Alexander Averianov

Alexander Averianov
Russian Academy of Sciences | RAS · Zoological Institute

Doctor of Biological Sciences

About

446
Publications
123,145
Reads
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5,204
Citations
Additional affiliations
June 2010 - present
Saint Petersburg State University
Position
  • Professor (Full)
January 2009 - present
January 2007 - present
Rheinische Friedrich-Wilhelms-Universität Bonn
Education
September 1983 - June 1988
Leningrad State University
Field of study
  • Zoology

Publications

Publications (446)
Article
A theropod femur from the Late Cretaceous Kansai locality in Tajikistan (Yalovach Formation, Santonian) is referred to the family Therizinosauridae based on a combination of the following derived characters: the bone is straight in the anteroposterior plane; the femoral head is directed dorsomedially; the fem-oral neck is constricted between the fe...
Article
A theropod femur from the Late Cretaceous Kansai locality in Tajikistan (Yalovach Formation, Santonian) is referred to the family Therizinosauridae based on combination of the following derived characters: the bone is straight in anteroposterior plane; the femoral head is directed dorsomedially; the femoral neck is constricted between the femoral h...
Article
Full-text available
A mammalian petrosal from the Lower Cretaceous Teete locality in Yakutia (Russia) shows a prominent and complex system of venous channels in the bony wall of the pars cochlearis surrounding the straight cochlear canal. This complex venous system is distinctive and more strongly developed than in other mammalian petrosals. A bony ridge is present on...
Article
A distal fragment of the wing metacarpal from the Lower Cretaceous Ilek Formation at the Novochernorechensk locality in Krasnoyarsk Territory, West Siberia, Russia, is nearly identical to the wing metacarpal of the dsungaripterid pterosaur Lonchognathosaurus acutirostris from the Lower Cretaceous of North-West China. It is identified here as cf. Lo...
Article
A nearly complete and articulated dinosaur skeleton from the Upper Cretaceous (Cenomanian-Santonian) Bayinshire Formation at Bayshin Tsav locality in Gobi Desert (Mongolia) represents a juvenile hadrosauroid. This specimen is clustered with Gobihadros mongoliensis from the same formation and locality on 68% of the 304 most parsimonious trees produc...
Article
The curved pencil-like sauropod teeth from the Bostobe Formation (Upper Cretaceous, Santonian-Campanian) of the Shakh-Shakh locality in Kazakhstan are referred to a representative of the Laurasian clade Opisthocoelicaudiidae known from the Upper Cretaceous (Campanian-Maastrichtian) of Mongolia. The teeth are slightly expanded near the apex and have...
Article
Ondogurvel alifanovi gen. et sp. nov. (Alvarezsauridae, Parvicursorinae) is described based on a partial postcranial skeleton from the Upper Cretaceous (Campanian) Barungoyot Formation at the Nemegt locality in Mongolia. The new taxon differs from all other alvarezsaurids in having metatarsals II and IV completely fused along their contact area. Th...
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Asiapator onchin gen. et sp. nov. is based on a dentary fragment from the middle Eocene (Irdinmanhan) Khaychin Formation at Khaychin Ula 3 locality, Mongolia. The preserved dentition includes di1 being replaced by i1, dp2 or p2, roots of dp4, and erupting p4. This is the first record of the Apatemyidae in Central Asia. The holotype of A. onchin pro...
Article
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The Campanian Beloe Ozero locality within the Rybushka Formation in Saratov Province, Russia, produces numerous isolated bones of sea turtles (clade Pan-Chelonioidea) identified as belonging to Protostega gigas, a giant protostegid, previously reliably known only from the Coniacian – Campanian of North America, and Pan-Chelonioidea indet. The Proto...
Article
The new spalacotheriid “symmetrodontan” Cifellitherium suderlandicum gen. et sp. nov. from the Barremian–Aptian of the Balve locality in northwestern Germany is the first record of spalacotheriids in Central Europe. The new taxon is based on one lower and two upper molars. Cifellitherium is similar to Spalacotherium, but differs by smaller styloc...
Article
A nearly complete and articulated dinosaur skeleton from the Upper Cretaceous (Cenomanian–Santonian) Bayinshire Formation at Bayshin Tsav locality in Gobi Desert (Mongolia) represents a juvenile hadrosauroid. This specimen is clustered with Gobihadros mongoliensis from the same formation and locality on 68% of the 304 most parsimonious trees produc...
Article
The curved pencil-like sauropod teeth from the Bostobe Formation (Upper Cretaceous, Santonian–Campanian) of the Shakh-Shakh locality in Kazakhstan are referred to a representative of the Laurasian clade Opisthocoelicaudiidae known from the Upper Cretaceous (Campanian–Maastrichtian) of Mongolia. The teeth are slightly expanded near the apex and have...
Article
Parvicursor remotus Karhu & Rautian, 1996 Karhu, A. A. & Rautian, A. S. 1996. A new family of Maniraptora (Dinosauria: Saurischia) from the Late Cretaceous of Mongolia. Paleontological Journal, 30, 583–592. [Google Scholar], based on a fragmentary skeleton from the Campanian Barungoyot Formation at Khulsan, in the Gobi Desert of Mongolia, is re-des...
Article
The multituberculate assemblage from the Early Cretaceous Khovoor locality in Mongolia, based on the study of 112 specimens in PIN collection, includes three taxa: arginbaatarid Arginbaatar dmitrievae Trofimov, 1980 (=Monobaatar mimicus Kielan-Jaworowska et al., 1987, new synonym) and two eobaatarids, Eobaatar magnus Kielan-Jaworowska et al., 1987...
Article
Five partially preserved mammaliaform petrosals recovered from Middle Jurassic sediments of the Berezovsk coal mine (Krasnoyarsk Krai, Russia) show similarities to other early mammaliaforms like the morganucodontan Morganucodon and the docodontan Haldanodon in having an inflated promontorium and a curved and apically inflated cochlear canal, but th...
Article
Khulsanurus magnificus gen. et sp. nov., is described based on a partial skeleton, including cervical and caudal vertebrae, scapulocoracoids, humerus, and pubis from the Late Cretaceous (Campanian) Barungoyot Formation at Khulsan locality in Gobi Desert, Mongolia. The new taxon differs from other alvarezsaurids by a combination of characters that i...
Article
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A new dromaeosaurid theropod dinosaur, Kansaignathus sogdianus gen. et sp. nov., is described based on a dentary from the Yalovach Formation (Santonian) at Kansai locality in northern Fergana Valley (Tajikistan) collected by Paleontological Institute of the Academy of Sciences of the USSR in 1963-1964. Dentary has 12 tooth alveoli and not downturne...
Article
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A new dromaeosaurid theropod dinosaur, Kansaignathus sogdianus gen. et sp. nov., is described based on a dentary from the Yalovach Formation (Santonian) at Kansai locality in northern Fergana Valley (Tajikistan) collected by Paleontological Institute of the Academy of Sciences of the USSR in 1963–1964. Dentary has 12 tooth alveoli and not downturne...
Article
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The early ceratopsian dinosaur Psittacosaurus sibiricus from the Early Cretaceous of Western Siberia, Russia, is one of the most advanced and largest (up to 2.5 m) members of the genus. Here we present a description of ontogenetic changes in the long-bone histology of this species. Analysis of a growth series of femora demonstrates significant hist...
Article
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The Langenberg Quarry near Bad Harzburg has yielded the first Jurassic stem therian mammal of Germany, recovered from Kimmeridgian (Late Jurassic) near shore deposits of a palaeo-island within the Lower Saxony Basin of the European archipelago. The new stem therian is represented by one lower and three upper molars. Hercynodon germanicus gen. et sp...
Article
A new specimen of the Early Cretaceous pterodactyloid pterosaur Istiodactylus latidens is described. This specimen in the Moscow Vernadsky State Geological Museum represents the associated anterior ends of the rostrum and mandibular symphysis but lacks collection data. It likely comes from the Barremian-Aptian Vectis Formation of southern England b...
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Первая находка птерозавра в России была сделана в 1911 г. в верхнем мелу Пензенской области. Найденный фрагмент шейного позвонка (го лотип Bogolubovia orientalis) относили последовательно к представителям птеранодонтид и аждархид. Новые находки в верхнем мелу Саратов ской обл. позволили установить одновременное существование двух таксонов птероза...
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Isolated stegosaurian teeth from the Early Cretaceous high-latitude (palaeolatitude estimate of N 62˚- 66.5˚) Teete locality in Yakutia (Eastern Siberia, Russia) are characterized by a labiolingually compressed, slightly asymmetrical and mesiodistally denticulated (9–14 denticles) crown, a pronounced ring-like cingulum, as well as a “complex networ...
Article
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Arnebolagus leporinus Lopatin and Averianov, 2008, known previously from a single tooth (P3) from the early Eocene Bumban Member of Naran Bulak Formation at Tsagan-Khushu locality in Mongolia, is redescribed based on additional specimens from the type locality. Phylogenetic relationships of Eocene stem lagomorphs from Asia and North America are rec...
Article
Multituberculata from the Lower Cretaceous (Barremian–Aptian) Balve locality in Germany are represented by two taxa, the pinheirodontid Bructerodon alatus gen. et sp. nov. and the eobaatarid Cheruscodon balvensis gen. et sp. nov. Bructerodon alatus gen. et sp. nov. is known from an upper premolar (P5) and an upper molar (M1) with a distinct distoli...
Article
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A detailed study of the dentition of Arnebolagus leporinus, the oldest lagomorphs, from the basal Eocene of Mongolia (age 55 million years) allows to consider it as an ancestor of the branch of “modern lagomorphs” (eulagomorphs). Within this branch, the ancestral group of modern hares, rabbits, and pikas has separated about 35 million years ago.
Article
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Dzharatitanis kingi gen. et sp. nov. is based on an isolated anterior caudal vertebra (USNM 538127) from the Upper Cretaceous (Turonian) Bissekty Formation at Dzharakuduk, Uzbekistan. Phylogenetic analysis places the new taxon within the diplodocoid clade Rebbachisauridae. This is the first rebbachisaurid reported from Asia and one of the youngest...
Preprint
An additional anterior caudal vertebra of a titanosaurian sauropod Tengrisaurus starkovi from the Lower Cretaceous (Barremian–Aptian) Murtoi Formation of Transbaikalia, Asiatic Russia, reveals new morphological details of this taxon. The new characters include dorsoventrally compressed cotyle and condyle of the centrum and a prominent ridge separat...
Article
Fragmentary remains, including two anterior premolars, axis fragment, a phalanx, and rib fragments from the Middle Eocene (late Bartonian) Ak-Kaya locality represent the first sirenian fossils from the Paleogene of Crimea. The lower p3 shows some similarity with p3 in the dugongid Eotheroides sp. from the Eocene of North America. The axis is simila...
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The Campanian Beloe Ozero locality within the Rybushka Formation in Saratov Province, Russia, is one of the richest and most diverse Upper Cretaceous pterosaur localities in Europe. It produces identifiable remains of Pteranodontidae indet. and Azhdarchidae indet., as well as bones which can be attributed to either of these groups. The pteranodonti...
Article
Six dinosaurian taxa, Therizinosauridae indet., Ornithomimosauria indet., Dromaeosauridae indet., Titanosauridae indet., cf. Mongolosaurus sp., and Psittacosaurus sp., are identified and described from late Barremian to middle Aptian strata of the Murtoi Formation at the Mogoito locality in Buryatia, western Transbaikalia, Russia. The Mogoito verte...
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The first finding of the docodontan (a lower molar of Sibirotherium sp.) from the Bol'shoi Kemchug 3 locality (Ilek Formation, Lower Cretaceous) in Krasnoyarsk Territory is described. Sibirotherium rossicum, one of the youngest docodontans in the geological record, is known from the Ilek Formation of the Shestakovo 1 locality (Kemerovo Region). Pre...
Article
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The first finding of the docodontan in the Bol’shoi Kemchug 3 locality (Ilek Formation, Lower Cretaceous) in the Krasnoyarsk Territory – the lower molar of Sibirotherium sp. Sibirotherium rossicum, one of the youngest docodontans in the geological record, is known from the Ilek Formation of the Shestakovo 1 locality (Kemerovo Region). Previously, o...
Chapter
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Eutheria includes one of three major clades of mammals, the extant members of which are referred to as placentals. With more than 5000 extinct and extant genera, including 1220 extant (living) genera and over 6000 extant species arrayed in 19 extant orders, placentals (extant eutherians) are the most taxonomically diverse of the three branches of e...
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To maintain stability and universality of the nomenclature we propose the generic name Riabininohadros gen. nov. as new to meet the conditions of availability and validity of the ICZN Code.
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We describe a new euharamiyidan taxon Cryoharamiya tarda, gen. et sp. nov. from the Lower Cretaceous of Yakutia, East Siberia, Russia. The Early Cretaceous Cryoharamiya is the youngest representative of Euharamiyida in the fossil record. This new taxon extends significantly the stratigraphic and geographic ranges for Euharamiyida.
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We describe in detail three braincases of the ankylosaur Bissektipelta archibaldi from the Late Cretaceous (Turonian) of Uzbekistan with the aid of computed tomography, segmentation, and 3D modeling. Bissektipelta archibaldi is confirmed as a valid taxon and attributed to Ankylosaurinae based on the results of a phylogenetic analysis. The topograph...
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Tashtykia primaeva gen. et sp. nov. and Tagaria antiqua gen. et sp. nov. (Multituberculata incertae sedis) are described based on isolated teeth from the Middle Jurassic (Bathonian) Itat Formation from the Berezovsk coal mine in Krasnoyarsk Territory, Western Siberia, Russia. Tashtykia primaeva gen. et sp. nov. is characterized by a P5 with three r...
Article
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The pterodactyloid family Lonchodectidae includes three genera, Lonchodectes Hooley, 1914, Lonchodraco Rodrigues et Kellner, 2013, and Ikrandraco Wang et al., 2014, and four species, Lonchodectes compressirostris (Owen, 1851), Lonchodraco giganteus (Bowerbank, 1846), Ikrandraco avatar Wang et al., 2014, and Ikrandraco machaerorhynchus (Seeley, 1870...
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Cladistic analysis of distribution of 18 morphological characters in Proteles, recent Hyaenidae and some other recent and extinct Aeluroidae has been performed. The monophyletic groups Protelidae (Proteles and Dinocrocuta) and Hyaenidae (Parahyaena, Hyaena and Crocuta} are distinguished. In the family Protelidae two subfamilies Protelinae (Proteles...
Article
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Salamanders (Caudata) are one of the three modern groups of amphibians known from the Middle Jurassic. The early stages of evolution of these amphibians are still poorly known, especially for stem taxa of Jurassic age. A new small-sized stem salamander, Egoria malashichevi gen. et sp. nov., from the Middle Jurassic (Bathonian) Itat Formation of the...
Article
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A new sauropod, Abdarainurus barsboldi gen. et sp. nov., is described based on several anterior and one middle caudal vertebrae and chevrons from the Late Cretaceous Alagteeg Formation at Abdrant Nuru, northern Gobi Desert, Mongolia. The new taxon is characterized by 16 unique or rare characters of the caudal skeleton, including opisthocoelous cent...
Article
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An upper “triconodont” molar from the Late Jurassic (late Kimmeridgian) of the Langenberg Quarry in northern Germany is attributed to Storchodon cingulatus gen. et sp. nov. of Morganucodonta. The molar is characterized by continuous lingual and buccal cingula, and a relatively large, buccally-shifted cusp D which is not integrated in the buccal cin...
Article
Kazachostylops occidentalis Nesov, 1987b, based on partial maxilla and dentary from the upper Paleocene Zhylga locality in South Kazakhstan, is redescribed. A new phylogenetic hypothesis of Arctostylopida is proposed based on phylogenetic analysis of 26 characters and 17 taxa. Kazachostylops is recovered as a sister taxon to the Arctostylopinae, th...
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Isolated sauropod teeth from the Early Cretaceous Teete locality in Yakutia (Eastern Siberia, Russia) are the only evidence that sauropods lived in high latitudes (palaeolatitude estimate of N 62°) in the Northern Hemisphere. The spatulate broad tooth crowns of adult individuals lack marginal denticles while these are present in a juvenile tooth. T...
Article
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Three new euharamiyidan taxa are described based on isolated teeth from the Middle Jurassic (Bathonian) Itat Formation of the Berezovsk coal mine in Krasnoyarsk Territory, Western Siberia, Russia. The newly erected Sharypovoia, gen. nov., is referred to the Shenshouidae based on a premolariform p4, in which the main cusp and basined heel are surrou...
Article
Three sauropod middle caudal vertebrae are described from the three different localities within the Lower Cretaceous (Barremian-Aptian) Ilek Formation in Krasnoyarsk Territory, Western Siberia, Russia. All vertebrae are strongly procoelous and can be referred to Lithostrotia indet. LMCCE 005/40 from Bol'shoi Kemchug 3 locality has a ventral groove...
Article
New postcranial bones (cervical and caudal vertebrae, chevron, fibula) of the proceratosaurid tyrannosauroid Kileskus aristotocus from the Middle Jurassic (Bathonian) Itat Formation at Berezovsk coal mine, Krasnoyarsk Territory, Western Siberia, Russia, reveal three possible autapomorphies of Kileskus: a deep anterior recess between the neural cana...
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Two new localities of Mesozoic mammals have been discovered: Bol'shoi Ilek and the Berezovaya River (Russia, Krasnoyarsk Territory, Lower Cretaceous, Ilek Formation). Bol'shoi Ilek locality has yielded an edentulous fragment of the maxillary of Docodonta indet. A fragment of dentary without teeth attributed to Mammalia indet. (presumably eutriconod...
Article
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Two new localities of Mesozoic mammals have been discovered: Bol’shoi Ilek and Berezovaya River (Russia, Krasnoyarsk Territory, Lower Cretaceous, Ilek Formation). Bol’shoi Ilek locality yields an edentulous fragment of the maxillary of Docodonta indet. A fragment of dentary without teeth attributed to Mammalia indet. (presumably eutriconodontan or...
Article
The Late Jurassic (late Kimmeridgian) multituberculate assemblage from the Langenberg Quarry in Northern Germany includes Paulchoffatiidae indet., the pinheirodontid Teutonodon langenbergensis, and Cimbriodon multituberculatus gen. et sp. nov. of the paulchoffatiid line. A fragmentary m1 that was formerly attributed to Eobaataridae indet. is now as...
Article
This paper describes fragmentary turtle material (isolated basisphenoid, shell bones, humerus and ilium) from the Lower Cretaceous (Berriasian–Barremian) Teete locality in Yakutia (Eastern Siberia, Russia). All the material is attributed to one taxon of basal turtles (Mesochelydia indet.), based on the morphology of the basisphenoid, shell bones an...
Article
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A new study of the holotype of Riabininohadros weberae from the Upper Cretaceous (upper Maastrichtian) of Crimea (Besh Kosh) revealed previously unknown fragments of femur, astragalus, and calcaneus. This taxon is characterized by a complex of primitive characters found in Iguanodontia and basal Ornithischia and its phylogenetic position can be def...
Article
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A new study of the holotype of Riabininohadros weberae from the Upper Cretaceous (upper Maas-trichtian) of Crimea (Besh-Kosh) reveals previously unknown fragments of femur, astragalus, and calcaneus. This taxon is characterized by a complex of primitive characters found in Iguanodontia and basal Ornithischia and its phylogenetic position can be def...
Article
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A sample of 136 isolated theropod teeth from nine vertebrate localities within the Lower Cretaceous (Barremian-Aptian) Ilek Formation in West Siberia, Russia, can be separated into five dental morphotypes referred to five or six theropod taxa based on morphological characters. The Morphotype A includes small to large lateral teeth with relatively l...
Article
Richardoestesia is an enigmatic theropod dinosaur originally described on the basis of a pair of dentaries (holotype of R. gilmorei Currie et al., 1990) and isolated distinctive teeth, which are abundant in the Upper Cretaceous of North America. Richardoestesia asiatica (Nesov, 1995) is represented by numerous isolated teeth from the Upper Cretaceo...
Article
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Currently, there are two sauropod taxa known from the Upper Cretaceous (Maastrichtian) Nemegt Formation of Gobi Desert, Mongolia: Nemegtosaurus from the Nemegt locality and Opisthocoelicaudia from the Altan Uul IV locality. Both taxa are represented by not overlapping elements (skull and partial postcranial skeleton respectively), which arises ques...
Article
Full-text available
Currently, there are two sauropod taxa known from the Upper Cretaceous (Maastrichtian) Nemegt Formation of Gobi Desert, Mongolia: Nemegtosaurus from the Nemegt locality and Opisthocoelicaudia from the Altan Uul IV locality. Both taxa are represented by not overlapping elements (skull and partial postcranial skeleton respectively), which arises ques...
Article
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Ornithopod dinosaur Riabininohadros weberae from the Upper Cretaceous (upper Maastrichtian) of the Crimean Peninsula (Besh-Kosh) shows a set of primitive characters of hind limbs observed in iguanodontids and basal ornithischians and is referred to as Styracosterna indet. The second dinosaur specimen from Crimea (Aleshino) is a fragmentary skeleton...
Article
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Isolated sauropod teeth and caudal vertebrae from the Middle Jurassic Itat Formation of the Berezovsk coal mine, Krasnoyarsk Territory, Russia, are attributed to the endemic Asiatic eusauropod family Mamenchisauridae based mostly on phenetic similarity. The few large anterior teeth lack marginal denticles. In small, possible juvenile posterior teet...
Article
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Volgatitan simbirskiensis, gen. et sp. nov., is described based on a series of anteri- or and middle caudal vertebrae from a single individual discovered in the Lower Cretaceous (upper Hauterivian, Speetoniceras versicolor ammonite Zone) marine deposits at Slantsevy Rudnik vertebrate locality near Ulyanovsk City, Russia. The new taxon is characteri...
Article
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Reexamination of the holotype of Riabininohadros weberae from the Upper Cretaceous (upper Maastrichtian) of the Crimean Peninsula (Besh-Kosh) allowed determination of previously unknown elements of the femur, astragalus, and calcaneus. This taxon shows a set of primitive characters observed in iguanodontids and basal ornithischians and is referred...