Alexander V. Timokhov

Lomonosov Moscow State University | MSU · Department of Entomology

Forest canopy is densely populated by phyto-, sapro-, and microbiphages, as well as predators and parasitoids. Eventually, many of crown inhabitants fall down, forming so-called ‘arthropod rain’. Although arthropod rain can be an important food source for litter-dwelling predators and saprophages, its origin and composition remains unexplored. We m...

A morphological and molecular analysis of Gryon Haliday (Platygastroidea, Scelionidae) was conducted to provide a taxonomic and phylogenetic context for a species under evaluation as a biological control agent of Bagrada hilaris (Burmeister) (Hemiptera, Pentatomidae). Our analysis revealed that Gryon is polyphyletic and that the biological control...

Обобщены и уточнены известные сведения о распространении, биотопической приуроченности и фенологии видов рода Bittacus Latreille 1805 в европейской части России и прилегающих регионах. Bittacus italicus (Müller 1766) впервые указан для Волгоградской области и Казахстана. Проанализированы диагностические признаки B. italicus и B. hageni Brauer 1860....

The available data on geographic distribution, habitat preferences, and phenology of species of the genus Bittacus Latreille, 1805 in European Russia and the adjacent regions of Kazakhstan are summarized and clarified. Bittacus italicus (Müller, 1766) is recorded for the first time from Volgograd Province of Russia and from Kazakhstan. The potentia...

Invertebrate phyto-, sapro-, and microbophages, as well as predators and parasitoids, densely populate tree crowns. Eventually, all crown inhabitants fall from the trees and become a food source for litter-dwelling predators, scavengers, and saprophages. However, the functional significance of the arthropod rain, i.e., the flux of invertebrates fal...

Abstract. A brief review of the proctotrupoid genus Vanhornia Crawford is given. A new species, Vanhornia yurii sp. nov. from Primorskiy Territory, Russia, is described and illustrated. Photographic illustrations of Vanhornia eucnemidarum Crawford, 1909 (specimens from USA), V. leileri Hedqvist, 1976 (specimens from the Russian Far East) and V. qui...

Karyotypes of four species of the genus Telenomus Haliday, 1833, namely, T. acrobates Giard, 1895, T. angustatus (Thomson, 1861), T. heydeni Mayr, 1879 and T. turesis Walker, 1836 were studied for the first time using chromosome morphometrics. Although all examined members of the genus appeared to have n = 10, substantial differences between karyot...

The new records of 24 species of scelionid wasps in the regions of Russia and adjacent countries, Georgia, Kazakhstan, and Mongolia, are given. Of them, Apegus kerteszi Kieffer, 1908 is recorded for the fauna of Russia for the first time. A new generic combinations is suggested: Phanuromyia moldoviana (Özdikmen, 2011), comb. nov. Doubtful distribut...

The new data on distributions of nine species of platygastrid wasps in the fauna of Russia are given. Of them, Leptacis laodice (Walker, 1836) is recorded in the fauna of Russia for the first time. New species are recorded for Leningradskaya, Moscow, Chelyabinsk and Irkutsk Provinces, Stavropol, Altai and Primorskiy Territories, Chuvashia, Mordovia...

Karyotypes of three species of the genus Trissolcus Ashmead, 1893, namely, T. kozlovi Rjachovsky, 1975, T. rufiventris (Mayr, 1907) and T. semistriatus (Nees von Esenbeck, 1834), were studied for the first time. Although all examined members of the genus appeared to have 2n = 20, chromosome morphometrics revealed substantial differences between kar...

The Agromyzidae fly Selachops flavocinctus Wahlberg, 1844 as host of the Alysiinae genera Asyntactus Marshall, 1898 (Alysiini) and Protochorebus Perepechaenko, 1997 (Dacnusini) is recorded for the first time. A new species from Central Urals, Protochorebus pervushini sp. nov., is described and illustrated. The new material for Asyntactus rhogaleus...

FIGURES 10 – 20. Protochorebus pervushini sp. nov. (10 – 12, 14 – 20, female, holotype; 13, male, paratype). 10. Habitus, dorsal view. 11. Head, front view. 12, 13. Head, dorsal view. 14, Mandible. 15. Basal segments of antenna. 16. Apical segments of antenna. 17. Mesosoma, dorsal view. 18. Mesosoma, lateral view. 19. Hind femur and tibia. 20. Ovip...

FIGURES 21 – 23. Protochorebus pervushini sp. nov. (female, holotype). 21. Fore and hind wings. 22. Metasoma, dorsal view. 23. Metasoma, lateral view.

FIGURES 24 – 34. Protochorebus kasparyani Perepechaenko (female, holotype). 24. Mandible. 25. Habitus, dorsal view. 26. Head, front view. 27. Head, dorsal view. 28. Apical segments of antenna. 29. Basal segments of antenna. 30. Fore and hind wings. 31. Mesosoma, lateral view. 32. Mesosoma, dorsal view. 33. Metasoma, dorsal view. 34. Apex of metasom...

FIGURE 1. View of spring shore vegetation aspect on Verkh-Isetsk pond (Yekaterinburg City).

FIGURES 2 – 9. Asyntactus rhogaleus Marshall (female). 2. Habitus, dorsal view. 3. Habitus, lateral view. 4. Head, dorsal view. 5, Mandible. 6. Head and mesosoma, dorsal view. 7. Median segments of antenna. 8. Metasoma, dorsal view. 9. Apical part of metasoma and ovipositor, ventral view.

FIGURES 45. Map of distribution of Selachops flavocinctus Wahlberg.

FIGURES 35 – 44. Selachops flavocinctus Wahlberg. 35. Adult habitus, lateral view. 36 Wing. 37. Puparium within culm of Carex acuta. 38. Cephalopharyngeal skeleton (extracted from puparium), dorsal view. 39. Cephalopharyngeal skeleton, lateral view. 40. Puparium habitus, lateral view. 41. Puparium habitus, dorsal view. 42. Puparium, anterior end, l...

Table S1. Overview of measurements of Anisopteromalus females, showing minimum, maximum, mean and standard deviation in µm (except for A. ceylonensis with n = 1).

Table S3. Pearson's product-moment correlation coefficients for all measurements of Anisopteromalus.

Figure S1. Matrix scatterplot of eight variables (in µm) of Anisopteromalus.

Table S2. List of Anisopteromalus specimens and outgroups included in the molecular analysis.

Appendix S1. Descriptions of species and lists of further material examined.

We use an integrative taxonomic approach to revise the genus Anisopteromalus. In particular, we apply multivariate ratio analysis (MRA), a rather new statistical method based on principal component analysis (PCA) and linear discriminant analysis (LDA), to numerous body measurements and combine the data with those from our molecular analysis of Cytb...

Plant, spider, beetle, and ground beetle communities were studied in different agrolandscape biotopes (a wheat field, a field margin, and a forest belt) of Krasnodar Territory. In these biotopes, borders between plant communities were distinct. On the contrary, arthropod communities were not usually confined to the certain biotopes. The high degree...

The investigations (three years) were carried out in agroecosystems (maize, vetch-oat mixture, and winter wheat), including their adjoining grassy margins. Insects were collected regularly during growing periods by standard entomological sweeping in different zones of agrosystems: in the center (about 150 m from field margins), along perimeter of f...

The investigations were carried out in a maize field (1994), vetch-oat field (1995), winter wheat field (1996) and their grassy margins (Moscow region). Staphylinid-beetles were regularly collected by pitfall traps placed in central parts of the fields, on their edges, and in grassy margins around them. The total number of staphylinid-beeltes was 3...

Investigations were conducted in Moscow oblast in 1996. Pitfall traps were used for collecting ground beetle adults in the winter wheat crop field (49 traps) and its grassy margins (32 traps). The effect of some abiotic and biotic factors on the distribution of 9 mass species of beetles was studied. The open space species (Harpalus rufipes, H. qffi...

The investigations were carried out in vetch-oat fields in 1995 and in winter wheat fields in 1996 starting immediately after snow melting. Ground beetles were regularly collected using pitfall traps placed by series in the central part of the fields, on their edges, and in grassy margins around them. The total number of the carabids collected is 2...