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Publications (219)
A study of Rovno amber has revealed two fossil leafy liverwort species of the genera Lejeunea and Odontoschisma (Marchantiophyta), which are new to the late Eocene flora of Rovno amber. One of these species, Odontoschisma dimorpha, is known from Baltic and Bitterfeld amber, whereas the other species, Lejeunea aristovii, is described here as new. Le...
Schäfer-Verwimp, A. & Winter, G. (2024): Preliminary checklist for the liverworts and hornworts of El Salvador. Frahmia 41:1-17 * A brief summary of the geology, physical features, climate and vegetation is given, followed by the checklist and a compilation of 13 further (previously unpublished) records based on herbarium specimens from various her...
Winter, G. & Schäfer-Verwimp, A. (2024): A small bryophyte collection made by Wilhelm Lötschert 1952-1953 in El Salvador. Frahmia 40:1-15 *. A small collection of bryophytes made by W. Lötschert in El Salvador in 1952-1953 and kept in the Herbarium Senckenbergianum (hb FR) was reexamined. In 1955, O. Burck had examined only the mosses and described...
Two leafy liverwort species from the late Eocene Rovno amber are described. They are placed in the extant genus Nipponolejeunea (Jubulaceae, Porellales, Marchantiophyta) that includes two extant East Asian species and one fossil species described from the Eocene Baltic amber. New fossil species N. rovnoi sp. nov. and N. solodovnikovii sp. nov. diff...
Premise
Bryophytes form a major component of terrestrial plant biomass, structuring ecological communities in all biomes. Our understanding of the evolutionary history of hornworts, liverworts and mosses has been significantly reshaped by inferences from molecular data, which have highlighted extensive homoplasy in various traits and repeated burst...
Schäfer-Verwimp, A. & Winter, G. (2023): Frullania hohenesteri (Marchantiophyta: Frullaniaceae)-a new species from Santa Catarina, South Brazil. Frahmia 39:1-14 *. Frullania hohenesteri is described and illustrated from Santa Catarina, South Brazil. It belongs to the subgenus Trachycolea sect. Irregulares because of its peculiar flattened and smoot...
Winter, G. & Schäfer-Verwimp, A. (2023): Frullania dulimensis (Marchantiophyta: Frullaniaceae) from Costa Rica with perianths. Frahmia 36:1-6 3. The rare neotropical liverwort Frullania dulimensis has been detected for the first time with perianths in Costa Rica, representing the first record for Central America. So far, it was known from the type...
New records of 18 species of mosses and liverworts are presented, with detailed notes on their taxonomy, distribution, and ecological characteristics. These findings provide valuable insights for advancing research in biogeography, ecology, and conservation.
The specimen previously identified as Marsupella microphylla from Brazil is reassessed and described as a new species, M. brasiliensis . The new species is characterized by paroicous inflorescence, bispiral elaters, scale-like, commonly unlobed leaves and very small leaf cells. Descriptions and drawings are provided along with a corresponding discu...
Bryophytes are emerging as the sister-group to extant vascular plants, and their current diversity highlights that their life cycle characterized by a dominating vegetative gametophyte and an unbranched sporophyte composes a successful alternative to that of vascular plants and their dominating sporophyte. The evolutionary history of hornworts, liv...
Bazzania species that look similar to B. subtilis which were collected by the first and third author in the Philippines are described and illustrated. Amongst these specimens B. acanonista and B. globuliformis were described as new to sience. Furthermore B. herzogiana, B. sikkimensis, B. vittata, and B. wiltensii, mentioned by other authors for the...
Schäfer-Verwimp, A. & Winter, G. (2022): Frullania azureomontana, a new species from Jamaica (Marchantiophyta: Frullaniaceae). Frahmia 30:1-12. * Frullania azureomontana is described and illustrated from elfin forest of the Blue Mountains of Jamaica. It stands out by distant lobules (subgen. Diastaloba s.l.), relatively small size, and dispersed oc...
Amber is renowned for the exceptional preservation state of its inclusions, allowing detailed morphological analysis and providing relevant environmental, palaeoecological, geographical, and geological information. Amber deposits predominantly known from North America, Europe, and Asia, are considered to be rare on the continents that formed Gondwa...
Prior to the advent of molecular work, the observable variation in vegetative reproduction has been used to classify Lejeunea into subgenera and sections. Thereby, the ability of developing caducous leaves was regarded as major factor. A reexamination of several Lejeunea specimens revealed that L. cocoes with caducous leaves shows considerable morp...
Marsupella sect. Stolonicaulon is not speciose and is a commonly neglected section within the genus, which currently includes three species with somewhat similar morphologies (wiry shoots with distanced leaves) and distributions in the mountains of tropical and subtropical regions (SE (Southeast) Asia, the Venezuelan Andes, and the high mountains o...
Extending the diversity of the bryoflora in Kachin amber (Myanmar), with the description of Radula patrickmuelleri, sp. nov. and R. tanaiensis, sp. nov. (Jungermanniopsida, Porellales, Radulaceae)
Abstract
The most prolific source of exquisitely preserved bryophyte fossils is amber, which often contains inclusions in a three-dimensional and lifeli...
Schäfer-Verwimp, A., Winter, G., Yao, K.-Y., Yang, J.-D. (2022): Plagiochila subtropica (Plagiochilaceae, Marchantiophyta) new to Taiwan. Frahmia 28:1-5. Plagiochila subtropica is reported for the first time for the bryophyte flora of Taiwan. A morphological description and figures based on Taiwanese plants are provided.
Introduction
Half a century since the creation of the International Association of Bryologists, we carried out a review to identify outstanding challenges and future perspectives in bryology. Specifically, we have identified 50 fundamental questions that are critical in advancing the discipline.
Methods
We have adapted a deep-rooted methodology of...
New and remarkable bryophyte records in the Province of South Tyrol, Italy. II.
Three species of bryophytes (Plagiothecium neckeroideum, Pohlia camptotrachela and Pohlia sphagnicola) are reported new to Italy, six are first records for the region of Trentino-Alto Adige/Südtirol (Cephaloziella stellulifera, Scapania verrucosa, Bryoerythrophyllum fer...
DNA-based analyses as well as the fossil record indicate that liverworts date back to the early Paleozoic, but diverse specimens of lifelike three-dimensional fossils only occur as amber inclusions from the mid-Cretaceous onwards. Some Albian–Cenomanian ambers preserve elements of a diverse biota that existed during the rise of angiosperms, a time...
New records of 23 species of mosses and liverworts are presented, with detailed notes on their taxonomy, distribution, and ecological characteristics. These findings provide valuable insights for advancing research in biogeography, ecology, and conservation.
Anisothecium rotundatum Broth. (= Dicranella rotundata [Broth.] Takaki) is newly combined under Diobelonella, and the latter genus is reported from Taiwan for the first time. Description of the plants from Taiwan along with photo plates and line drawings of the stem and leaf cross sections of the genera Diobelonella and Dichodontium are provided. D...
Among liverworts, the epiphytic lifestyle is not only present in leafy forms but also in thalloid liverworts, which so far has received little attention in evolutionary and biogeographical studies. Metzgeria, with about 107 species worldwide, is the only genus of thalloid liverworts that comprises true epiphytes. In the present study, we provide th...
The number of available liverwort fossils substantially increased within the past decade, which is mainly due to new findings from Cretaceous and Cenozoic amber deposits. Many of them, however, are fragmentary and not predestined for consideration in evolutionary analyses. here, we list those liverwort fossils that we suggest as suitable for calibr...
ABSTRACT Porotrichum squarrosum has never been reported from Brazil and it is a new record. While revising herbarium specimens we found a specimen named Pireella squarrosa (currently Porotrichum), identified by Massimo Mastracci. We have compared this specimen with records from literature and several hundreds of specimens and found that among speci...
Frullania dasyueshanensis sp. nov. is described from Dasyueshan National Forest Recreation area and other localities in Taiwan. The new species was collected in mountain rainforests between 2230 and 2650 meter.
It differs from all known Frullania species in Taiwan by its clavate-cylindrical terete perianth and can be clearly distinguished from the...
Jungermannia obcordata LEHM. & LINDENB. was published 1834 by LEHMANN and transferred to Frullania in Synopsis Hepaticarum 1845. YUZAWA (1988) transferred 4 species as synonyms sub Frullania caulisequa (NEES) MONT., among them Frullania obcordata (LEHM. & LINDENB.) LEHM. & LINDENB. probably without having seen the type of Frullania caulisequa. The...
Understanding the biogeographical and diversification processes explaining current diversity patterns of subcosmopolitan-distributed groups is challenging. We aimed at disentangling the historical biogeography of the subcosmopolitan liverwort genus Lejeunea with estimation of ancestral areas of origin and testing if sexual system and palaeotemperat...
Seven species (Cephalozia ambigua, Harpanthus flotowianus, Isopaches decolorans, Schljakovianthus quadrilobus, Solenostoma pumilum, Andreaea nivalis and Dicranella staphylina) are reported for the first time for South Tyrol, three of them (Schljakovianthus quadrilobus, Solenostoma pumilum and Dicranella staphylina) as new for the region of Trentino...
Abstract
Seven species (Cephalozia ambigua, Harpanthus flotowianus, Isopaches decolorans, Schljakovianthus quadrilobus, Solenostoma pumilum, Andreaea nivalis and Dicranella staphylina) are reported for the first time for South Tyrol, three of them (Schljakovianthus quadrilobus, Solenostoma pumilum and Dicranella staphylina) as new for the region of...
During a recent visit of the Maltese Islands Malta and Gozo in January 2018, some 200 collections of bryophytes have been done representing one species of hornworts, 14 species of liverworts and 49 species of mosses, among these six first records (Riccia ciliata Hoffm., Targionia lorbeeriana Müll.Frib., Amblystegium serpens (Hedw.) Schimp., Cheilot...
From recent fieldwork in Taiwan some 11 bryophytes, including 8 liverworts (Cheilolejeunea krakakammae, Cephaloziella divaricata, Diplophyllum obtusatum, Douinia plicata, Geocalyx graveolens, Lejeunea micholitzii, Metzgeria crassipilis, Spruceanthus mamillilobulus), and 3 mosses (Daltonia marginata, Ulota intermedia and Zygodon reinwardtii) are rep...
Frullaniaceae are the most diverse family of leafy liverworts preserved in amber and are known from several deposits dating from the Miocene to the Early Cretaceous. In the fossil record, Frullania is represented by 15 species as well as the extinct genera Pseudofrullania, Protofrullania and Kaolakia. Here, we describe another species of Frullania...
New synonyms, morphological characterizations and national records for three neotropical Lejeunea species, including comments on distribution and ecology, are presented. Lejeunea cristulata (Steph.) M.E.Reiner & Goda (= L. resupinata [Steph.] Steph., syn. nov.) is newly recorded for Venezuela, Peru and Bolivia. Lejeunea diversicuspis Spruce (= L. m...
Molecular evidence supports the transfer of Conoscyphus Mitt. from Lophocoleaceae to Acrobolbaceae, which is unexpected on the basis of morphological evidence and further disrupts the morphological circumscription of Acrobolbaceae. Conoscyphus differs from other Acrobolbaceae in possessing a stem perigynium and a conspicuous perianth that forms a t...
On the presence of Mittenothamnium reduncum (Hypnaceae) in Argentina. Mittenothamnium
reduncum is recorded for the first time for the bryological flora of Argentina. A brief description, illustration,
and discussion of the species are provided. Morphological differences between this taxon and M. reptans,
the most frequent species in the area, are p...
An inclusion in Miocene Dominican amber is described as Cheilolejeunea lamyi, sp. nov. The fossil resembles the extant Cheilolejeunea rigidula but differs in the partly subacute leaf lobes, often apiculate underleaf lobes, and the slightly more acute lobes of the bracteole. The two other Cheilolejeunea species in Dominican amber, Cheilolejeunea ant...
A joint bryological expedition was conducted during mid-October 2016 between the authors and colleagues with the Taiwan Endemic Species Research Institute (TAIE). 1012 bryological collections were obtained. Species new for selected counties and for Taiwan were documented. A set of these collections are deposited at TAIE.
Fossil tree resins preserve a wide range of animals, plants, fungi and microorganisms in microscopic fidelity. Fossil organisms preserved in an individual piece of amber lived at the same time in Earth history and mostly even in the same habitat, but they were not necessarily parts of the same interacting community. Here, we report on an in situ pr...
The new species Lejeunea flagellifera from Brazil is described and illustrated. This species is characterized by its large size, the large, inflated and rectangular leaf lobule and by flagelliform branches with caducous leaves. The specimens were found in montane ombrophilous forests, growing on bark and living leaves of trees.
Classification of incompletely preserved fossils is a challenging task. We describe a moss fossil preserved in a piece of Miocene Dominican amber and assign it to the monospecific extant genus Helicophyllum. According to the published divergence time estimates, Helicophyllum originated in the Late Cretaceous, but so far no Helicophyllum fossils hav...
Lepidolejeunea grandiocellata, sp. nov. is described for two specimens from the Blue Mountains of Jamaica. This delicate species is characterized by ocelli in basal parts of leaves, underleaves, bracteoles and bracts often being conspicuously larger than the surrounding cells, whereas ocelli in central and upper parts of these structures are more o...
DNA-based divergence time estimates suggested major changes in the composition of epiphyte lineages of liverworts during the Cretaceous; however, evidence from the fossil record is scarce. We present the first Cretaceous fossil of the predominantly epiphytic leafy liverwort genus Radula in ca. 100 Myr old Burmese amber. The fossil's exquisite prese...
Molecular phylogeny reconstruction has motivated recircumscription of all families and most genera within the Lophocoleinae, and in Plagiochila, the largest genus of this lineage, has refined the sectional classification as well. Here, we extend this ongoing revision in the first study focusing on species from Australasia, a region to date underrep...
We present molecular data demonstrating that the New Zealand endemic Plagiochila caducifolia represents an isolated population of Plagiochila (sect. Arrectae) spinulosa, a species previously thought confined to the eastern Holarctic but which also occurs in Lesotho, southern Africa. In New Zealand, the wide distribution of P. spinulosa in undisturb...
An inclusion in Cretaceous amber from Myanmar is described as a new fossil species in the extant liverwort genus Frullania. The name Frullania pinnata is proposed for the taxon that is characterized by entire underleaves, paired with a conspicuous pinnate branching pattern, two distinctive and stable morphological features. Entire underleaves are k...
A revision of the Baltic and Bitterfeld amber fossils assigned to Cylindrocolea dimorpha (Cephaloziellaceae) has yielded evidence of the presence of multicellular, bifid underleaves, which have not previously been reported for this species and conflict with the current circumscription of the family. We transfer the fossil species to Odontoschisma (...
A chloroplast DNA phylogeny of Radula substantiates the presence of three Radula species in Austria, R. complanata, R. lindenbergiana and R. visianica, all members of subg. Radula. Radula visianica has been observed at a few localities in the Austrian Alps, and is resolved in a sister relationship with a clade comprising R. complanata, R. jonesii,...
Three new species of the genus Cheilolejeunea are described and illustrated: C. cyrtolejeuneoides sp. nov. and C. grosseoleosa sp. nov. both from Brazil, and C. azureomontana sp. nov. from Jamaica. Cheilolejeunea cyrtolejeuneoides is similar to C. holostipa and C. insecta but mainly differing by autoicous plants and deeply bifid underleaves; Cheilo...
A revision of the Baltic and Bitterfeld amber fossils assigned to Cylindrocolea dimorpha (Cephaloziellaceae) has yielded evidence of the presence of multicellular, bifid underleaves, which have not previously been reported for this species and conflict with the current circumscription of the family. We transfer the fossil species to Odontoschisma (...
Recent findings established the Albian-Cenomanian amber of Myanmar (Burmese amber) as an important source of Mesozoic cryptogams. Here, we describe an inclusion within Burmese amber as Protofrullania cornigera gen. et sp. nov. in the extant family Frullaniaceae (Porellales, Jungermanniopsida). Diagnostic characters include its incubous leaves consi...
Allopolyploidy is an important mode of speciation in angiosperms, ferns and mosses, but has been considered of little importance in liverworts. The few known examples include taxa of the simple thalloid genus Pellia, the complex thalloid genera Corsinia, Reboulia and Targionia and the leafy liverworts Barbilophozia, Calypogeia and Porella. The dipl...
The new species Lejeunea longidentata from Brazil is described and illustrated. This species is characterized by asymmetrically ovate-falcate leaf lobes, ventral lobe margin often upcurved, lobe apex often recurved and by the narrowly elongate, curved, banana-shaped lobule tooth-cell. The specimen was found in Araucaria forest in Minas Gerais growi...
Molecular and morphological data support the recognition of seven species of Chiastocaulon in Australia, of which four, namely C. braunianum and C. geminifolium comb. nov. from tropical north-eastern Queensland and C. proliferum and C. flamabilis sp. nov. from Tasmania (and New Zealand), represent new records. The other three species accepted for A...