Alexander AverianovRussian Academy of Sciences | RAS · Zoological Institute
Alexander Averianov
Doctor of Biological Sciences
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491
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Introduction
Additional affiliations
June 2010 - present
January 2009 - present
January 2007 - present
Rheinische Friedrich-Wilhelms-Universität Bonn
Education
September 1983 - June 1988
Publications
Publications (491)
А detailed morphological characterization of deciduous teeth in 7 genera and
8 species of recent carnivorous marnmals of the farnily Procyonidae is given. There
were three ways of adaptation of deciduous teeth in representatives of the farnily,
i. е. ornnivorous, in Procyon, Nasua, and Nasuella, frugivorous, in Bassarycion
and Potos, and retaining...
A tegotheriid docodontan Sibirotherium rossicum from the Early Cretaceous of Western Siberia, Russia, was considered to have six lower premolars, as in the tegotheriids Agilodocodon and Microdocodon from the Middle Jurassic of China. The micro-computed tomography of two dentary specimens with a supposed submolariform ultimate deciduous premolar (dp...
According to the degree of development of the main cusps and the presence of a small lingual part, the upper molar of a docodontan from the Bol’shoi Kemchug 4 locality (Ilek Formation, Lower Cretaceous) in the Krasnoyarsk Territory corresponds to M1 of Agilodocodon scansorius Meng et al., 2015 from the Middle Jurassic of China. The described tooth...
Eutriconodonta are an important group of early crown mammals with a wide distribution in the Jurassic-Cretaceous of the Northern Hemisphere and few occurrences in the Southern Hemisphere. Three taxa of eutriconodontans are known from the Early Cretaceous high-latitude Teete vertebrate assemblage in Yakutia, Russia: Sangarotherium aquilonium (Eutric...
According to the degree of development of the main cusps and the presence of a small lingual part, the upper molar of a docodontan from the Bol’shoi Kemchug 4 locality (Ilek Formation, Lower Cretaceous) in the Krasnoyarsk Territory corresponds to M1 of Agilodocodon scansorius Meng et al., 2015 from the Middle Jurassic of China. The described tooth...
The isolated cervical vertebra PIN 329/16 from the Aptian Ilek Formation at Shestakovo 1 locality in Western Siberia, Russia, is unusual because of its elongated neural arch, extremely constricted neural arch around its midlength, and by possessing a shallow postzygapophyseal centrodiapophyseal fossa placed adjacent to the postzygapophyses. It diff...
The ontogenetic status of the known specimens of alvarezsaurid theropod dinosaurs is revised. We present histological data that the holotype of Parvicursor remotus from the Upper Cretaceous Barungoyot Formation of Mongolia is not an adult individual, as previously thought, but a juvenile less than one year old. The miniaturization took place only i...
A palaeophiid sea snake Pterosphenus muruntau Averianov, 1997 was based on two trunk vertebrae and several vertebral fragments from the middle Eocene (Bartonian) coastal marine deposits of Dzheroi 2 locality in the Central Kyzylkum Desert, Uzbekistan. The species was considered a no-men dubium by Rage and coauthors in 2003 based on alleged poor pre...
In Russia, sauropod remains have been found in the Middle Jurassic of Western Siberia (Mamenchisauridae), the Lower Cretaceous of the Volga region (Volgatitan simbirskiensis), Transbaikalia (Tengrisaurus starkovi), and Western Siberia (Sibirotitan astrosacralis), and the Upper Cretaceous of the Far East (Opisthocoelicaudidae). Most finds from the L...
There are about 1000 valid species level taxa of dinosaurs in the world, 85% of them were described after 1990. Twelve taxa originate from Russia, while 10 of them were also established after 1990. Over the past 30 years, the number of publications devoted to the dinosaurs of Russia has increased 6.6 times. The first finds on the territory of Russi...
Two isolated metatarsals (III and IV) of theropod dinosaurs from the Bostobe Formation (Upper Cretaceous, Santonian-Campanian) of the Shakh-Shakh locality in Kazakhstan are assigned to the representatives of the families Caenagnathidae and Dromaeosauridae, respectively. The metatarsal III of Caenagnathidae indet. confirms the presence of oviraptoro...
Two isolated metatarsals (III and IV) of theropod dinosaurs from the Bostobe Formation (Upper Cretaceous, Santonian–Campanian) of the Shakh-Shakh locality in Kazakhstan are assigned to the representatives of the families Caenagnathidae and Dromaeosauridae, respectively. The metatarsal III of Caenagnathidae indet. confirms
the presence of oviraptoro...
Isolated sacrum fragments, manual ungual phalanges, and a third metatarsal from the Upper Cretaceous (Santonian) Yalovach Formation at the Kansai locality in Tajikistan, show similarities to those elements in various dromaeosaurid theropods and are referred to Kansaignathus sogdianus, previously described on the basis of a dentary from the same loc...
A new hyaenodont Maocyon peregrinus , gen. et sp. nov., is described based on a partial skull and associated mandible from the upper Eocene Youganwo Formation at Maoming locality in Guangdong Province, China. It shows certain similarities with the Hyainailouroidea in the skull structure, including anteroposteriorly extended jugal/squamosal suture,...
The Upper Cretaceous (Turonian) Bissekty Formation of Uzbekistan has yielded many isolated bones and teeth representing a variety of non-avian theropod dinosaurs. A pedal phalanx II-2 indicates the presence of a dromaeosaurid theropod that attained a larger body size than any previously known member of that clade. The same formation also yielded a...
The paper presents a brief biography and scientific achievements of a renowned Russian
mammologist Alexei Vladimirovich Abramov (b. 1962) in relation to his 60th anniversary. The paper is illustrated by photos obtained from colleagues who personally know A.V. Abramov and from his family archive.
A distal fragment of tibiotarsus from the Late Cretaceous Kansai locality (Yalovach Formation, Santonian) in Tajikistan was assigned to oviraptorosaurian theropods of the family Avimimidae by the fusion of the tibia and proximal tarsal bones into tibiotarsus, reduction of the fibula and fusion of its distal end with the lateral condyle of the tibio...
The previously unknown second cervical vertebra (axis) of the sauropod Sibirotitan astrosacralis Averianov et al., 2018 from the Early Cretaceous locality Shestakovo 1 (Ilek Formation, Aptian) in Kemerovo oblast, Russia, makes it possible to supplement the morphological characterization of this dinosaur species. The described axis is characterized...
A distal fragment of tibiotarsus from the Late Cretaceous Kansai locality (Yalovach Formation, Santonian) in Tajikistan was assigned to oviraptorosaurian theropods of the family Avimimidae by the fusion of the tibia and proximal tarsal bones into tibiotarsus, reduction of the fibula and fusion of its distal end with the lateral condyle of tibiotars...
Ранее неизвестный второй шейный позвонок (аксис) завропода Sibirotitan astrosacralis Averianov et al., 2018 из раннемелового местонахождения Шестаково 1 (илекская свита, апт) в Кемеровской области, Россия, позволяет дополнить морфологическую характеристику этого вида динозавров. Для описанного аксиса характерны такие уникальные признаки, как плоска...
Bogolubovia orientalis, the first pterosaur taxon described from Russia, was known for more than a hundred years from a single specimen, the posterior fragment of the middle cervical vertebra from the Upper Cretaceous (upper Campanian) Rybushka Formation at Malaya Serdoba locality in Penza oblast, Russia. First classified as a pteranodontid, Bogolu...
A theropod femur from the Late Cretaceous Kansai locality in Tajikistan (Yalovach Formation, Santonian) is referred to the family Therizinosauridae based on a combination of the following derived characters: the bone is straight in the anteroposterior plane; the femoral head is directed dorsomedially; the fem-oral neck is constricted between the fe...
A theropod femur from the Late Cretaceous Kansai locality in Tajikistan (Yalovach Formation, Santonian) is referred to the family Therizinosauridae based on combination of the following derived characters: the bone is straight in anteroposterior plane; the femoral head is directed dorsomedially; the femoral neck is constricted between the femoral h...
The revised dinosaur assemblage from the Lower Cretaceous Murtoi Formation at Mogoito locality in Transbaikalia, Russia, includes lithostrotian titanosaur Tengrisaurus starkovi known from caudal vertebrae, Sauropoda indet. represented by isolated teeth and metatarsal IV, theropods Ornithomimosauria, Therizinosauria, Dromaeosauridae, Theropoda indet...
Международной командой палеонтологов из России (Москва, Санкт-Петербург и Томск) и Германии (Бонн), исследующей позвоночных юрского периода в Красноярском крае России (Березовский карьер около г. Шарыпово), описаны новые виды млекопитающих из группы эухарамийид, многие представители которой имели летательную перепонку и могли планировать с дерева н...
Tashtykia primaeva и Tagaria antiqua из средней юры Березовского карьера около города Шарыпово в Красноярском крае заполняют морфологическую дистанцию между другими мультитуберкулятами и среднеюрскими переходными формами от эухарамийид к многобугорчатым – кермакодоном из Англии и мегаконом из Китая. В связи с открытым разнообразием среднеюрских мул...
Nemegtosaurus mongoliensis надежно отличается от Opisthocoelicaudia skarzynskii чертами строения спинных позвонков и бедренной кости, и должен сохранить свое самостоятельное положение. Таким образом, в нэмэгетинское время позднего мела в Монголии существовало не менее двух родов завропод – нэмэгтозавр и опистоцеликаудия.
Из местонахождения Тээтэ (начало позднего мела) на юго-западе Якутии описаны изолированные зубы гигантских длинношеих динозавров-завропод из группы макронарий, живших в высоких широтах (северная палеоширота 62°). Это самая северная находка завропод в мире. Находка зуба детеныша свидетельствует, что завроподы и другие динозавры не просто приходили н...
Необычным хвостом отличался описанный из позднего мела Монголии новый завроподный динозавр – абдараинур (Abdarainurus barsboldi Averianov et Lopatin, 2020). Собственно, только по серии из девяти хвостовых позвонков (восьми передних и одного среднего) он и известен, и его родовое название означает «хвост из Абдараин-Нуру» (русская транскрипция место...
Изучение двух хорошо сохранившихся зубов из коллекции Палеонтологического института им. А.А. Борисяка РАН (сборы А.К. Рождественского, 1957 г.) позволило уточнить таксономическую принадлежность завропод из местонахождения Шах-Шах в Северо-Восточном Приаралье (Кызылординская область, Казахстан). Вероятнее всего, они принадлежат представителю Opistho...
Описана зубная кость нижней челюсти нового хищного динозавра семейства дромеозаврид (Dromaeosauridae), найденная в 1960-х гг. в местонахождении Кансай на севере Ферганской долины (Таджикистан). Название нового динозавра – кансайгнат согдианский (Kansaignathus sogdianus), дано по месту находки («челюсть из Кансая») и по Согдиане – древней историческ...
Из верхнемеловой барунгойотской свиты местонахождения Хульсан в Монголии описан частично сохранившийся скелет нового рода и вида мелких хищных динозавров-альваресзаврид – Khulsanurus magnificus. Его название означает «великолепный хвост из Хульсана». Новый таксон отличается от других представителей семейства Alvarezsauridae комбинацией признаков, к...
Переизучение голотипа Parvicursor remotus Karhu et Rautian, 1996 позволило выявить ряд ранее не отмеченных морфологических особенностей парвикурсора и на этом основании пересмотреть его родственные связи. Оказалось, что цератоник очень слабо отличается от парвикурсора и может быть его младшим синонимом, что разрушает предложенную ранее (Agnolin et...
Новый род и вид динозавров Ondogurvel alifanovi (Alvarezsauridae, Parvicursorinae) описан по неполному скелету из верхнемеловых отложений местонахождения Нэмэгт в пустыне Гоби Южной Монголии. Родовое название «ондогурвэл» образовано от монгольских слов «ондо» (яйцо) и «гурвэл» (ящерица), что отражает предполагаемое питание мелких теропод альваресза...
1971 г. Совместной советско-монгольской палеонтологической экспедицией были обнаружены и собраны многочисленные остатки утконосых динозавров, включая почти полный скелет ювенильной особи, в сеноман-сантонской баинширэинской свите местонахождения Байшин-Цав в пустыне Гоби в Монголии. Этот скелет был помещен в экспозицию Палеонтологического музея им....
В 2018 г. российские и немецкие палеонтологи сообщили об открытии в Якутии, в динозавровом местонахождении Тээтэ, нового высокоширотного комплекса млекопитающих, предположительно самого древнего из известных (датируется началом мелового периода). Северная палеоширота, на которой обитали эти животные, оценивается в 63–70°. Изучение остатков млекопит...
Исследователи из России и Германии, изучающие позвоночных начала мелового периода из полярного местонахождения Тээтэ на юго-западе Якутии (северная палеоширота предположительно 62°), описали криохарамийю (Cryoharamiya tarda), новый род и вид млекопитающих из мезозойской группы эухарамийид, поверхностно сходных с грызунами небольших растительноядных...
Изучение богатейших материалов из местонахождения Ховур в Северной Гоби (112 экземпляров в коллекции Палеонтологического института им. А.А. Борисяка РАН) позволило получить новые данные о челюстной и зубной морфологии раннемеловых мультитуберкулят Монголии, ревизовать их систематический состав и реконструировать филогенетические связи. Наиболее нео...
Быстрая эволюция плацентарных началась после вымирания динозавров. Их более ранняя, мезозойская, история, пока недостаточно изучена. В частности, еще мало данных о древнейших плацентарных раннего мела – эпохи, когда эта группа впервые появляется в ископаемой летописи. Уникальным источником сведений о раннемеловых млекопитающих Азии является местона...
Древнейший представитель отряда зайцеобразных (Lagomorpha), описанный
в 2008 г. из Монголии Arnebolagus leporinus, имеет возраст около 55 миллионов лет. Родовое название было ему дано в честь звезды Арнеб – альфы созвездия Зайца. Оно подчеркивает древность и первостепенную важность находки для изучения эволюции группы. Изучение дополнительных матер...
Апатемииды были небольшой своеобразной группой мелких плацентарных млекопитающих палеогена Северного полушария. Они жили на деревьях и питались насекомыми-ксилофагами, которых извлекали из коры и древесины с помощью сильно удлиненных пальцев передних конечностей и мощных увеличенных резцов. Остатки апатемиид весьма редки, что в целом характерно для...
A mammalian petrosal from the Lower Cretaceous Teete locality in Yakutia (Russia) shows a prominent and complex system of venous channels in the bony wall of the pars cochlearis surrounding the straight cochlear canal. This complex venous system is distinctive and more strongly developed than in other mammalian petrosals. A bony ridge is present on...
A distal fragment of the wing metacarpal from the Lower Cretaceous Ilek Formation at the Novochernorechensk locality in Krasnoyarsk Territory, West Siberia, Russia, is nearly identical to the wing metacarpal of the dsungaripterid pterosaur Lonchognathosaurus acutirostris from the Lower Cretaceous of North-West China. It is identified here as cf. Lo...
A nearly complete and articulated dinosaur skeleton from the Upper Cretaceous (Cenomanian-Santonian) Bayinshire Formation at Bayshin Tsav locality in Gobi Desert (Mongolia) represents a juvenile hadrosauroid. This specimen is clustered with Gobihadros mongoliensis from the same formation and locality on 68% of the 304 most parsimonious trees produc...
The curved pencil-like sauropod teeth from the Bostobe Formation (Upper Cretaceous, Santonian-Campanian) of the Shakh-Shakh locality in Kazakhstan are referred to a representative of the Laurasian clade Opisthocoelicaudiidae known from the Upper Cretaceous (Campanian-Maastrichtian) of Mongolia. The teeth are slightly expanded near the apex and have...
A new alvarezsaurid theropod, Dzharaonyx eski, gen. et sp. nov., is documented by a variety of dissociated but well-preserved postcranial bones from the Upper Cretaceous (Turonian) Bissekty Formation at Dzharakuduk, Uzbekistan. It has opisthocoelous posterior dorsal vertebrae lacking infrapostzygapophyseal fossae. The posterior caudal vertebrae are...
Ondogurvel alifanovi gen. et sp. nov. (Alvarezsauridae, Parvicursorinae) is described based on a partial postcranial skeleton from the Upper Cretaceous (Campanian) Barungoyot Formation at the Nemegt locality in Mongolia. The new taxon differs from all other alvarezsaurids in having metatarsals II and IV completely fused along their contact area. Th...
Asiapator onchin gen. et sp. nov. is based on a dentary fragment from the middle Eocene (Irdinmanhan) Khaychin Formation at Khaychin Ula 3 locality, Mongolia. The preserved dentition includes di1 being replaced by i1, dp2 or p2, roots of dp4, and erupting p4. This is the first record of the Apatemyidae in Central Asia. The holotype of A. onchin pro...
The Campanian Beloe Ozero locality within the Rybushka Formation in Saratov Province, Russia, produces numerous isolated bones of sea turtles (clade Pan-Chelonioidea) identified as belonging to Protostega gigas, a giant protostegid, previously reliably known only from the Coniacian – Campanian of North America, and Pan-Chelonioidea indet. The Proto...
The new spalacotheriid “symmetrodontan” Cifellitherium suderlandicum gen. et sp. nov. from the Barremian–Aptian
of the Balve locality in northwestern Germany is the first record of spalacotheriids in Central Europe. The new taxon is
based on one lower and two upper molars. Cifellitherium is similar to Spalacotherium, but differs by smaller styloc...
A nearly complete and articulated dinosaur skeleton from the Upper Cretaceous (Cenomanian–Santonian) Bayinshire Formation at Bayshin Tsav locality in Gobi Desert (Mongolia) represents a juvenile hadrosauroid. This specimen is clustered with Gobihadros mongoliensis from the same formation and locality on 68% of the 304 most parsimonious trees produc...
The curved pencil-like sauropod teeth from the Bostobe Formation (Upper Cretaceous, Santonian–Campanian) of the Shakh-Shakh locality in Kazakhstan are referred to a representative of the Laurasian clade Opisthocoelicaudiidae known from the Upper Cretaceous (Campanian–Maastrichtian) of Mongolia. The teeth are slightly expanded near the apex and have...
Parvicursor remotus Karhu & Rautian, 1996 Karhu, A. A. & Rautian, A. S. 1996. A new family of Maniraptora (Dinosauria: Saurischia) from the Late Cretaceous of Mongolia. Paleontological Journal, 30, 583–592. [Google Scholar], based on a fragmentary skeleton from the Campanian Barungoyot Formation at Khulsan, in the Gobi Desert of Mongolia, is re-des...
The multituberculate assemblage from the Early Cretaceous Khovoor locality in Mongolia, based on the study of 112 specimens in PIN collection, includes three taxa: arginbaatarid Arginbaatar dmitrievae Trofimov, 1980 (=Monobaatar mimicus Kielan-Jaworowska et al., 1987, new synonym) and two eobaatarids, Eobaatar magnus Kielan-Jaworowska et al., 1987...
Five partially preserved mammaliaform petrosals recovered from Middle Jurassic sediments of the Berezovsk coal mine (Krasnoyarsk Krai, Russia) show similarities to other early mammaliaforms like the morganucodontan Morganucodon and the docodontan Haldanodon in having an inflated promontorium and a curved and apically inflated cochlear canal, but th...
Khulsanurus magnificus gen. et sp. nov., is described based on a partial skeleton, including cervical and caudal vertebrae, scapulocoracoids, humerus, and pubis from the Late Cretaceous (Campanian) Barungoyot Formation at Khulsan locality in Gobi Desert, Mongolia. The new taxon differs from other alvarezsaurids by a combination of characters that i...
A new dromaeosaurid theropod dinosaur, Kansaignathus sogdianus gen. et sp. nov., is described based on a dentary from the Yalovach Formation (Santonian) at Kansai locality in northern Fergana Valley (Tajikistan) collected by Paleontological Institute of the Academy of Sciences of the USSR in 1963-1964. Dentary has 12 tooth alveoli and not downturne...
A new dromaeosaurid theropod dinosaur, Kansaignathus sogdianus gen. et sp. nov., is described based on a dentary from the Yalovach Formation (Santonian) at Kansai locality in northern Fergana Valley (Tajikistan) collected by Paleontological Institute of the Academy of Sciences of the USSR in 1963–1964. Dentary has 12 tooth alveoli and not downturne...
The early ceratopsian dinosaur Psittacosaurus sibiricus from the Early Cretaceous of Western Siberia, Russia, is one of the most advanced and largest (up to 2.5 m) members of the genus. Here we present a description of ontogenetic changes in the long-bone histology of this species. Analysis of a growth series of femora demonstrates significant hist...
The Langenberg Quarry near Bad Harzburg has yielded the first Jurassic stem therian mammal of Germany, recovered from Kimmeridgian (Late Jurassic) near shore deposits of a palaeo-island within the Lower Saxony Basin of the European archipelago. The new stem therian is represented by one lower and three upper molars. Hercynodon germanicus gen. et sp...
A new specimen of the Early Cretaceous pterodactyloid pterosaur Istiodactylus latidens is described. This specimen in the Moscow Vernadsky State Geological Museum represents the associated anterior ends of the rostrum and mandibular symphysis but lacks collection data. It likely comes from the Barremian-Aptian Vectis Formation of southern England b...
Первая находка птерозавра в России была сделана в 1911 г. в верхнем мелу Пензенской области. Найденный фрагмент шейного позвонка (го
лотип Bogolubovia orientalis) относили последовательно к представителям птеранодонтид и аждархид. Новые находки в верхнем мелу Саратов
ской обл. позволили установить одновременное существование двух таксонов птероза...
Isolated stegosaurian teeth from the Early Cretaceous high-latitude (palaeolatitude estimate of N 62˚- 66.5˚) Teete locality in Yakutia (Eastern Siberia, Russia) are characterized by a labiolingually compressed, slightly asymmetrical and mesiodistally denticulated (9–14 denticles) crown, a pronounced ring-like cingulum, as well as a “complex networ...
Arnebolagus leporinus Lopatin and Averianov, 2008, known previously from a single tooth (P3) from the early Eocene Bumban Member of Naran Bulak Formation at Tsagan-Khushu locality in Mongolia, is redescribed based on additional specimens from the type locality. Phylogenetic relationships of Eocene stem lagomorphs from Asia and North America are rec...
Multituberculata from the Lower Cretaceous (Barremian–Aptian) Balve locality in Germany are represented by two taxa, the pinheirodontid Bructerodon alatus gen. et sp. nov. and the eobaatarid Cheruscodon balvensis gen. et sp. nov. Bructerodon alatus gen. et sp. nov. is known from an upper premolar (P5) and an upper molar (M1) with a distinct distoli...
A detailed study of the dentition of Arnebolagus leporinus, the oldest lagomorphs, from the basal Eocene of Mongolia (age 55 million years) allows to consider it as an ancestor of the branch of “modern lagomorphs” (eulagomorphs). Within this branch, the ancestral group of modern hares, rabbits, and pikas has separated about 35 million years ago.
Dzharatitanis kingi gen. et sp. nov. is based on an isolated anterior caudal vertebra (USNM 538127) from the Upper Cretaceous (Turonian) Bissekty Formation at Dzharakuduk, Uzbekistan. Phylogenetic analysis places the new taxon within the diplodocoid clade Rebbachisauridae. This is the first rebbachisaurid reported from Asia and one of the youngest...
An additional anterior caudal vertebra of a titanosaurian sauropod Tengrisaurus starkovi from the Lower Cretaceous (Barremian–Aptian) Murtoi Formation of Transbaikalia, Asiatic Russia, reveals new morphological details of this taxon. The new characters include dorsoventrally compressed cotyle and condyle of the centrum and a prominent ridge separat...
Fragmentary remains, including two anterior premolars, axis fragment, a phalanx, and rib fragments from the Middle Eocene (late Bartonian) Ak-Kaya locality represent the first sirenian fossils from the Paleogene of Crimea. The lower p3 shows some similarity with p3 in the dugongid Eotheroides sp. from the Eocene of North America. The axis is simila...
The Campanian Beloe Ozero locality within the Rybushka Formation in Saratov Province, Russia, is one of the richest and most diverse Upper Cretaceous pterosaur localities in Europe. It produces identifiable remains of Pteranodontidae indet. and Azhdarchidae indet., as well as bones which can be attributed to either of these groups. The pteranodonti...
Six dinosaurian taxa, Therizinosauridae indet., Ornithomimosauria indet., Dromaeosauridae indet., Titanosauridae indet., cf. Mongolosaurus sp., and Psittacosaurus sp., are identified and described from late Barremian to middle Aptian strata of the Murtoi Formation at the Mogoito locality in Buryatia, western Transbaikalia, Russia. The Mogoito verte...
