 localities that have provided mostly isolated vertebrae. Due to the poor diagnostic value of these fossils, it was recently claimed that all the European species described prior to the 21st century are not...
Data
List of references consulted to score characters for the phylogenetic analysis. (PDF)
Data
Matrix used in the phylogenetic analyses. (NEX)
Data
Lists of the apomorphies of V. marathonensis, of the synapomorphies of the clade including V. marathonensis and the subgenus Indovaranus, and of Varanus (based on analysis 1B). (PDF)
Data
List of extant comparative specimens of Varanus. (PDF)
Data
New characters added to the original list of Conrad et al. [11]. (PDF)
Data
Results of the phylogenetic analysis 1A. (PDF)
Data
Results of the phylogenetic analysis 2A. (PDF)
Poster
Full-text available
The late early Miocene snake assemblages are well-documented from Central European localities of the MN 4 and MN 5 Zones. However, diversified snake fauna is still poorly known from MN 1 to MN 3 Zones. German Wintershof-West locality (MN3b) provided both abundant fauna with a relatively diversified snake assemblage including Boidae: Bavarioboa sp.,...
Conference Paper
Full-text available
Diplocynodon ratelii is a fossil alligatoroid known from the Miocene of Europe. It is characterized by the presence of a crest-like thickening on the margin of the external naris; the anterior tip of the nasals reach externally (but do not bisect) the posterior rim of the external naris; a concavoconvex frontopari-etal suture; a shorter mandibular...
Article
In the Iberian Peninsula, Miocene apes (Hominoidea) are generally rare and mostly restricted to the Vallès-Penedès Basin. Here we report a new hominoid maxillary fragment with M2 from this basin. It was surface-collected in March 2017 from the site of Can Pallars i Llobateres (CPL, Sant Quirze del Vallès), where fossil apes had not been previously...
Book
Full-text available
¿Qué sabemos realmente sobre la historia de la vida en la Tierra? En general nos parece que la paleontología se dedica únicamente al estudio de los dinosaurios, o eso nos ha hecho creer el cine, pero esta ciencia ha conseguido explicarnos con un detalle sorprendente el origen de la vida y su evolución a lo largo de millones de años. Porque no todo...
Article
Full-text available
El conocimiento sobre las tortugas gigantes fósiles de Europa se ha incrementado considerablemente en los últimos años y ha comportado la descripción del género Titanochelon para incluir a las especies del Neógeno europeo. Sin embargo, la inclusión de la tortuga gigante de las Islas Baleares, “Testudo” gymnesica, en este género se ha llevado a cabo...
Poster
Full-text available
La variación del número de dedos y falanges de las extremidades anteriores es un fenómeno común en tortugas, aunque la variación en el grado de desarrollo difiere en género e incluso a nivel de especie. Se ha estudiado la variación en el número de dedos y falanges de las extremidades de la población de tortuga mediterránea (Testudo hermanni hermann...
Article
The single extant species of the anuran genus Latonia lives in Israel, but in the fossil record the genus is known mainly from Europe, spanning from the Oligocene to the early Pleistocene. Here we describe new remains of Latonia from the early to late Miocene of the Vallès-Penedès Basin (NE Iberian Peninsula), coming from the following localities:...
Poster
Full-text available
Czujan’s sandpit (Mikulov, Czech Republic) represents a unique locality of middle Miocene mammals, and further is known as the type locality of Tethytragus stehlini. Czujan’s sandpit is mentioned only occasionally in published sources, however, Neogene mammals from the Mikulov area come from several sites of different stratigraphical age. Therefore...
Conference Paper
Full-text available
The Miocene record of the Vallès-Penedès Basin (NE of the Iberian Peninsula) has provided a rich and diverse herpetofauna (amphibians and reptiles), which is known in the literature since the early 20th Century, thanks to the pioneering work of Bergounioux and, subsequently, Crusafont and Hoffstetter. However, the early Miocene localities have rece...
Conference Paper
Full-text available
The two fossiliferous karst localities of Mokrá-Western Quarry, Czech Republic (MWQ, 1/2001 Turtle Joint and MWQ, 2/2003 Reptile Joint) have provided a rich herpetofauna (amphibians and squamates) from the Early Miocene (MN4b). Here, we describe the unpublished turtle material recovered from above-mentioned sites, as well as two other localities of...
Conference Paper
Full-text available
The knowledge about the European extinct alligatoroids significantly increased during the last ten years, based on the erection of the two new species and the revision of the previously published taxa. The remains of the genus Diplocynodon are quite common in the northwest localities of Bohemian area, but they have only been referred at genus level...
Article
We describe crocodylian remains from the Early Miocene (MN4) site of Els Casots (Subirats, Vallès-Penedès Basin, NE of the Iberian Peninsula). Referral to Diplocynodon (Alligatoroidea: Diplocynodontidae) is justified by several cranial and postcranial features, including: (1) the subequal and confluent alveoli of the maxilla (fourth and fifth) and...
Article
Full-text available
We provide a taxonomic review of the extinct testudinid Testudo catalaunica, based on published and unpublished material from several Miocene (late Aragonian and early Vallesian) sites of the Vallès-Penedès Basin (north-east Iberian Peninsula). We show that Testudo catalaunica irregularis is a junior subjective synonym of T. catalaunica, and furthe...
Conference Paper
Full-text available
We report the crocodylian remains from the early Miocene (MN4) site of Els Casots (Subirats, Vallès-Penedès Basin, NE Iberian Peninsula). They are attributed to the extinct alligatoroid Diplocynodon, which includes nine valid species from the Tertiary of Europe (Delfino and Smith, 2012; Martin et al., 2014). Among available remains (skulls, teeth,...
Article
Full-text available
Background. The Late Cretaceous is a keystone period to understand the origin and early radiation of Crocodylia, the group containing all extant lineages of crocodilians. Among the taxa described from the latest Cretaceous of Europe, the genus Allodaposuchus is one of the most common but also one of the most controversial. However, because of its f...
Conference Paper
Full-text available
The chelonian fauna of Sicily is currently limited to a single autochthonous tortoise, Testudo hermanni hermanni, whose shell length reaches approximately 20 cm. Even if the Balkan subspecies of the Hermann's tortoise, Testudo hermanni boettgeri, is significantly larger (up to 35 cm), the Sicilian Pleistocene fossil record testifies for the presenc...
Conference Paper
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Nel maggio 2014, durante lo scavo archeologico di una sepoltura multipla all'interno della grotta Zubbio di Cozzo San Pietro (PA), attribuito, secondo la cultura materiale, al Bronzo Antico (metà III mill. a.C.), sono stati rinvenuti reperti faunistici di grandi e piccoli vertebrati. Particolarmente interessanti sono i resti di una testuggine di di...
Article
Full-text available
The taxonomy of the freshwater turtle subgenus Ptychogaster (Temnoclemmys) (Testudines,Geoemydidae, Ptychogasterinae) is revised based on new and previously-published mate-rial from the Vallès-Penedès Basin (NE Iberian Peninsula). Additional remains of subgenusPtychogaster, first reported from this basin, are also described. Although the oldest rem...
Article
Full-text available
Although skulls of extinct Testudinidae are generally much scarcer than shell remains, when available they provide important data for resolving taxonomic and phylogenetic problems, as illustrated here by two well-preserved giant tortoise skulls from the early Vallesian (MN9, Late Miocene) of Ecoparc de Can Mata (ECM; els Hostalets de Pierola, Vallè...
Chapter
Full-text available
The history of turtle lineage began around 225 Mya. Since then, this group of reptiles developed diverse ecological strategies and colonized a wide range of environments, from marine to fully terrestrial habitats. In spite of its their great ecological diversity, the bauplan of turtles is peculiar and little variable, with a body encased in a rigid...
Article
Abstract. Testudo lunellensis Almera and Bofill, 1903 from the Middle Pleistocene of Cova de Gràcia (Park Güell, Barcelona, Spain) is a valid species belonging to the clade of the extant Testudo hermanni – a diagnostic feature being the narrowed vertebral scutes. Thanks to still unpublished material, T. lunellensis is diagnosed for the first time a...
Article
Cheirogaster includes medium to large-sized extinct terrestrial tortoises from Europe. Here we report two skulls from the Vallesian (MN9, Late Miocene) of Ecoparc de Can Mata (ECM; els Hostalets de Pierola, Catalonia, Spain), attributed to Cheirogaster richardi, whose cranial anatomy was previously unknown. This taxon was considered a subjective ju...
Article
Full-text available
Monitor lizards have a wide current distribution embracing Africa, Asia, and Australia, but the fossil record indicates that the genus Varanus inhabited Europe at least from the Early Miocene to the Pliocene. The European fossil record is limited to about 35 localities that have provided mostly isolated vertebrae. The following European species hav...
Article
Full-text available
Se dan a conocer los resultados preliminares de la intervención paleontológica preventiva realizada en el marco de los trabajos de construcción del Ecoparc de Can Mata (=Ecoparc 4, ECM; Vallesiense inferior, MN9), en el término municipal de els Hostalets de Pierola (cuenca del Vallès-Penedès). La intervención se realizó desde 2007 a 2010, y fue mot...
Article
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Two news skulls of the giant tortoise Cheirogaster (Testudines: Testudinidae) from the early Vallesian (MN 9, late Mioce-ne) of Ecoparc de Can Mata (els Hostalets de Pierola, Vallès-Penedès Basin, Catalonia, Spain) are preliminarily described. The cranial features of the specimens confi rm an attribution to the genus Cheirogaster and suggest that t...
Article
Full-text available
In the Iouaridène Syncline (High Atlas, Morocco) 28 sites have been studied, with 781 dinosaur footprints listed. The theropod tracks are the most abundant in the area. The sites are located in the red layers of the Iouaridène Formation of Oxfordian-Kimmeridgian age (Upper Jurassic). Two trackways and a pair of theropod footprints have been found i...

Projects

Projects (4)
Project
This project focusses on the study of the fossil remains from the Late Miocene site of Venta del Moro, Valencia Spain.
Project
This project explores the presence of marmot’s remains and their meaning in a paleontological and archaeological context. The present study has been focused on three objectives which are: 1. To find out the taxon’s number of Marmota genus that is present in Europe during Quaternary and which is the cause of the marmots’ diachronic distribution during this period; 2. To check the alpine marmot’s presence as a cold fauna paleoenvironmental indicator; 3. To find out if the marmots’ rests concentrations in the deposits are as a result of the human anthropic activity