Alexander Shmakov

Alexander Shmakov
Altai State University | ASU · Faculty of Biology

About

152
Publications
40,054
Reads
How we measure 'reads'
A 'read' is counted each time someone views a publication summary (such as the title, abstract, and list of authors), clicks on a figure, or views or downloads the full-text. Learn more
1,481
Citations
Citations since 2016
87 Research Items
1396 Citations
2016201720182019202020212022050100150200250300
2016201720182019202020212022050100150200250300
2016201720182019202020212022050100150200250300
2016201720182019202020212022050100150200250300
Additional affiliations
September 1977 - present
Altai State University
Position
  • Professor (Full)

Publications

Publications (152)
Article
Full-text available
New data on the distribution of 40 species from the families Asteraceae, Cystopteridaceae, Dryopteridaceae, Euphorbiaceae, Poaceae, Rosaceae, and Woodsiaceae in Mongolia are presented in the article. For the first time, Centaurea cyanus, Cystopteris almaatensis, Potentilla × habievii, Potentilla salsa, and Potentilla smirnovii are indicated for the...
Article
Full-text available
In the course of field research, analysis of previously published works and review of materials stored in well-known collection funds of Siberia (NS, KRAS, SSHZ), information on the distribution of rare adventitious and native plant species in the Upper Yenisei basin was clarified. For the first time, Arctium lappa, Armoracia sisymbrioides, Bupleur...
Article
Full-text available
В ходе полевых исследований, анализа ранее опубликованных работ и просмотра материалов, хранящихся в известных коллекционных фондах Сибири (NS, KRAS, SSHZ), уточнены сведения о распространении на территории бассейна Верхнего Енисея редких адвентивных и аборигенных видов растений. Во флоре Тывы впервые отмечены Arctium lappa, Armoracia sisymbrioides...
Article
Full-text available
The article presents data on the type specimens of the names of 29 validly published taxa of the genus Potentilla L. described from the Caucasus and some adjacent territories, kept in the Herbarium of the Komarov Botanical Institute RAS (LE). Listed for each taxon are its Latin name with a nomenclatural citation, category of the type specimen(s), f...
Article
Full-text available
Based on morphological characters, a new species Potentilla friesenii collected from alpine belt of Kuraisky Ridge (Altai Mts.) is described. The novelty belongs to Potentilla sect. Chrysanthae; it is most closely related to P. asiatica but differs from the latter by styles not or somewhat widened and smooth to slightly verrucose (vs. always somewh...
Article
The article presents new data on the distribution in the territory of Western Altai of 20 species of vascular plants from the families Asteraceae, Betulaceae, Boraginaceae, Crassulaceae, Huperziaceae, Grossulariaceae, Lamiaceae, Onagraceae, Orchidaceae, Papaveraceae, Polygonaceae, Ranunculaceae, Rosaceae, Saxifragaceae и Violaceae. All of them were...
Article
Full-text available
This paper continues consideration of the spores of three paleotropical fern genera – Taenitis, Syngramma, and Austrogramme (Pteridoideae, Pteridaceae) from South-Eastern Asia and Oceania. At the second stage, we carried out a comparative scanning electron microscopy study of spores of three species of Austrogramme, four species of Syngramma, and s...
Article
Full-text available
Background: The herbarium of the South-Siberian Botanical Garden of Altai State University (ALTB) houses the largest collection of plants from the Altai Mountain Country (AMC), an area that extends across Russia, Kazakhstan, Mongolia and China. The collection of ALTB includes more than 450,00 specimens, making it the seventh largest in Russia and...
Article
In this article, we research the phylogenetic position of the rare endemic banana, Musa huangbaioa, which was described only in Chinese journal in 1987. This banana was found at the foot of the Mount Emei in Sichuan Province and has remarkable morphological features, e. g., undulated petiole margins, ribbed fruits and irregular form of the seeds, w...
Article
Full-text available
For the first time, the Potentilla laciniosa is reported for the flora of Kazakhstan and Altai Mountain Country with the exact location. After a large disjunction of the range, the species was found in the Kyzylbeltau mountains of the Urdzhar region in the southwestern part of the Tarbagatai ridge. Some morphological characters of the new finding a...
Article
Full-text available
The Dzungarian Gobi (DzG), one of 16 phytogeographical regions in the country, is located in the southwestern part of Khovd province in western Mongolia. It comprises some of Mongolia’s largest reserves, namely the Great Gobi B Strictly Protected Area and the National Park Bulgan gol-Ikh Ongog. We conducted a comprehensive survey of the area’s flor...
Article
Based on morphological characteristics revealed for the first time, the species independence of the Crimean endemic Potentilla depressa is confirmed. In contrast to the closely related P. humifusa, the nutlets of P. depressa are wrinkled because of small longitudinal grooves. A special outgrowth looking like a thickened light yellow ring is formed...
Article
Full-text available
Allium toksanbaicum from South East Kazakhstan is described as a new species. Molecular markers reveal a close relationship to A. obliquum and some other central Asian species of the section Oreiprason. We investigated the phylogenetic relationship of the new species based on sequences of two chloroplast spacers (rpl32-trnL and trnQ-rps16) and the...
Article
Full-text available
В ходе полевых исследований, просмотра материалов, хранящихся в гербариях ЦСБС СО РАН (NS), КГПУ им. В. П. Астафьева (KRAS), государственного природного биосферного заповедника «Саяно-Шушенский» (SSHZ) и государственного природного заповедника «Азас», уточнены сведения о распространении в бассейне Верхнего Енисея редких видов растений. Впервые во ф...
Article
During field research, viewing materials stored in the herbaria of the Central Siberian Botanical Garden SB RAS (NS), Krasnoyarsk State Pedagogical University named after V. P. Astafyev (KRAS), the “Sayano-Shushensky” State Natural Biosphere Reserve (SSHZ) and the “Azas” State Nature Reserve, clarified information on the distribution of rare plant...
Article
Full-text available
Constituting one of Earth’s major biomes, steppes are characterised by naturally treeless extra-tropical vegetation. The formation of the Eurasian steppe belt, the largest steppe region in the world, began in Central Asia during the Neogene. In the glacial stages of the Pleistocene, steppe displaced forest vegetation, which in turn recolonised the...
Article
Full-text available
Potentilla radiata, Potentilla rhipidophylla. Аннотация. Приводятся новые данные о распространении 12 таксонов из рода Potentilla L., обнаружен-ных в Казахстане. Растения были собраны на территории Восточно-Казахстанской области (ВКО) в пределах Алтайской горной страны (АГС). Собранные с Калбинского хребта P. radiata Lehm. и с плоскогорья Укок P. r...
Article
Full-text available
In the course of field research, analysis of previously published works and viewing materials stored in the herbarium of the Central Siberian Botanical Garden SB RAS named after I. M. Krasnoborov (NS) we clarified an information on the distribution of rare adventive plant species in the territory of the Upper Yenisei basin. Cosmos bipinnatus, Ipomo...
Article
In this article, we compare the phylogenetic position of various specimens of the Japanese banana (Musa basjoo) originating from the greenhouses of the St. Petersburg Botanical Garden of Peter the Great and from the Kyoto Garden using the analysis of marker sequences ITS1–5.8S rRNA–ITS2 and trnL–trnF. Samples of M. basjoo grown in the Peter the Gre...
Article
Full-text available
The article presents a new data on distribution of 12 taxa of Potentilla L.in Kazakhstan. The plants were gathered on the territory of the East Kazakhstan Region (EKR) within the Altai Mountain Country (AMC). P. radiata Lehm. from the Kalbinsky ridge and P. rhipidophylla Soják from the Ukok Plateau proved to be the new species for Kazakhstan. P. ra...
Article
Full-text available
Analysis of ploidy and DNA content of Asplenium septentrionale s. l. from the Crimea, the Caucasus and Europe showed the presence of diploid forms. This allowed for the first time to draw a conclusion about the growth of Asplenium caucasicum (Fraser-Jenkins et Lovis) Viane in Europe and Russia, described from Asia. For the first time Asplenium onop...
Article
Finding morphological differences between cytotypes that are stable throughout their geographical range is important for understanding evolution of polyploid complexes. The ancient monocot lineage Acorus includes two groups, of which A. calamus s.l., an important medicinal plant, is a polyploid complex with a centre of diversity in Asia. European p...
Poster
Full-text available
To solve one of the tasks of the «Flora of Altai» project on monitoring of rare and endemic plant species, we propose modern approaches and methods of biodiversity informatics. The empirical basis for the two GBIF datasets was the herbarium material of the ALTB with more than 450 thousand specimens
Article
Full-text available
The spores of the following species from genus Cystopteris Bernh from the Trans-Ili Alatau and AltaiSayan regions were studied by scanning electron microscopy (SEM): Cystopteris almaatensis Kotukhov, Cystopterisdickieana R. Sim, Cystopteris gureevi Stepanov, Cystopteris fragilis Bernh. The characters of their perisporium wererevealed. The results o...
Article
The article presents the results of measuring the DNA content by flow cytometry for 6 species of thegenus Trapa L.: Trapa manshurica Fler. (2C = 1.03 pg), T. hankensis Pshenn. (2C = 1.47 pg), T. incisa Siebold & Zucc.(2C = 0.83 pg), T. maximowiczii Korsh. (2C = 0.94 pg), T. pseudoincisa Nakai (2C = 2.18 pg). For T. japonica Fler. twocytotypes 1.12...
Article
Full-text available
Summary. Revision of herbarium material stored in the Herbarium of Altai State University (ALTB, Barnaul) resulted in revealing new localities of the following six species of Oxytropis DC.: O. saposhnikovii Kryl., O. longi-bracteata Kar. et Kir., O. martjanovii Kryl., O. kusnetzovii Kryl. et Steinb., O. ladyginii Kryl., O. lapponica (Wahl.) Ј. Gay....
Article
Full-text available
This work gives the list of representatives of the family Liliaceae Juss., growing on the territory of Altai Mountain Country (AMC) according the materials of Herbarium ALTB including data from digital collections NS (NSK), MW, E, CAS, PE, B and G. As the analysis' result of literary sources, herbarium funds and observations of representatives in n...
Article
Full-text available
The system of the genus Potentilla L. s. str. and its species composition is given for the flora of the Altai Mountain Country (AMC). The structure of the genus is represented by 11 sections. Multifidae with 33 species is the most multitudinous of them. The sections of Fasciculato-pilosae and Fragarioides include one species for each. The general c...
Article
Full-text available
The paper proposes new data and contributions related to distribution of some species from the family Rosaceae in Kyrgyzstan. Five species of the genus Potentilla are newly recorded for the flora of Kyrgyzstan, of which Potentilla tuvinica is a new taxon for the Tian Shan Mountain Range. One species from the genus Fragarias-trum and one from the ge...
Article
Full-text available
This research is the first comprehensive analysis of the intrageneric relationships of Actiniopteris and Onychium based on original data including spore morphology, phylogeny and phylogeography. Actiniopteris and Onychium are members of the large subfamily Pteridoideae of the family Pteridaceae. The Pteridaceae family is considered one of the most...
Article
Full-text available
The article presents the results of measuring the DNA content by flow cytometry of S. sanguinolenta and S. borealis species in 50 populations from natural habitats of Russia and Mongolia. Two probable cytotypes for S. sanguinolenta, as well as single samples with multiple changes, and two cytotypes for S. borealis were identified. Samples with the...
Article
Full-text available
Potentilla. Аннотация. Приводятся новые дополнения к флоре сосудистых растений Казахстана. Материалы статьи содержат информацию о распространении 16 таксонов из рода Potentilla L., собранных на территории данной страны. Впервые для флоры Казахстана приводится 5 видов: P. crebridens Juz., P. × angarensis Popov, P. pen-niphylla Soják, P. tergemina So...
Article
The genus Allium is one of the largest genera in the family of Amaryllidaceae. It comprises over 1000 species, and the number is still increasing. The new intergeneric classification system, based on molecular sequence data, subdivided the genus into 15 subgenera and more than seventy sections. In some sections, the phylogenetic position of species...
Article
Full-text available
Janthina, флористическая на-ходка, Fabaceae, Oxytropis. Аннотация. В ходе обработки гербарного материала, хранящегося в Гербарии Алтайского госуниверситета (ALTB, г. Барнаул), выявлен редкий вид, собранный на территории Республики Алтай-Oxytropis krylovii Schipz., который впервые приводится для флоры России. Ранее этот вид был известен только из фл...
Article
Full-text available
Potentilla chamaeleo var. tytthantha, Potentilla rudolfii. Аннотация. Приводятся новые данные о распространении трёх видов из рода Potentilla, обнаруженных на территории Алтайской горной страны. P. pamirica, собранный на плоскогорье Укок, является новым видом для флоры России; P. tobolensis впервые приводится для флоры Республики Алтай; для P. rudo...
Article
Full-text available
5 Государственный природный биосферный заповедник «Саяно-Шушенский», ул. Заповедная, 7, пос. Шушенское, Ключевые слова: границы ареала, заповедники, миграции, редкие аборигенные виды, чужеродные виды. Аннотация. В ходе полевых исследований, анализа ранее опубликованных работ и просмотра материалов, хранящихся в гербариях Ботанического института РАН...
Article
Full-text available
Синтаксономическое разнообразие высокогорной растительности северо-восточной части Тывы представлено 10 ассоциациями и 1 субассоциацией, отнесенных к 6 классам эколого-флористической классификации. Впервые описано 2 ассоциации и 1 субассоциация. Асс. Сallianthemo sajanensis–Geranietum krylovii ass. nova hoc loco отнесена к провизорному союзу Violo...
Article
Full-text available
Алтайская горная страна (АГС) является самым высоким поднятием Северной Азии, её уровеньбиоразнообразия в три раза богаче Западной Сибири. Биоразнообразие территории АГС исследуется на протяжении двухсот лет. При этом наибольшее число находок приходится на последние 30 лет с момента детальногоизучения Алтайской горной страны специалистами Южно-Сиби...
Article
Full-text available
Алтайская горная страна (АГС) является самым высоким поднятием Северной Азии, уровень её биоразнообразия в три раза богаче Западно-Сибирской равнины. Биоразнообразие АГС исследуется на протяжении 200 лет. При этом, наибольший объем собранного материала по АГС приходится на последние 30 лет с момента начала детального изучения флоры и фауны Алтайско...
Article
Full-text available
For the first time we have conducted a study of three species of the genus Selaginella P. Beauv. from Russia: S. sanguinolenta (L.) Spring, S. helvetica (L.) Link and S. borealis (Kaulf.) Rupr. using the method of scanning electron microscopy (SEM). In a comparative analysis of the morphology of micro-and megaspores of S. san-guinolenta, S. helveti...
Article
Full-text available
Проведен анализ относительного содержания ДНК образцов Selaginella borealis и Selaginella sanguinolenta из различных регионов. Выявлен высокий полиморфизм в относительном содержании ДНК, который связан видимо с различиями в основном хромосомном числе между различными популяциями. Также выявлено одно полиплоидное растение.
Article
Full-text available
The article provides an additional information on the flora of the State National Natural Park “Altyn-Emel”. The natural park is located in the Almaty Region of the Republic of Kazakhstan and occupies the territory on the south-western part of the Dzungarian Alatau ridge. For the first time, 4 species of the genus Allium L. and 5 species of the gen...
Article
Full-text available
The techniques of biotechnology are considered promising and highly-demanded for propagation and conservation of rare species of medicinal plants. The variety of natural forms of Rhodiola rosea makes it difficult to develop a single efficient in vitro protocol for this valuable and rare medicinal plant. The optimization of in vitro methods for spec...
Article
Full-text available
The synopsis reflects data on the diversity, synonymy and distribution of genera and species of the family Woodsiaceae. The largest species diversity of the family is located in the mountain systems of America (northern and central) and Eurasia (central and eastern). The family Woodsiaceae is represented by 7 genera (Cheilanthopsis, Eriosoriopsis,...
Article
Full-text available
A taxonomic study of the genus Campanula L. in the flora of Altai is presented. Based on the data obtained, 14 Campanula species, belonging to 3 subgenera and 7 sections, grow in the territory of the Altai Mountain Country. The subgenus Campanula includes 4 sections and 8 species and is the most diverse in the flora of Altai. An original key is pre...
Article
Full-text available
The synopsis presents the most comprehensive data on the diversity, synonymy and distribution of species of the genus Saussurea DC. in Eurasia. The greatest species diversity of the genus occurs in the mountain systems of Central and SouthEast Asia, the Himalayas and Tibet. To date more than 1200 taxa of the species and subspecies level has been de...
Article
Full-text available
The synopsis of the genus Cystopteris Bernh., consisting of 32 species and 5 hybrids, is given in the article. An information on the type material, its localization and the general distribution of species is presented for all taxa. Two new combinations are designated: Cystopteris huteri (Hausm. ex Milde) Shmakov, comb. et stat. nov. and sect. Khokh...
Article
Full-text available
A study of spores of the single species of the genus Parahemionitis Panigrahi was performed using the method of scanning electronic microscopy (SEM). Spores of Parahemionitis arifolia (Burm. f.) Panigrahi are tetrahedral trilete, roundish-triangular in polar position, with micro-wrinkled exospore and sculptured perispore. Sculpture of perispore is...
Article
Full-text available
A comparative study of spores of seven species: Taenitis blechnoides (Willd.) Sw., T. hookeri (C. Chr.) Holttum, T. pinnata (J. Sm.) Holttum, Syngramma alismifolia (Presl) J. Sm., S. lobbiana (Hook.) J. Sm., S. quinata (Hook.) Carruth., and Austrogramme boerlageana (Alderw.) Hennipman from South-Eastern Asia was performed using the method of scanni...
Article
Full-text available
NS) и государственного природного биосферного заповедника «Саяно-Шушенский» (SSHZ), уточнены сведения о видовом составе флоры Западного Саяна. Впервые в горной системе обнаружены восемь видов: Bupleurum sibiricum, Ephedra regeliana, Epilobium roseum, Lotus krylovii, Saussurea serratuloides, Setaria faberi, Viola jeni-seensis, V. schauloi. У 28 редк...
Article
Full-text available
The critical revision of Trollius L. (Ranunculaceae) in the Far East of Russia was made, in which nine species were recognized. The identification key and taxonomical synopsis of the genus have been provided. Synony-my, geographical distribution and coeno-ecological peculiarities of each species of these nine species are presented. For the first ti...
Article
Full-text available
p> Trichoderma isolates (SSBGT07, SSBGT08, SSBGT09, SSBGT10) were isolated from the soil samples of the South-Siberian Botanical Garden and identified using morphological observation and ITS region analysis as Trichoderma harzianum , T. asperellum, T. ghanense , and T. longibranchiatum . Antagonistic activity against Cladosporium sp. and Botrytis s...
Article
Full-text available
As a result of expedition work in 2010–2016 in the Altai Krai and the Altai Republic, new records of Stipa grandis P. A. Smirn. were found for Western Siberia. Also new locations of rare and protected species of the genus Stipa were found: 5 for Stipa lessingiana Trin. et Rupr., and one by one – S. dasyphylla (Lindem.) Czern. ex Trautv., S. pulcher...
Article
Full-text available
p> Simplicillium lanosoniveum isolate SSBG2 was isolated from the diseased C. hesper collected from Schefflera octophylla in greenhouse of the South-Siberian Botanical Garden, and identified based on morphological observation and ITS region analysis. The infected plants were inoculated with conidial suspension of the isolate SSBG2 in concentrations...
Article
Full-text available
Comparative investigation of the spore surface ultrastructure of six specimens of Pityrogramma calomelanos (L.) Link (Pteridoideae, Pteridaceae) was carried out with using of scanning electronic microscopy (SEM). Specimens of Pityrogramma calomelanos were collected in the different parts of its range: in South China (Hainan Island), Nepal, Honduras...
Article
Full-text available
The ultrastructure of the spore surface of Onychium ipii Ching (Pteridoideae, Pteridaceae) was investigated by using of scanning electron microscopy. Spores of Onychium ipii are trilete, tetrahedral, with hemispherical distal side and convex proximal one. Equatorial diameter 42.4-48.1 μm, polar axis 35.7-40.6 μm. Distal and proximal sides are separ...
Article
Full-text available
The ultrastructure of the spore surface of 6 species Cystopteris Bernh. subgenus Cystopteris (Cystopteridaceae): C. almaatensis Kotukhov, C. altajensis Gureeva, C. dickieana R. Sim, C. fragilis (L.) Bernh. C. gureevae Stepanov, C. protrusa (Weatherby) Blasdell was investigated by using of scanning electron microscopy. On the basis of differences in...
Article
Full-text available
p>For the territory of the Altay Mountain are the location of new fern species belonging to families of Athyriaceae, Cryptogrammaceae, Cystopteridaceae, Dryopteridaceae, Hypolepidaceae, Onocleaceae, Polypodiaceae, Thelypteridaceae, Woodsiaceae.</p
Article
Full-text available
p>Scanning electron microscopy (SEM) was used to perform a comparative study for four representatives of the subfamily Ceratopteridoideae (J. Sm.) R.M. Tryon from East Asia: Ceratopteris thalictroides Brongn., C. pteridoides (Hook.) Hieron., Acrostichum aureum Linn., and A. speciosum Willd. The analysis of the external morphology of the representat...
Article
Full-text available
We used the method of scanning electron microscopy (SEM) to study the representatives of the genus Anogramma from South-West Asia and the Himalayas. To confirm the presence of Cerosora microphylla (Hook.) R.M. Tryon in China (hence, the absence of Anogramma microphylla (Hook.) Diels), we revealed the features of external morphology of spores that a...