Abdelouahed Lagnaoui

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Université Hassan 1er · Ecole Supérieure de l'Education et de la Formation

A well-known accident in the Chefchaouen district of Morocco has highlighted the vulnerability of the region’s valuable geoheritage to environmental contamination. To assess the potential risks and inform effective mitigation strategies, we conducted a geoenvironmental assessment study. By integrating diverse data sources, including geological ma...

A well-known accident in the Chefchaouen district of Morocco has highlighted the vulnerability of the region’s valuable geoheritage to environmental contamination. To assess the potential risks and inform effective mitigation strategies, we conducted a geoenvironmental assessment study. By integrating diverse data sources, including geological maps...

Northeastern of Morocco is known by the presence of several volcanic outcrops of a Neogene to Plio-quaternary age. Miocene magmatic activity in Ras Tarf (Al Hoceima region), Gourougou (Nador region) and Guilliz (Taourirt-Guercif region) areas consists mainly of shoshonitic andesites, highly potassic calc-alkaline and potassic calc-alkaline volcanic...

It is important to understand Martian samples and their exact habitats, including extreme conditions, to improve future exploration of Mars. The Mars laboratory of Eco Astronomy Inc., represented by the Department of Research and Innovation’s Advances in Multidisciplinary Studies Unit, has developed a standard Mars Regolith Simulant called “HAYA"—E...

Late Devonian strata of the eastern Anti-Atlas are gaining a reputation to yield three-dimensionally preserved and articulated skulls of gnathostomes including that of osteichthyans, chondrichthyans and placoderms. This articulated preservation facilitates direct comparisons, yields new anatomical information, and thus helped recognizing several ne...

This study delves into the sedimentological framework of late Paleozoic tetrapod footprints discovered in the upper Pennsylvanian (? Gzhelian) of the Oued Zat Basin. The footprints, attributed to the ichnogenera Batrachichnus and Limnopus, provide a unique window into the sedimentary dynamics of this ancient environment. Our analysis reveals that t...

Morocco is an increasingly important place for the understanding of the evolution ofearly vertebrates; particularly terrestrial, mainly due to recent discoveries of abundant and diversevertebrate ichnofossil assemblages. Since the Devonian, the vertebrate ichnological record shows a great diversity in lifestyle of the extinct vertebrate groups, thr...

In this paper, we report a new vertebrate ichnoassemblage from the mid- to late Permian red beds of the Ikakern Formation (Tourbihine Member, T2) from the Argana Basin of Morocco, to improve knowledge of paleobiology and palaeobiogeography. The ichnofossils comprise than 50 footprints organized on four trackways and assigned to the ichnogenus Hyloi...

Throughout the Silurian and Devonian, cartilaginous fish successively evolved their specialized skeletal and dental characteristics, and increasingly refined their sensory systems. The Late Devonian shark taxon Maghriboselache mohamezanei gen. et sp. n. from the eastern Anti-Atlas of Morocco is known from multiple specimens reserving most of its sk...

Supplementary materials including: Taphonomy Supplementary Table 1 (list of specimens) Description of specimens Remarks on phylogenetic analyses Taxon and Character lists Supplementary Figs. 1 to 55

Africa is an increasingly important place for the understanding of the evolution of life in marine and non-marine environment, mainly due to recent discoveries of abundant and diverse vertebrate and invertebrate ichnological assemblages. New findings have been described from several stratigraphic intervals in several countries helped to establish a...

Within the next decade, it is speculated that humans will build habitats on Mars to conduct research and make long-term settlements, including harbor life stations. The practical intention that underpins our research is to test the ability of human's thinking patterns to develop micro-business approach via space-type terrestrial tourism. One such e...

In the last decade, the numerous discoveries of the late Palaeozoic vertebrate ichnoassemblages in continental deposits of Morocco increase the importance of this place for vertebrate palaeoecology, ichnotaxonomy, biostratigraphy and trans-regional correlation. The vertebrate ichnoassemblage, described herein, comes from the uppermost part of the u...

The late Permian continental succession of the Argana Basin yields a diverse and abundant ichnological record. Thus, the Ikakern Formation became one of the most diverse ichnofaunal assemblages of the late Palaeozoic in a global scale. The Ikakern Formation is an up to 2,000 m thick succession of alluvial fan conglomerates (Ait Driss Member, T1) gr...

Like other vertebrate groups, chondrichthyans diversified during the Devonian. Although mostly known from their teeth, some conservation deposits like the northern American Cleveland Shale or the northern African Thylacocephalan Layer yield complete skeletons of early chondrichthyans. Based on several nearly complete skeletons and some 3D crania, w...

Foum Larjamme geosite from Bani Geopark project, south-east of Morocco, is characterised by the co-occurrence of geological, archaeological and historical heritage. However, the archaeological heritage is particularly more appreciated by the general public as it attests to the pre-existence of human beings. An integration analysis of these archaeol...

From Middle-Late Triassic continental deposits of the Timezgadiouine Formation (Argana Basin, Morocco), 17 lithofacies are identified, with alluvial fan, fluvial, floodplain and lacustrine associations. Invertebrate traces include Archaeonassa fossulata, Arenicolites isp., cf. Arenituba, Beaconites isp., cf. Camborygma, Cruziana problematica, Diplo...

Abstract Oblongichnus vulpesi n. isp. is herein described as amygdaloid, oval or cleft-shaped burrows with a lining of variable thickness and an oblong to quadrangular elongated shaft from the mid-Holocene of the Destacamento Río Salado Member. Valves of the solenid bivalve species Solen tehuelchus were found within the burrows, indicating that thi...

Tridactyl theropod and ornithischian dinosaur tracks and trackways from Imilchil and Isli formations (Middle–?Late Jurassic, Bajocian–?) of the central High Atlas region (Morocco) are described. The Imilchil Formationconsists of brackish marine-continental deposits, and the Isli Formation is a continental red-bed succession. Considering numerous ne...

Trace fossils occur in several strata of the Devonian and Carboniferous of the eastern Anti-Atlas, but they are still poorly documented. Here, we describe a fossil swimming trace from strata overlying the Hangenberg Black Shale (correlation largely based on lithostratigraphy; Postclymenia ammonoid genozone, ca. 370 Ma old). We discuss the systemati...

The late Ordovician glaciation gave birth to tunnel valleys and their filling in the fields that made up Northern Gondwana at that time. A valuable geoheritage value is attributed to these valleys as they constitute an important part of the Earth history evolution. This study focuses on the scientific value of the Hirnantian tunnel valleys through...

We describe the swimming trace Undichna from the latest Devonian of Morocco and interpret it as a trace of an early chondrichthyan.

The observation of tubular structures within Endurance Crater, Mars, has been reported by Joseph et al (2021a,b) who hypothesized these may be mineralized and fossilized remnants of tube worms that in the ancient and recent past flourished within lakes of water heated by thermal vents. The discovery of what may be spherical hematite in this same vi...

Newly discovered dinosaur footprints from the Folkestone Formation (Lower Greensand Group) of the Lower Cretaceous (Albian) of Folkestone, Kent in S.E. England are described. The footprints can be referred to ankylosaur, theropod and ornithopod dinosaurs as potential trackmakers. The occurrence of dinosaur footprints, along with fossil wood and oys...

Complete trackways of the Late Triassic archosaur ichnogenus Brachychirotherium are extremely rare and thus far mostly known from North America, though isolated imprints are abundant. Complete step cycles have been found on the upper and lower bedding surfaces in laminated mudstones and finegrained sandstones of the lowermost part of the Machraa Ab...

Here we present the first comprehensive ichnotaxonomic analysis of a Late Triassic playa system from the Sidi Saïd Maachou Basin, Coastal Meseta (Western Meseta, Morocco). The Late Triassic deposits consist of sediments deposited in various nonmarine environments dominated by dry red-bed facies. These deposits yielded, so far, an ichnoassemblage co...

The Messinian diatomaceous deposits are widely exposed in the Lower Chelif Basin (NW-Algeria). The upper part of the Diatomaceous Formation is intercalated by four diatomaceous beds rich in Zoophycos and Chondrites. Both these traces were produced by the members of endo-benthonic fauna adapted to nutrient exploitation within sediments. In the Diato...

Zagora Province in south eastern Morocco is characterised by its desert environment as well as its culture, people, biodiversity and geology inherited from the Anti-Atlas chain. But the value of this geodiversity is not widely recognised. Ordovician and Quaternary deposits are the most dominant outcrops in the province. Within the former, the Fezou...

Morocco is increasingly engaged in a process of protection and promotion of its Geoheritage, aligned on the strategies adopted by several countries. This commitment, reinforced by the quality of its geodiversity, has brought it to be recognized as the first African and Arab country to set up a labeled UNESCO Geopark in 2014, the M'goun Geopark. Sin...

An archosaur trackway consisting of 10 successive pes imprints has been recovered from the Late Triassic Irohalene Member of the Timezgadiouine Formation (Argana Basin, Morocco). Footprints are tetradactyl-pentadactyl, and show an elongated ‘heel’, probably enclosing the trace of digit V. The trackway lacks a manus impression, which could be due to...

The Faïdja Formation displays mixed siliciclastic-carbonate deposits distributed in three members, which are organised from bottom to top as follows: the Clayey Sandstone Faïdja Member; the Clayey Limestone Bel Aoura Member and the Sandy Claystone Douaouda Member. Sedimentological data indicate the evolution of a subsiding marine environment from t...

This paper presents a comprehensive ichnological analysis of invertebrate traces, that are possibly produced by bivalve organisms, and that have been found in the Lower and Upper Kazanian deposits of the Volga Region. The reported invertebrate ichnofauna include the ichnogenera Lockeia, Oblongichnus, Protovirgularia, Ptychoplasma, and Arenituba. Th...

This paper presents the progress of the ichnological analysis of the Lower and Upper Kazanian deposits of the Pechishchi and Sentyak sections from the Volga-Kama Basin (East-European Platform, Russian Federation). The ichnological data consists of a diverse invertebrate ichnofauna, including seven ichnogenera assigned to eight ichnotaxa, which are:...

The Al Atrous region is belonging to Tafilat area, which is the junction zone between the ranges of Ougarta in the Southeast and the Anti-Atlas in the West. It is part of the Paleozoic domain of the Eastern Anti-Atlas. Limited in the North by the Cretaceous outcrops of the sub-Atlas zone of Aoufous, in the South by those of Kem-Kem, in the East by...

The reported invertebrate ichnofossil assemblages come from the Lower and Upper Kazanian marine strata of the Marine-continental (transitional) formations of the Volga and Kama River regions, which is considered as a part of the Russian Plate (East European Platform). The distribution of the invertebrate ichnoassemblages described here is different...

Here, we provide an overview on the progress of the ichnological analysis of the Lower and Upper Kazanian (Middle Permian) deposits from Volga region. Thus, the invertebrate trace fossils provide elements for the palaeoenvironmental and palaeoecological interpretations, and give the behavioural insight of the invertebrate fauna of the study area. A...

A new dinosaur tracksite is reported from continental red beds of the Jurassic (Late Bathonian-?Callovian) Isli Formation along the northern flank of the Aït Ali ou Ikkou Syncline of the Imilchil area, Central High Atlas, Morocco. The succession was deposited in a fluvio-lacustrine environment, and contains at least fourteen track-bearing levels. T...

Trackways of Late Triassic Brachychirotherium (archosaur) are known from few specimens, mostly from North America. Complete step cycles have been found on the upper and lower bedding surface in laminated mudstone and fine sandstone of the upper Oued Oum Er Rbiaa Formation (Late Triassic). This is the first trackway of the ichnogenus in North Africa...

A new dinosaur tracksite was discovered in continental red beds of the Isli Formation (late Bathonian-?Callovian) along the northern flank of the Aït Ali ou Ikkou Syncline in the Imilchil area, Central High Atlas, Morocco. The area has an extraordinary scientific potential, as the layers, which have been deposited in a fluvio-lacustrine environment...

Trackways of Late Triassic Brachychirotherium (archosaur) are known from few specimens, mostly from North America. Complete step cycles have been found on the upper and lower bedding surface in laminated mudstone and fine sandstone of the upper Oued Oum Er Rbiaa Formation (Late Triassic). This is the first trackway of the ichnogenus in North Africa...

Trackways of Late Triassic Brachychirotherium (archosaur) are known from few specimens, mostly from North America. Complete step cycles have been found on the upper and lower bedding surface in laminated mudstone and fine sandstone of the upper Oued Oum Er Rbiaa Formation (Late Triassic). This is the first trackway of the ichnogenus in North Africa...

Middle to Late Triassic redbeds of the Timezgadiouine Formation (T4-T5, Anisian-Carnian) in the Irohalene area (Argana Basin, Western High Atlas, Morocco) consist of cyclical clastic sedimentary rocks, which are mainly composed of conglomerates, sandstones, siltstones, claystones and mudstones, that were deposited in an alluvial plain to mudflat en...

Complete trackways of the Late Triassic archosaur ichnogenus Brachychirotherium are extremely rare and thus far mostly known from North America, though isolated imprints are abundant. Complete step cycles have been found on the upper and lower bedding surfaces in laminated mudstones and finegrained sandstones of the lowermost part of the Machraa Ab...

Complete trackways of the Late Triassic archosaur ichnogenus Brachychirotherium are extremely rare and thus far mostly known from North America, though isolated imprints are abundant. Complete step cycles have been found on the upper and lower bedding surfaces in laminated mudstones and finegrained sandstones of the lowermost part of the Machraa Ab...

Complete trackways of the Late Triassic archosaur ichnogenus Brachychirotherium are extremely rare and thus far mostly known from North America, though isolated imprints are abundant. Complete step cycles have been found on the upper and lower bedding surfaces in laminated mudstones and fine-grained sandstones of the lowermost part of the Machraa A...

Complete trackways of the Late Triassic archosaur ichnogenus Brachychirotherium are extremely rare and thus far mostly known from North America, though isolated imprints are abundant. Complete step cycles have been found on the upper and lower bedding surfaces in laminated mudstones and fine-grained sandstones of the lowermost part of the Machraa A...

The folded Middle Atlas Mountains has rich geomorphologic landscapes and remarkable geological sites, little known by the general public. In this paper, we propose an approximately 60 km geological trail, including several geosites mostly staggered in the sinuous valleys of the Atchane and Guigou rivers, between the Boulemane, Skoura and El Mers. T...

A burrow previously assigned to the ichnotaxon Palaeophycus insignis is redescribed, and a new ichnotaxon Olongichnus solodukhoi n. igen. n. isp is erected for this flattened, slightly curved, multi-oriented, smooth burrow that has a thick lining with internal and external very fine-grained mucuous layers. It shows subrectangular to oblong form in...

Quadruped trackways of large pentadactyl footprints are reported from the Middle Triassic (Anisian-Ladinian) Cerro de las Cabras Formation of the Cuyo Basin, Mendoza Province, central-western Argentina. The track-bearing strata are interpreted as deposited by sheetfloods in a mixed flat where water was ponded in a playa-lake setting. The vertebrate...

Horseshoe crabs (Chelicerata, Xiphosura) from Mesozoic deposits are extremely rare in the fossil record of the African continent. Here we report new evidence of the occurrence of xiphosurans in North Africa. These are horseshoe crab traces, including the ichnogenera Kouphichnium (repichnia) and Selenichnites (fodinichnia and/or domichnia), which ha...

Forelimb posture in sauropod dinosaurs is still poorly understood. Although a laterally directed (semisupinated) manus is the plesiomorphic condition in sauropodomorphs, the sauropod track record prevailingly shows anterolateral to anterior manus orientations, suggesting a high degree of manus pronation. The ?Middle Jurassic Tafaytour tracksites de...

The rise and early evolution of Dinosauromorpha is documented by skeletal remains, beginning during the Middle Triassic (Anisian). Small lacertoid footprints from Olenekian- Anisian deposits of North America and Europe (Rotodactylus, Prorotodactylus) have also been attributed to this group, but this relationship is still debated. Mesaxonic tridacty...

An archosaur trackway consisting of 10 successive pes imprints has been recovered from the Late Triassic Irohalene Member of the Timezgadiouine Formation (Argana Basin, Morocco). Footprints are tetradactyl-pentadactyl, and show an elongated ‘heel’, probably enclosing the trace of digit V. The trackway lacks a manus impression, which could be due to...

The Middle Atlas Mountains have rich geomorphologic landscapes and remarkable geological sites, little known by the general public. In this work, we propose an approximately 60 km long geological trail, including several geosites mostly staggered in the sinuous valleys of the Atchane and Guigou rivers, between Boulemane, Skoura and El Mers. These g...

Xiphosurans (limulacea, limulids) from Mesozoic deposits are extremely rare in the fossil record of the African continent. Here we report new findings of xiphosuran traces, including the ichnogenera Kouphichnium (repichnia) and Selenichnites (fodinichnia), which have been discovered in Middle Jurassic strata of the Imilchil area (Central High Atlas...

The Late Carboniferous Souss Basin of south-central Morocco exhibits an approximately 1,800 m thick succession of fluvial and lacustrine deposits that have yielded diverse fossil remains of plants, insects, conchostracans, ostracods, jellyfish, fishes, and few tetrapod footprints. Recent exploration of ichnofossils of the Souss Basin led to the dis...

The Late Triassic fluvio-lacustrine sediments of the upper part of the Oued Oum Er-Rbiaa Formation in the Sidi Saïd Maachou Basin, Moroccan Meseta, have recently been known for a quite diverse record of fossils, including plant impressions, root traces, invertebrate and vertebrate traces as well as fish scales (Saber et al., 2007; Hminna et al., 20...

The Irohalene Member of the Timezgadiouine Formation (T5, Upper Triassic, Carnian) near Irohalene village (Argana Basin, western High Atlas) is well known for its rich vertebrate fauna, comprising both body and ichnofossils. A diverse ichnoassemblage with tetrapod footprints was described by Lagnaoui et al. (2012, 2016). Recent discoveries from thi...

This volume is dedicated to the results of the First International Congress on Continental Ichnology (ICCI-2015), which was held in the Faculty of Sciences, Choua€ıb Doukkali University, El Jadida, Morocco, from 21st to 25th April, 2015 (Saber et al. 2015; Lagnaoui and Saber 2015). The first two days were devoted to keynotes, oral presentations, po...

Most of the structural studies of the intracontinental High Atlas belt of Morocco have dealt with the central part of the belt, whose basement does not crop out. Here we study the Alpine deformation of the North Subatlas Zone, which is the part of the Western High Atlas (WHA) Paleozoic Massif that involves both Paleozoic basement units and remnants...

The late Triassic-early Jurassic volcanism of Sidi Saïd Maachou basin belongs to the costalMeseta and the Central Atlantic Magmatic Province (CAMP). The volcanic pile conformably overlies the red siltstones of Machraa Boujamaa Formation. This set includes a stack of several lava flows 40 to 80 m thick. The petrographic study shows that the textures...

The late Triassic-early Jurassic volcanism of Sidi Saïd Maachou basin belongs to the costalMeseta and the Central Atlantic Magmatic Province (CAMP). The volcanic pile conformably overlies the red siltstones of Machraa Boujamaa Formation. This set includes a stack of several lava flows 40 to 80 m thick. The petrographic study shows that the textures...

The late Triassic-early Jurassic volcanism of Sidi Saïd Maachou basin belongs to the costal Meseta and the Central Atlantic Magmatic Province (CAMP). The volcanic pile conformably overlies the red siltstones of Machraa Boujamaa Formation. This set includes a stack of several lava flows 40 to 80 m thick. The petrographic study shows that the texture...

The late Triassic-early Jurassic volcanism of Sidi Saïd Maachou basin belongs to the costalMeseta and the Central Atlantic Magmatic Province (CAMP). The volcanic pile conformably overlies the red siltstones of Machraa Boujamaa Formation. This set includes a stack of several lava flows 40 to 80 m thick. The petrographic study shows that the textures...

Footprints of Crocodylomorpha (Archosauria, Pseudosuchia) are rare in Mesozoic ichnoassemblages. However, those of small terrestrial forms (Batrachopus) occur in the Late Triassic-Early Jurassic. Crocodylopodus trackways of semi-aquatic crocodylomorphs have been found in the Jurassic-Cretaceous. We discovered tetradactyl pes (8 cm long) and associa...

Small bird-like tracks have recently been discovered at three outcrops of the Imilchil Formation (Middle Jurassic, Bajocian-Bathonian) in the Central High Atlas of Morocco. The track-bearing strata are part of a marine-continental transitional succession, the studied surfaces being sandy marls and limestones of a brackish depositional environment....

The Late Palaeozoic deposit from Morocco yielded a diverse and rich palaeontological heritage. New findings have been described from seven stratigraphic intervals in six localities: (1) Westphalian deposits of Sidi Kassem basin contain of the ichnogenera Batrachichnus, Dimetropus, Hylopus and Notalacerta, associated with invertebrate traces assigne...

La feuille d’Imi-N-Tanout de la carte géologique du Maroc au 1/50.000 est située à l’extrémité occidentale du Massif ancien du Haut Atlas occidental. Elle est séparée du Haut Atlas de Marrakech, d’ouest en est, par les feuilles d’Addouz, Azegour et Amezmiz du présent programme de cartographie géologique. La feuille d’Imi-N-Tanout présente trois dom...

ABSTRACT Abstract The Amezmiz sheet of the geological map of Morocco 1:50 000 is located in the eastern part of the western High Atlas, at its join to the Marrakech Atlas. It offers three contrasting morphological areas, with, i) in the south, a young mountainous domain corresponding to the axis of the Atlas chain, deeply cut by the Agoundis and Nf...

In the last decade, Morocco becoming a more and more important place for vertebrate and invertebrate ichnology by the numerous discoveries of Late Palaeozoic and Early Mesozoic vertebrate and invertebrate ichnassemblages in continental red-beds. The ichnoassemblages, described herein, come from uppermost part of the up to 1800 m thick red-bed Ikake...