Alexander V. Fateryga

T. I. Vyazemsky Karadag Scientific Station – Nature Reserve of RAS – Branch of A. O. Kovalevsky Institute of Biology of the Southern Seas of RAS

The book describes native orchids of the Crimea. A short overview, color photographs, and a map showing distribution within the Crimea are presented for each of the 45 species. A key to identification of all taxa is also provided. The book is dedicated for specialists in nature conservation, students, and everybody who is interested in the Crimean...

The Palaearctic complex of anthidiine bees closely related to Pseudoanthidium scapulare has long been a source of unresolved taxonomic and systematic issues. Until now, the number of species in the complex and their geographical distributions were largely unclear, thus complicating the compilation of accurate species checklists and hindering conser...

Celonites ivanovi sp. nov. is described as a new species from Dagestan where it has been recorded from dry habitats in a small area on the northern side of the Greater Caucasus. Celonites cagrii sp. nov. is described from Erzurum Province in east Turkey. As in other members of the C. cyprius-group, the females of both species were observed to visit...

Celonites kozlovi Kostylev, 1935, and C. sibiricus Gusenleitner, 2007, coexist in semi-deserts of the Altai Mountains. The trophic relationships of these pollen wasp species to flowers are largely unknown. We observed the flower visits and behaviors of wasps on flowers; pollen-collecting structures of females were studied using SEM; the taxonomic p...

Ancistrocerus capra was described by de Saussure in 1857 from North America and then was synonymized with the Palaearctic A. antilope (Panzer, 1798) by Bequaert in 1944. Although these species share a combination of two characters (impunctate and shining metapleuron and lateral surface of the propodeum and a bifurcate apex of the aedeagus) not know...

Two nests of Hoplitis curvipes are described from Apulia (Italy) and Dagestan (Russia). Both nests consisted of two brood cells placed side by side under a stone. The cells were neither attached to each other nor to the substrate. They were constructed from leaf fragments, which were imbricately arranged, forming a cone-like structure; each leaf fr...

Heinrich Friese described Anthidium spiniventris [sic] from Palestine in 1899, and A. melanopygum as a “variety” of it from Turkey in 1917. While A. melanopygum was subsequently recognized as a subspecies of A. spiniventre, a morphological examination of new material of both taxa suggests that these taxa represent distinct species. This was also co...

Isodontia nigella (F. Smith, 1856) is native to the Eastern Palaearctic, Oriental region, and Australia. A sheaf of reed canes installed as a trap nest in the Crimea was occupied with 73 nests of this species in 2021. The nests contained one to eight cells separated by partitions made of packed fragments of grass stalks and blades; the closing plug...

Hoplitis astragali sp. nov., a member of the H. monstrabilis species group, and H. dagestanica sp. nov., a member of the H. adunca species group, are described. The former species is known from Dagestan in Russia, Azerbaijan, and Turkmenistan, the latter only from Dagestan. Nests of H. astragali are described. Females of this species excavated burr...

Onychopterocheilus (Asiapterocheilus) proshchalykini Fateryga, sp. nov. is described from the Altai Republic (Russia). The new species is closely related to O. (A.) kiritshenkoi (Kostylev, 1940) but distinctly differs from it by the structure of the clypeus, the female vertex and scutellum, the male genitalia, as well as the coloration. In addition...

Celonites montanus Mocsáry, 1906, described from the Alai Range in Kyrgyzstan, is synonymized with C. kostylevi Panfilov, 1961, syn. nov. and C. haemorrhoidalis Gusenleitner, 2012, syn. nov. (both also described from Kyrgyzstan). The species is reported from Kazakhstan (Almaty Province) for the first time. It was collected at nototribic flowers of...

The knowledge of the solitary Vespidae of Uzbekistan is very poor and a preliminary list of species is provided. The list contains 105 species of 34 genera: seven species of three genera in the subfamily Masarinae, 96 species of 29 genera in Eumeninae s. str., one species in Raphiglossinae and one in Zethinae. Among them, eight genera and 27 specie...

Cephalanthera caucasica had been reported previously from the Samur Forest in Dagestan (Russia), but that report was considered doubtful: a possible misidentification of supposed hybrids between C. damasonium and C. longifolia as C. caucasica was speculated. Moreover, this species was recently reduced to a synonym of C. kotschyana in the «Plants of...

The following bee species new to the Kazantip Nature Reserve are reported: Colletes hederae Schmidt & Westrich, 1993, C. similis Schenk, 1853, Andrena marginata Fabricius, 1776, as is one species of vespid wasp, Euodynerus posticus (Herrich-Schäffer, 1841). In late September/early October, bees were recorded on two forage species: Galatella villosa...

The data on 19 species of megachilid bees collected in the Nakhchivan Autonomous Republic of Azerbaijan in 2019-2022 are reported. Fifteen species are new for Azerbaijan; four other species, previously known by literature records, are confirmed for Azerbaijan. Anthidium gussakovskiji Mavromoustakis, 1939 is excluded from the fauna of the republic....

Two nests of Ancistrocerus oviventris (Wesmael, 1836) are described. One nest from Poland was attached to a concrete fence and consisted of 10 mud cells. The second nest from the Crimea was attached to a stone and consisted of three mud cells covered with an additional layer of mud and gravel particles. Females provisioned their cells with moth cat...

Taraxacum hybernum Steven (Asteraceae family) native to Southeast Europe is a promising rubber-bearing dandelion species. In the 1930s, several phenotypically distinct forms were described in this dandelion, such as brown-achened and pinkish-achened. More than 70 years after termination of research of temperate zone rubber-bearing plants we were ab...

Four new species are described: Pseudepipona (Deuterepipona) kostylevi Fateryga, sp. nov. (Turkmenistan, Uzbekistan, Kazakhstan), P. (D.) nikolayi Fateryga, sp. nov. (Kazakhstan), P. (D.) popovi Fateryga, sp. nov. (Turkmenistan), and P. (D.) vladimiri Fateryga, sp. nov. (Kazakhstan). A new synonymy is proposed: Pseudepipona (Deuterepipona) superba...

New records of Celonites tauricus Kostylev, 1935 are reported from Chios, Rhodes, Samos (Greece), Dagestan (Russia), Georgia, and the main portion of Azerbaijan (previous records were made from the Nakhchivan Autonomous Republic only). Flower visits of imagines were observed at six species of Lamiaceae, four of them being recorded for the first tim...

Nesting of a Katamenes sichelii (de Saussure, 1852) female was observed with timekeeping of its activity. The nest was located on a rock outcrop. The cell walls were built by the wasp from small pebbles that were glued to each other and to the substrate with earthen mastic made by moistening dry clay with regurgitated liquid. The vault and neck of...

The Pseudepipona herrichii-group includes species of the genus Pseudepipona de Saussure, 1856, subgenus Pseudepipona s. str., without long setae on the gena, the scutum, and the propleura, with the transverse carina of the propodeum forming laterally a tooth or at least a blunt angle, and with the preapical tooth of the male mandible of approximate...

With this paper we continue a new annual series, the main purpose of which is to make significant floristic findings from Russia and neighboring countries more visible in Russia and abroad. In total, this paper presents new records for 48 vascular plant species from 6 Eurasian countries, obtained during field explorations, as well as during taxonom...

Quartinia mongolica (Morawitz, 1889), a species hitherto known only from the type series collected in China (Gansu), is reported from Mongolia (Khovd Province). The species is closely related to Q. funebris Kostylev, 1935 known from Turkmenistan. They differ in coloration, integumental sculpture, and the structure of the male genitalia. Lectotypes...

Two natural interspecific hybrids of Ophrys mammosa Desf. s. l. and O. oestrifera M. Bieb. (O. × aghemanii Renz) are reported from Russia: a very rare O. mammosa subsp. mammosa × O. oestrifera from the Crimea (vicinity of Zelenogorye) and a relatively abundant O. mammosa subsp. caucasica (Woronow ex Grossh.) Soó × O. oestrifera from the Krasnodar T...

Orobanche ingens is an endemic species from the Caucasus, especially the Greater Caucasus, parasitising on large Apiaceae (usually Heracleum). This species was misclassified over the years and little was known about its range and habitats. Here, we clarify the typification, as well as provide notes about the taxonomy of this species. Additionally,...

Nesting of a Katamenes sichelii (de Saussure, 1852) female was observed with the timekeeping of its activity. The nest was located on a rock outcrop. The walls of a cell were built by the wasp from small pebbles attached to each other and to the substrate with the help of earthen mastic made by moistening dry clay with regurgitated liquid. The vaul...

A new invasive bee species, Megachile sculpturalis Smith, 1853 was revealed nesting in Simferopol (Crimea) in 2019. Bees occupied a "bee hotel" located on the 5 th floor of a building on the outskirts of the town. These bees were transferred to one and two new localities in 2020 and 2021, respectively. A total of 86 nesting females were found in th...

This is the fourteenth of a series of miscellaneous contributions, by various authors, where hitherto unpublished data relevant to both the Med-Checklist and the Euro+Med (or Sisyphus) projects are presented. This instalment deals with the families Apocynaceae, Compositae, Crassulaceae, Cyperaceae, Euphorbiaceae, Gramineae, Leguminosae, Nyctaginace...

Supplementary figures: Fig. S1. Nerium oleander L.; Fig. S2. Mirabilis jalapa L. (photographs by P. Novák)

New combinations are validated: Jacobaea erucifolia subsp. grandidentata (Ledeb.) V.V. Fateryga & Fateryga, comb. nov. (Asteraceae), Bituminaria bituminosa subsp. pontica (A.P. Khokhr.) V.V. Fateryga & Fateryga, comb. nov. (Fabaceae) and Phlomis herba-venti subsp. taurica (Hartwiss ex Bunge) V.V. Fateryga & Fateryga, comb. & stat. nov. (Lamiaceae).

Raphiglossa rasnitsyni Fateryga, sp. n. is described from Turkmenistan. The new species resembles R. eumenoides S.S. Saunders, 1850 in its body size and proportions of the metasomal tergum 1 but strongly differs from it by having a narrow propodeal concavity which is less wide than the metasomal tergum 1 at base, anterior margin of clypeus deeply e...

The nest structure of Eustenancistrocerus amadanensis was studied in Crimea. Females nested in the ground, in level or gently sloping spots with sparse vegetation. The soil in which the nests were built varied broadly in granulometric composition, from sandy loams to dense clay loams resembling clay in their properties. Of 11 nests studied, 5 had b...

Nests of Pareumenes quadrispinosus (de Saussure, 1855) were obtained at Me Linh Station for Biodiversity (323 nests) and Tam Dao Town (283 nests), Vinh Phuc Province, as well as at Phu Luong, Thai Nguyen Province (9 nests) and Kim Boi, Hoa Binh Province (62 nests). The wasps nested in segments of bamboo canes and reed stems, 5 to 18 mm in internal...

An account on 35 species in the family Vespidae (subfamilies Masarinae, Eumeninae s. str., and Raphiglossinae) is presented. Genera Jugurtia de Saussure, 1854, Quartinia André, 1884, Brachypipona Gusenleitner, 1967, and Cyrtolabulus van der Vecht, 1969, as well as 17 species and one subspecies are new to Azerbaijan. Three additional species are new...

Two females of Leptochilus regulus (de Saussure, 1855) were observed nesting in reed stalks of a Fabre’s hive serving as a block of trap nests in Crimea, with the timekeeping of all their nesting behaviours. The building material used by the females to separate the nesting cavity into the cells consisted of pellets of dry soil, gravel particles, an...

A list of 54 bee species of the family Megachilidae with data on their distribution and relative abundance in Kemerovskaya Oblast is presented. Five species, Megachile fulvimana Eversmann, 1852, M. nigriventris Schenck, 1870, Osmia inermis (Zetterstedt, 1838), O. nigriventris (Zetterstedt, 1838) and Pseudoanthidium tenellum (Mocsáry, 1881), are rec...

The presented dataset contains information about findings of vascular plant species found in Russia and adjacent countries and published in Botanica Pacifica Journal. In total, this dataset includes 38 species belonging to 34 genera and 23 families. The study is based on data obtained during field explorations, as well as during taxonomic revisions...

With this paper we continue a new annual series, the main purpose of which is to make significant floristic findings from Russia and neighboring countries more visible in Russia and abroad. In total, this paper presents new records for 38 vascular plant species from 7 Eurasian countries, obtained during field explorations, as well as during taxonom...

The data on 38 species of megachilid bees in the tribes Lithurgini, Dioxyini, and Megachilini collected in the Nakhchivan Autonomous Republic of Azerbaijan mainly in 2018–2020 are given. Seventeen species are new to Azerbaijan, four other species are new to the Nakhchivan Autonomous Republic and as a result of the present investigation the fauna of...

Two new genera are described for three species in the "red" group of the North American eumenine wasps from the genus Odynerus Latreille, 1802: Bohartodynerus Fateryga, gen. n. for O. margaretellus Rohwer, 1915 (type species) and O. cinnabarinus Bohart, 1939, and Parkerodynerus Fateryga, gen. n. (type species O. erythrogaster Bohart, 1939). Boharto...

The nest structure of Eustenancistrocerus amadanensis was studied in the Crimea. Females nested in the ground, on horizontal or gently sloping spots with sparse vegetation. The granulometric composition of the soil, in which the wasps nested, varied broadly from sandy loams to dense clay loams close by their properties to clay. Eleven nests were st...

The genus Holosteum L. distributed mostly in temperate Eurasia has very confused taxonomy. Modern estimation of the species richness varies from one species worldwide to seven species occurring just in Eastern Europe. One of the most problematic species of the genus is H. glutinosum (M. Bieb.) Fisch. et C. A. Mey. usually treated as a subspecies or...

The orchid genera Epipactis Zinn and Ophrys L. are well-known by their complicated taxonomy and extensive debates over species richness within them. These genera are represented in Turkmenistan by two species each. Two of them, namely E. turcomanica K. P. Popov et Neshat. and O. kopetdagensis K. P. Popov et Neshat., were hitherto accepted as specie...

The nesting biology of Chalybion turanicum (Gussakovskij, 1935) has been studied, with a total of 31 nests being examined. All studied nests were located inside the old nest cells of Sceliphron destillatorium (Illiger, 1807). Each nest of Ch. turanicum consisted of a single cell. Females hunted for spiders, with 18 species in five families being id...

New additions to the knowledge of the subfamily Eumeninae in Russia are provided. Stenodynerus rossicus Fateryga & Kochetkov, sp. nov. is described from Amurskaya Province and Altai Republic. Three species of eumenine wasps are reported from Russia for the first time: Onychopterocheilus kiritshenkoi (Kostylev, 1940), Pterocheilus quaesitus (Morawit...

New data on 22 species of bees of the family Megachilidae from the North Caucasus and the south of European Russia are reported. Six species are new to Russia: Hoplitis curvipes (Morawitz, 1871), Osmia cinerea Warncke, 1988, O. ligurica Morawitz, 1868, O. cyanoxantha Pérez, 1879, Protosmia glutinosa (Giraud, 1871), and Coelioxys mielbergi Morawitz,...

A list of 121 plant species in 33 families on which eumenine wasps were recorded feeding on nectar in Crimea is given. Forage plants were ascertained for 58 wasp species. The largest numbers of forage plant species belonged to the families Apiaceae (18), Asteraceae (17), and Lamiaceae (13). The largest numbers of wasp species were recorded on fl ow...

Nesting of Leptochilus limbiferus (Morawitz, 1867) was observed in the Krasnodar Territory, Russia. Twelve nests were studied; all of them were located in empty shells of a terrestrial snail species, Xeropicta derbentina (Krynicki). The building material used by the wasps was pellets of dry soil and gravel particles, as well as shell bits, and some...

Data on 34 species of anthidiine bees collected in Nakhchivan Autonomous Republic of Azerbaijan are reported. Sixteen species are new to Azerbaijan as a whole, and six other species are new to Nakhchivan Autonomous Republic. A total of 46 species of Anthidiini are currently known from Azerbaijan, 41 of them are known from Nakhchivan Autonomous Repu...

This is the twelfth of a series of miscellaneous contributions, by various authors, where hitherto unpublished data relevant to both the Med-Checklist and the Euro+Med (or Sisyphus) projects are presented. This instalment deals with the families Asparagaceae (incl. Hyacinthaceae), Boraginaceae, Cactaceae, Caryophyllaceae, Chenopodiaceae, Compositae...

A list of 121 plant species in 33 families on which feeding of eumenine wasps on nectar was recorded in the Crimea is given. Forage plants were ascertained for 58 wasp species. The largest numbers of forage plant species belonged to the families Apiaceae (18), Asteraceae (17), and Lamiaceae (13). The largest numbers of wasp species were recorded on...

Bionomics of Euodynerus fastidiosus (de Saussure) was studied in Crimea. One nest located in an old burrow of Cicadatra querula (Pallas) and 13 nests located in the cells of old nests of Sceliphron spp. were examined. The nests contained 1–4 cells separated by partitions made of soil mastic which the wasps prepared by mixing dry earth with water. T...

The distribution of Celonites kozlovi Kostylev, 1935 and C. sibiricus Gusenleitner, 2007 is summarized. Celonites kozlovi is firstly reported from Russia and C. sibiricus is firstly reported from Kazakhstan. The lectotype of C. kozlovi is designated and its type locality in Mongolia is specified in detail (to the south of Mt. Dund Saikhany Nuruu, Ö...

Twenty species of the orchid genus Cephalanthera are known in the World and seven species occur in Russia. One of them, restricted in Russia to the Black Sea coastal area of the Krasnodarsky Krai in the North Caucasus, has a long story of misidentifications and taxonomic confusions. The present study revealed that the correct name for this species...

Bionomics of Euodynerus fastidiosus (de Saussure) were studied in the Crimea. One nest located in an old burrow of Cicadatra querula (Pallas) and 13 nests located in cells of old nests of Sceliphron spp. were examined. The nests contained 1–4 cells separated by partitions made of soil mastic prepared by wasps by mixing dry earth with water. Lateral...

Brachypipona kurzenkoi Fateryga, sp. n. is described by females from Aktobe, Jambyl, and Almaty provinces of Kazakhstan. The new species is closely related to B. longicornis (Morawitz, 1895) but differs from it by pronotal carina weakly developed and not acutely producing laterally, epicnemial suture not carinate, mesepisternum more sparsely punctu...

Nesting of Stenancistrocerus obstrictus was studied in the Republic of Dagestan, Russia. Ten nests were observed, six of them were dissected and their structure was studied. The nests were located in preexisting holes in a clayey cliff. Females searched for a place for the nest during more than a day. The holes with nests were 2.5–10.5 cm in length...

This is the eleventh of a series of miscellaneous contributions, by various authors, where hitherto unpublished data relevant to both the Med-Checklist and the Euro+Med (or Sisyphus) projects are presented. This instalment deals with the families Anacardiaceae, Asparagaceae (incl. Hyacinthaceae), Bignoniaceae, Cactaceae, Compositae, Cruciferae, Cyp...

Megachile sculpturalis Smith, 1853 was revealed nesting in a trap nest and a “bee hotel” in Simferopol. The nests were built of conifer resin with addition of mud and sawdust. Females visited inflorescences of Eryngium campestre L., Inula helenium L., and Carduus acanthoides L. while pollen samples taken from one foraging female contained only poll...

The variety Limodorum abortivum var. viride is known from five localities of the Crimean South Coast. Communities of Quercus pubescens represent all of them, while L. abortivum var. abortivum grows in various forest types. Limodorum abortivum var. viride differs from the nominotypical variety by shorter, thinner, and completely green floral stem an...

The list of orchids in the flora of the Crimea numbers 45 species. Comments are provided for taxa reported in earlier publications (2012–2018) but absent in the present list due to their synonymization or other nomenclatural changes. Taxa added and excluded in comparison with the list published by A. V. Yena (2012) are particularly listed.

New additions to the knowledge of the subfamily Eumeninae in Russia are provided. Leptochilus (Lionotulus) leleji Fateryga, sp. nov. is described from Altai Republic. Males of Ancistrocerus hangaicus Kurzenko, 1977 and Jucancistrocerus (Eremodynerus) minutepunctatus Giordani Soika, 1970 are described for the first time; the latter species is transf...

This is the tenth of a series of miscellaneous contributions, by various authors, where hitherto unpublished data relevant to both the Med-Checklist and the Euro+Med (or Sisyphus) projects are presented. This instalment deals with the families Amaranthaceae, Campanulaceae, Caryophyllaceae, Chenopodiaceae, Compositae, Crassulaceae, Cruciferae, Cyper...

Trap nests for aculeate Hymenoptera were exposed during 15 years (2002–2016) in 80 localities of Crimea. A total of more than 500 trap nests containing about 25 000 cavities were used. As a result, 6 895 nests of 97 species from 9 families of Hymenoptera were obtained: Pompilidae (5 species / 118 nests), Vespidae (27/1201), Ampulici-dae (1/6), Sphe...

New data on the distribution of 37 species are reported. Eleven species are new to Russia: Lithurgus tibialis Morawitz, 1875, Anthidium spiniventre Friese, 1899, Icteranthidium ferrugineum (Fabricius, 1787), Hoplitis carinata (Stanek, 1969), Coelioxys acanthura (Illiger, 1806), C. decipiens (Spinola, 1838), Megachile albonotata Radoszkowski, 1886,...

A review of 80 vascular plant taxa is given. There are 47 species and subspecies (including five hybrids) added to the flora of the Karadag State Nature Reserve (Crimean Peninsula). Among them, 29 taxa have been reported from the Karadag Mountains for the first time and the 18 remaining taxa have already been listed in different references. At the...