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Introduction
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February 1996 - December 2020
February 1993 - January 1996
Education
August 1991 - December 1993
March 1989 - July 1991
March 1982 - December 1986
Publications
Publications (254)
The first cladistic analysis of the Tricommatinae is presented here based on a matrix with 88 terminals and 117 characters. The subfamily, hitherto with 28 genera and 53 species, is not recovered as a clade, instead it is composed of two nonsister clades, corresponding to the two main physiognomies of Tricommatinae sensu lato: (1) most of the diver...
For the first time a hypothesis of homology is proposed for the macrosetae which compose the armature of the distal truncus penis of the Gonyleptoidea. Previous attempts to name them in the literature have not been continued because they started by referring to macrosetae based on their position, which may change widely, and all of them were very n...
The cosmetids are conspicuous harvestmen, remarkably diverse in size, shape and colour. However, the effectiveness of all these morphological traits for diagnosing groups is scarcely explored in the literature. Since the early 20th century, the family Cosmetidae has been divided into two subfamilies, Cosmetinae and Discosomaticinae, and there has b...
This study presents a comprehensive survey of the extant diversity of harvestmen (Arachnida: Opiliones), a pivotal taxon
for biogeography and conservation studies, across Earth's geopolitical entities (here treated as "countries"). The rationale
for recognizing more than 200 of these countries lato sensu worldwide is provided. The four most diverse...
Background
K92 is a clade of Gonyleptidae harvestmen of which the greatest diversity lies within the large non-monophyletic subfamily Gonyleptinae. K92 formally includes five subfamilies: Caelopyginae Sørensen, 1884, Gonyleptinae Sundevall, 1833, Hernandariinae Sørensen, 1884, Progonyleptoidellinae Soares & Soares, 1985 and Sodreaninae Soares & Soa...
Knowledge of the family Assamiidae in the Eastern Himalayas is primarily concentrated from Nepal, while Bhutan, Xizang, and northeastern India remain much less studied. Herein, a new genus, Linzhiassamiagen. nov., is described from Xizang, China, along with two new species, Linzhiassamia medogensissp. nov. and Linzhiassamia zayuensissp. nov. These...
The limited temporal completeness and taxonomic accuracy of species lists, made available in a traditional manner in scientific publications, has always represented a problem. These lists are invariably limited to a few taxonomic groups and do not represent up-to-date knowledge of all species and classifications. In this context, the Brazilian mega...
The limited temporal completeness and taxonomic accuracy of species lists, made available in a traditional manner in scientific publications, has always represented a problem. These lists are invariably limited to a few taxonomic groups and do not represent up-to-date knowledge of all species and classifications. In this context, the Brazilian mega...
The limited temporal completeness and taxonomic accuracy of species lists, made available in a traditional manner in scientific publications, has always represented a problem. These lists are invariably limited to a few taxonomic groups and do not represent up-to-date knowledge of all species and classifications. In this context, the Brazilian mega...
Despite the significant amount of systematic research on Gonyleptidae since its original description almost 200 years ago by Sundevall in 1833, there are still unresolved phylogenetic relationships that require further investigation. Current knowledge of the unranked clade DRMN (Gonyleptidae), consisting of Discocyrtus Holmberg, 1878 sensu stricto,...
The limited temporal completeness and taxonomic accuracy of species lists, made available in a traditional manner in scientific publications, has always represented a problem. These lists are invariably limited to a few taxonomic groups and do not represent up-to-date knowledge of all species and classification. In this context, the B Brazilian meg...
In this study, the understanding of the harvestman genus Carlotta Roewer, 1943, is expanded. Carlotta currently comprises two species found in central Espírito Santo state. The descriptions of the existing Carlotta species have been supplemented, including genital morphology. Four new species are herein described-Carlotta beckeri sp. nov., Carlotta...
The reduced temporal validity of species lists made available in a traditional manner in scientific publications has always represented a problem. These lists are invariably limited to a few taxonomic groups and do not represent the up-to-date knowledge of all species and classifications. The Catálogo Taxonômico da Fauna do Brasil (CTFB), made publ...
Georeferenced localities for Opiliones recorded from Paraná. Copyright notice: This dataset is made available under the Open Database License (http://opendatacommons.org/ licenses/odbl/1.0/). The Open Database License (ODbL) is a license agreement intended to allow users to freely share, modify, and use this Dataset while maintaining this same free...
The knowledge of the diversity and distribution of Opiliones in Brazil can be considered suitable. However, while the harvestmen fauna of the southern Atlantic Forest is also well-known, several gaps remain and they are still far from complete, notably those in Paraná. Thus, investigations on regional assemblages focused on primary data are critica...
Catálogo Taxonômico da Fauna do Brasil: se ng the baseline knowledge on the
animal diversity in Brazil
The gonyleptids are one of the most diverse groups of harvestmen, exhibiting a vast range of sizes, shapes and behaviors. However, its internal classification, earlier based on meristic characters, particularly in larger groups such as Pachylinae and Discocyrtus, has been called into question by new proposals that consider morphological and molecul...
The genus Neocranaus Roewer, 1913 is revisited, its composition is expanded from two to five species and a new generic diagnosis is presented. Neocranaus albiconspersus Roewer, 1913, type species of the genus, is redescribed. The genus Tolimaius Roewer, 1915 syn. nov. is considered as a junior subjective synonym of Neocranaus, its sole member being...
This study, based on the discovery and examination of male specimens of Liomma laeve Roewer, 1959, proposes the reclassification of the monotypic genus Liomma Roewer, 1959. Currently placed in Grassatores incertae sedis, we hereby transfer it to the Nomoclastidae family. In this work, we present a reevaluation of the genus and a comprehensive redes...
The former subfamily Prostygninae was recently elevated to family rank, and its phylogenetic relationships were investigated based on molecular data. In this study, we provide a revised morphological diagnosis for the family, focusing on characters from the exomorphology and male genital morphology. Morphological data supporting their inclusion in...
The systematics of gonyleptid harvestmen has recently been undergoing critical changes, especially regarding the
Pachylinae and the K92 clade. In a recent cladistic analysis of the latter, the authors refrained from describing some of
the detected new species to minimize the bulk of the alpha taxonomy in the manuscript. Herein, as a supplement and...
The American species of Cosmetidae, in spite of being clearly distinguished in the older literature, were mixed-up by Roewer.
A small group of species historically included in the genus Cynorta C.L. Koch, 1839 has been identified and described in this study. Based on a phylogenetic analysis using morphological data, we propose that these species be recognized as a non-monotypic genus, for which the available name Abria Sørensen, 1932 is used. Abria is newly diagnosed and...
A new genus and species of Neotropical Grassatores, Hevelia crucis gen. et sp. nov., is described and illustrated
based on material of both sexes from Colombia. After comparison with the greater groups in Zalmoxoidea, especially the
putative early derivative families Guasiniidae and Icaleptidae, this new taxon is considered a Zalmoxoidea incertae s...
The genus Lacronia Strand, 1942 is herein revised, and a maximum parsimony phylogenetic
analysis of morphological characters (30 taxa, 115 characters, 2962 scorings) is performed to test its
monophyly. As a result, Lacronia is herein made monophyletic by means of the inclusion of Discocyrtus
boraceae B. Soares, 1942, Discocyrtus niger Mello-Leitão,...
Fortia, gen. nov., a new genus of Stygnidae with two new Colombian species, is diagnosed and described. Two possibly sympatric species Fortia jedi, sp. nov., and Fortia sith, sp. nov. (both from the Sierra Nevada de Santa Marta, Magdalena, Colombia), are described and illustrated. Relationships of the new genus are discussed.
Rhaucoides Roewer, 1912 is herein revisited and re-diagnosed based on somatic and genital morphology,
including the differences of both expanded and unexpanded glans complex shown for the first time in Cosmetidae. The
genus is characterized mainly by abdominal ornamentation, coloration and by having the two basalmost tarsomeres of leg
I fused. The...
The holotype of Avima leucobunus Roewer 1949, type-species of Avima, is studied, and its genital morphology is described for the first time. The Avima albiornata species group is here proposed to include three species, i.e. Avima albidecorata (Šilhavý, 1979), A. albiornata (Goodnight et Goodnight, 1947) and A. intermedia (Goodnight et Goodnight, 19...
After the type species of Paradiscocyrtus Mello-Leitão, 1927 (Paradiscocyrtus neglectus Mello-Leitão,
1927) was recognized and redescribed, the difference between it and the other two species of the
genus proved to be remarkable. Herein, a maximum parsimony phylogenetic analysis with morphological
characters (2519 scorings; 28 taxa; 105 characters)...
A description of the pattern of structure and organization of the penial macrosetae was recently put forward for Gonyleptoidea Sundevall, 1833 and demonstrated to occur also in other families of Grassatores Kury, 2002, such as Assamiidae Srensen, 1884, Epedanidae Srensen, 1886 and Pyramidopidae Sharma, Prieto & Giribet, 2011. This set of homology h...
The monotypic genus Bunostigma Mello-Leitão, 1935 is herein revisited. A detailed redescription of its type-species, Bunostigma singulare Mello-Leitão, 1935 from Rio de Janeiro state, eastern Brazil, is herein presented, providing in depth knowledge of this genus. Bunostigma is most closely related to Cryptogeobius Mello-Leitão, 1935, and as this g...
Twelve taxonomic acts are proposed in Gonyleptoidea. In Agoristenidae, 2 specific synonymies are proposed in Avima Roewer and Ocoita González-Sponga. Medellinia bordoni Soares and Avram syn. nov. is transferred from Holocranaus Roewer (Cranaidae) to Manaosbiidae attr. nov. and considered a junior subjective synonym of Rhopalocranaus bordoni Šilhavý...
During a study of Discocyrtus Holmberg, 1878 and related genera, several species of alleged Pachylinae, mostly Discocyrtus, from southern Brazil were suspected of belonging in Roeweriinae. Herein, a maximum parsimony phylogenetic analysis of morphological characters was performed to test this hypothesis. Accordingly, the following taxonomic changes...
Following the results of a maximum parsimony cladistic analysis of 30 terminal taxa against 117 morphologic characters (2978 scorings), the new genus Krateromaspis gen. nov. is described in Neopachylinae. This new genus accommodates three species from Argentina, Brazil and Paraguay, formerly included in Discocyrtus Holmberg, 1878: Krateromaspis ant...
Cosmetus Perty, 1833 is the type genus of the most diverse subfamily of Laniatores, Cosmetinae. The genus currently includes 22 valid species with a disconnected distribution, including coastal forests of NE Brazil, dry forests of the Ecuadorian Pacific, Chocó-Darien moist forests and the Venezuelan Andes. The morphology among species in the genus...
The English counterpart of Cuvier's Le Règne Animal, organized by Edward Griffith in several volumes, contained not only a translation of the French original, but also new material added by invited collaborators. The part on Arachnida, with new material contributed by George R. Gray, contained descriptions of five species of Opiliones, one of them...
The genus Spinopilar Mello-Leitão, 1940 currently has seven species distributed in eastern Brazil, Argentina and
Paraguay. Three species are hitherto known from Rio de Janeiro state (RJ), in the Brazilian Atlantic Forest. In this paper,
five species from Rio de Janeiro state (RJ) are studied as follows: (1) Spinopilar armatus Mello-Leitão, 1940, th...
Some taxonomic changes are made in Amazonian Stygnidae, based on reinterpretation of generic boundaries: (1) the hitherto monotypic genus Jime Pinto-da-Rocha & Tourinho, 2012 is newly recorded from Venezuela: Jime praecursor spec. nov. (Tobogán de la Selva, Amazonas state) is described and a new generic diagnosis is provided. The genital chaetotaxy...
A study of several Argentinean/southern Brazilian nominal species currently included in the genus Discocyrtus Holmberg, 1878 was conducted. We restore Opisthoplatus Holmberg, 1878 from synonymy of Discocyrtus after almost 140 years to include three species that are redescribed here. Opisthoplatus is referred to the recently described subfamily Neop...
Kury A. B., Mendes A. C., Cardoso L., Kury M. S., Granado A. de A., Giribet G., Cruz-López J. A., Longhorn S. J. (2021). WCO: World Catalogue of Opiliones (version 2021-03-23). In: Catalogue of Life, et al. (2021). Species 2000 & ITIS Catalogue of Life, 2021-04-05. Digital resource at www.catalogueoflife.org. Species 2000: Naturalis, Leiden, the Ne...
The sixth species of Gonyleptellus is described from the mountains of Nova Friburgo in Rio de Janeiro State, south-eastern Brazil. Gonyleptellus camelopardalis sp. nov. more closely resembles G. pustulatus and G. angeloi due to the proapical apophysis curved backwards with a secondary branch, and more specifically the latter because of the reduced...
The present paper describes two new species of Plistobunus Pocock, 1903, from China: Plistobunus gracilipenis sp. nov. from Yunnan Province, and P. weiguang sp. nov. from Guangxi Province. Their discovery greatly expands the distribution of this genus to the northwest and further inland in China.
The "World Catalogue of Opiliones" (WCO) is a collaborative effort to comprehensively index the Earth's species of harvestmen. This paper announces one component of the WCO, "WCO-Lite" a website available at https://wcolite.com/. WCO-Lite provides a graphic user interface for a second component of the WCO, "Opiliones of the World", a database on th...
This is minor version 1.2 of the WCO-Lite website, containing one more layer of this nomenclatural/taxonomic endeavor.
Version 1.2 contains bibliographic references for taxa nomina (exhaustive up to suprageneric taxa) and features a new author in the project team.
A morphological maximum parsimony analysis (2273 scorings; 25 taxa; 107 characters) is performed here with eight species of Discocyrtus Holmberg, 1878 to test their monophyly. According to the results obtained herein, only the four species Discocyrtus crenulatus Roewer, 1913, Discocyrtus flavigranulatus B. Soares, 1944 and Discocyrtus moraesianus M...
WCO-Lite (with the word “light” restyled as “lite”) is a slenderer, more agile, online version of WCO, aiming to be a public-ready, authoritative reference for taxonomy of all subtaxa of harvestmen worldwide. WCO-Lite is being developed using an integrated web-based workbench for taxonomists called TaxonWorks (http://taxonworks.org/), created by Th...
WEBSITE:
Kury, A.B., Mendes, A.C., Cardoso, L., Kury, M.S. & Granado, A.de A. (2020) World Catalogue of Opiliones. WCO-Lite version 1.1. Online at: https://wcolite.com/.
Complete list of valid taxon nomina of harvestmen from suborder to subspecies; exhaustive taxonomic survey up to December 2018
The genus nomen Larifuga Loman, 1898 was originally published without stated grammatical gender. No possible indications of gender are provided by the specific epithets, as all of them are nouns in the genitive. This genus nomen has consistently been treated as feminine in the literature. However, as shown here, it is a masculine classical Latin wo...
In the context of making a Catalog of World Opiliones accessible online, five vexing cases of incorrect authorship, dating and presumption of misidentification are herein treated, all in the suborder Laniatores. Pachylicus gracilis comb. nov. is proposed for Cerroa gracilis Roewer, 1928.
The harvestman genus Vima Hirst, 1912 is revised and newly diagnosed. A new species from Caqueta´
(Colombia), Vima panita sp. nov., is described and illustrated. This is the second species of Vima, and it represents the first
record of the genus from Colombia expanding its range by 1,500 km to the southwest. Scanning electron micrograph
images of t...
The old genus Libitia Simon, 1879 of small Andean harvestmen is revisited. The monotypic genus Libitiella Roewer, 1947 is herein considered a junior subjective synonym of Libitia. Accordingly, Libitiella bipunctata (Sørensen, 1932) is restored to the combination Libitia bipunctata. The species Libitia cordata and Libitia bipunctata comb. nov. are r...
The species Brachylibitia ectroxantha was described by Mello-Leitão in 1941 and since then it was never again reported or studied. Its type is certainly lost and his identity was until now a mystery; it is currently considered as species inquirenda and transferred to Eulibitia in a recent revision of the genus. Herein, we designate a neotype and pr...
Hyporheic zone is an active ecotone constituted by the interstitial spaces between the particles of the riverbed. The use of morphological traits can be useful to detect organisms distribution patterns in these areas. Information concerning hyporheic communities are scarce as well on chironomid vertical distribution. Fauna samples from the hyporhei...
Usatama gen. nov. is described to include a new species of Kimulidae from Colombia. This is the first record of the family from the country. The new genus presents a weakly armed ocularium, feeble sexual dimorphism, and unique genital structure with three parallel horseshoe-shaped girdles in pars distalis and absence of cochleariform macrosetae.
The family nomen Sironidae, widely attributed to Simon (1879), but recently stated to have been made available 40 years before by C. L. Koch (1839), is now shown to be even older, dating of Leach (1816), in a publication never acknowledged in the arachnological literature.
The genus Gonyleptellus Roewer, 1930 is revised and reordered; a new species Gonyleptellus angeloi sp. nov. is described from Nova Friburgo, Rio de Janeiro, three additional species are transferred from other genera and one species is removed, resulting in five species in this genus. The following nomenclatural acts are proposed herein: Gonyleptes...
IS19035 Accepted 20 June 2019
The laniatorean family Phalangodidae has been largely reduced to a core of Holarctic species, however many taxa were never formally transferred to other families. We examined a group of harvestmen related to Beloniscus Thorell, 1891, from Southeast Asia and determined the nature of the relationships of the "Beloniscu...
The penial and tarsal morphology of Protimesius is studied and morphological data support the revalidation of Obidosus with the subsequent new combinations (20 species) and restored combinations (two species). Emended diagnoses are proposed for both genera, and two cave-dwelling new species of Protimesius are described and illustrated: P. lucifer s...
Discocyrtoidespinorum sp. nov. is described from the conifers forest in Paraná State, Southern Brazil. This is the second species of Discocyrtoides Mello-Leitão, 1923, the other being known from São Paulo State. It is remarkable for having greenish/yellowish spots in the dorsal scutum highly contrasting with the brown background, a feature rare in...
The genus Neocynorta is here revisited, resulting in the increase of its diversity from three species to 13, maintaining only the type species N. virescens from its former conformation. Two subjective synonyms of Neocynorta are proposed: Forfexia and Szczurekia, transferring their members to Neocynorta (F. marmorata, S. frigida, S. petrae and S. vi...
This work is an in depth survey of all known extant and fossil Opiliones family-series nomina from the nomenclatural viewpoint, based on a corpus of 3788 bibliographic sources and aiming to serve as a reliable reference on the taxonomy of the order. A total of 194 available family-series nomina (5 fossil, 189 extant) are listed with dates and sourc...
The generic name Paecilaema C.L. Koch, 1839 is anchored to the nomen dubium Cosmetus u-flavum Perty, 1833, and the fixation of the latter is proposed by designating a neotype. The choice was made on a potential topotype, being a common species in metropolitan Rio de Janeiro city, SE Brazil, which has been described as new five times and is currentl...
The harvestman genus Propachylus Roewer, 1913 is herein revalidated from the synonymy of Discocyrtus
Holmberg, 1878. Its type species, Propachylus singularis Roewer, 1913 has a convoluted taxonomic history, connected to
Discocyrtus fornicatus Sørensen, 1884 (currently known as Metadiscocyrtus fornicatus). The monotypic genus
Metadiscocyrtus Roewer,...
The phylogeny of the genus Roquettea Mello-Leitão, 1931 and its position in Discosomaticinae are investigated. The monophyly of an expanded Roquettea is for the first time tested and corroborated. The internal relations of Roquettea are elucidated by means of a morphological maximum parsimony cladistic analysis. Synapomorphies for the genus include...
Gonyleptes curvipes Kollar, in C.L. Koch, 1839 has taxonomic problems: it is a primary junior homonym of Gonyleptes curvipes Guérin-Méneville, 1837; the holotype of G. curvipes Kollar was cited twice by Roewer (1913) under two genera, one being a misidentification of an undescribed species and a new combination under Discocyrtus Holmberg, 1878; Koc...
The correct spelling of a subfamily name based on the genus name Leptopsalis Thorell, 1882 should be Leptopsalidinae, therefore the original incorrect spelling of the subfamily name Leptopsalinae Clouse, 2012, is herein emended to Leptopsalidinae Clouse, 2012.