Annales Zoologici

Online ISSN: 1734-1833
Print ISSN: 0003-4541
Publications
Cassida mishmiensis sp. nov.: (1) habitus dorsal; (2) habitus lateral; (3) head. 
Habitus dorsal: (4) Cassida mishmiensis sp. nov.; (5) Cassida dohertyi Spaeth; (6) Cassida corruptrix Spaeth; (7) Cassida nigrohumeralis Borowiec et Ghate; (8) Cassida desultrix (Spaeth); (9) Cassida sigillata (Gorham); (10) Cassida tuberculata Medvedev et Eroshkina; (11) Cassida postarcuata (Chen et Zia); (12) Cassida truncatipennis (Spaeth); (13) Cassida cherrapunjiensis Maulik; (14) Cassida manipuria Maulik. 
Cassida atrosignata sp. nov.: (15) habitus dorsal; (16) habitus lateral; (17) head. 
Three species of Cassida Linné, 1758 from NE India are described as new to the science: Cassida mishmiensis sp. nov. (Arunachal Pradesh), C. atrosignata sp. nov. (Arunachal Pradesh) and C. pacholatkoi sp. nov. (Meghalaya). Type material of Thlaspida obenbergeri Spaeth, 1928 has been studied and a lectotype designation is given here. Thlaspida obenbergeri is transferred to the genus Cassida (comb. nov.) and a following new synonym is proposed: Cassida crucipennis Borowiec, 2003 = Cassida obenbergeri (Spaeth, 1928).
 
Lochmaea crataegi group is established for four previously described species: L. crataegi (Forster, 1771), L. limbata Pic, 1898, L. setulosa (Sahlberg, 1913) and L. machulkai Roubal, 1926. The species from Asia Minor, Near East and Caucasus (L. setulosa and L. machulkai ) are revised on the base of type material. L. machulkai is removed from synonymy and the lectotype is designated. Male genitalia of L. machulkai and L. limbata are illustrated for the first time. L. limbata is newly recorded for Iran and Jordan, L. setulosa for Turkey and L. machulkai for Azerbaijan, Armenia, Gruzia, Iran and Turkey.
 
Type material, reputed to be lost, of Tenebrio cornutus Fabricius, 1775 and Tenebrio atratus Fabricius, 1775 (Coleoptera, Tenebrionidae), together with two possible syntypes of Tenebrio brunneus Fabricius, 1798 (Coleoptera, Cerambycidae, Parandrinae) and the type of Tenebrio laevis Olivier, 1795 were discovered in the collection of William Hunter (1718-1783) in the Hunterian Museum (Zoology), University of Glasgow. Lectotype and paralectotype are designated for Tauroceras cornutus (Fabricius, 1775); lectotype for Zophobas atratus (Fabricius, 1775); lectotype for Merinus laevis (Olivier, 1795).
 
The last larval stage of Heliotaurus ruficollis Fabricius, 1781 (Tenebrionidae: Alleculinae, Omophlini), a subterranean detritivore, is described in detail. The larva of H. ruficollis has the distinguishing characters of the subfamily Alleculinae and the tribe Omophlini, and it may be distinguished from the larvae of Omophlus species by the more rounded apex of tergum 9 and the fact that the ventral portion of tergum 9 is at the same level than sternum 9.
 
Hesperus rufipennis. 9–13. Mature larva, 14. 1 st instar larva. (9) Head in dorsal aspect (Na – nasale, Gl – gland); (9A) the microstructure of neck; (10) head in lateral aspect; (10A) right stemmata cluster; (11) head in ventral aspect (Ap – apotome); (11A) tentorial pit; (12) right antenna in dorsal aspect (I–IV antenal segment); (13) apical part of right antenna (Sa – sensory appendage, So – solenidia); (14) right antenna in dorsal aspect (I–IV antenal segment).  
Hesperus rufipennis. (1) Egg; (2) chorion microsculpture from outside surface (srd-short ridge); (3) chorion microsculpture from inside surface. 4-8. Mature larva. (4) habitus (dorsal aspect); (5) middle seta of pronotum; (6) lateral seta of abdominal tergite I; (7) middle seta of abdominal tergite III; (8) middle seta of abdominal segment IX.
The egg, three larval instars and pupa of Hesperus rufipennis (Gravenhorst) are described for the first time, with the illustrations of structural features provided. Differences in the morphological structure of its larval instars (L1-3 ) cover: the number of setae on profemur and protibia, number of setae on tarsungulus, chaetotaxy of abdominal tergites, the structure of antenna and urogomphi, body colour and measurements. Diagnostic characters of immature stages of H. rufipennis are listed. Modifications of the keys to known eggs, larvae and pupa of the European Staphylininae species are proposed. Some data on its biology (life cycle, duration of immature stages, ecological preferences) are also given. It is a univoltine species with summer larvae and wintering imagines. Considering its environmental preferences and biology, this rove beetle may be of essential importance as a good umbrella species and environmental state indicator as well.
 
Balaustium murorum (Hermann, 1804) is redescribed. Characteristics of active postlarval forms and first characteristics of larvae, supplemented with data on biology of the species, are provided. Female of B. murorum, from which larvae were obtained by experimental rearing, has been designated as neotype. Key characters, hitherto used by different authors in order to distinguish between members of Balaustium are discussed. Seventeen species known from larvae are presently assigned to Balaustium, at the total number of 41 members of the genus known worldwide.
 
The taxonomy of the ant subgenus Dendrolasius Ruzsky, 1912 is reviewed on the base of the investigation of types and of non-type material of several species. L. fuji is described as a new species, which includes former L. fuliginosus (Latreille, 1798) from the East Palaearctic. L. nipponensis Forel, 1912, proposed by Espadaler et al. 2001 as the replacement name for "oriental fuliginosus ", actually is a good species and the senior synonym of L.crispus Wilson, 1955; L. orientalis Karawajew, 1912 is revived from synonymy and is considered as the senior synonym of L. teranishii Wheeler, 1928; L. capitatus Kusnetzov-Ugamsky, 1928 is considered as a good species, different from L. crispus. A key to Dendrolasius wo rker s and queens from the Eastern Palaearctic is also given.
 
Two new species, namely Baetis (Baetis) mirkae sp. nov. (larva, female subimago, and male imago) and Baetis (Rhodobaetis) irenkae sp. nov. (larva and male imago), from Cyprus are described and their critical diagnostic characters illustrated. The former is a representative of the subgenus Baetis s. str. (the B. lutheri species-group) showing close relationships mainly to B. (B.) lutheri Müller-Liebenau, 1967 and B. (B.) vardarensis Ikonomov, 1962; the latter is classified within the subgenus Rhodobaetis Jacob, 2003, being related mainly to B. (R.) ilex Zimmermann, 1978. Affinities of these new species to all representatives of respective related species-group and Rhodobaetis are discussed in detail and brief notes to their biology and distribution are presented. Based on data available so far, a detailed biogeographical analysis directed mainly to chorology and faunistic origin of 8 Palaearcticrepresentatives of the B. lutheri species-group and 26 species of Rhodobaetis (incl. B. irenkae sp. nov. described below) is offered.
 
The morphology of the third larval stage of Morica hybrida Charpentier, 1825 (Tenebrionidae: Pimeliinae, Akidini) is described and illustrated. The description is based on diagnostic characters of Tenebrionidae classification on head, legs and ninth abdominal segment. The larva of Morica hybrida shows numerous affinities with previously described larvae of Pimeliinae and is clearly included within tribe Akidini. The larva of M. hybrida shows a high similarity with the larva of M. favieri, although they differed by the presence or absence of additional small spines in distal part of ninth abdominal segment, and the shape of the margin of labrum. In addition, similarities with larvae of Akis spp. corroborates the proximity of these two Mediterranean genera of Akidini.
 
The types of all species belonging to the genus Goniadera Perty, 1830 are studied. The lectotype of Melandrya repanda Fabricius, 1801, is designated. Eight new species of Goniadera are described: Goniadera ardoini sp. nov., Brasil, G. barclayi sp. nov., Brasil and Peru, G. girardi sp. nov., G. jaegeri sp. nov., G. merkli sp. nov., Brasil, G. onorei sp. nov. from Ecuador, G. peseudorepanda sp. nov. from Costa Rica and Mexico and G. floresi sp. nov. from Argentina. Goniadera simplex Fairmaire, 1889 is transferred to Aemyone Bates, 1868, but this genus is degraded to sub-genus of Goniadera. The name Goniadera (Aemyone) simplex (Fairmaire, 1889) comb. nov., is proposed. The genus Opatresthes Gebien, 1928 is studied and considered as another sub-genus of Goniadera. Goniadera tuberculifera Fairmaire, 1889 is trasferred to this genus, the new name Goniadera (Opatresthes) tuberculifera (Fairmaire, 1889) comb. nov., is proposed. A new species from Nicaragua: Goniadera (Opatresthes) maesi sp. nov. is described. A key to separate all sub-genera and species is provided. Etaceta aeneicolor Fairmaire, by monotypy is excluded from the tribu Goniaderini and its position in the tribu Lupropini sensu Ardoin (1969) is confirmed. New synonymies are recognised: Goniadera simplex Fairmaire (= Aemyone bordoni Marcuzzi, 1994) syn. nov., Aemyone cariosa Fairmaire, 1873 (= Anaedus striatipennis Pic, 1917) syn. nov. and (Aemyone silvanae Marcuzzi, 1994) syn. nov.; Aemyone crenata Champion, 1886 becoming homonymic with Goniadera crenata Perty, 1830 is changed in Goniadera (Aemyone) championi nom. nov. French La totalité des types des espèces du genre Goniadera Perty, 1830 sont étudiés. Le lectotype de Melandrya repanda Fabricius, 1801 est désigné. Huit espèces nouvelles sont décrites: Goniadera ardoini sp. nov. du Brésil, G. barclayi du Brésil et Pérou, G. girardi sp. nov., G. jaegeri sp. nov. et G. merkli sp. nov., Brésil, G. onorei sp. nov. du Equateur, pseudorepanda sp. nov. du Costa Rica et du Mexique, et G. floresi sp. nov., d'Argentine. Goniadera simplex Fairmaire, 1889 est transféré dans le genre Aemyone Bates, 1868, considéré comme un sous-genre de Goniadera, le nom Goniadera (Aemyone) simplex (Fairmaire) comb. nov. est proposée. Goniadera tuberculifera Fairmaire, 1889 est transférée dans le genre Opatresthes Gebien, 1928, qui est égalément considéré comme un sous-genre de Goniadera. Le nouveau nom Goniadera (Opatresthes) tuberculifera (Fairmaire, 1889) comb. nov. est proposé. Une nouvelle espèce du Nicaragua, Goniadera (Opatresthes) maesi sp. nov. est décrite. Une clé pour séparer tous les sous-genres et espèces est donnée. Etaceta aeneicolor Fairmaire est exclu de la tribu des Goniaderini et sa position dans celle des Lupropini sensu Ardoin (1969) est confirmée. Des synonymies nouvelles sont établies: Goniadera simplex Fairmaire (= Aemyone bordoni Marcuzzi, 1994) syn. nov., Aemyone cariosa Fairmaire, 1873 (= Anaedus striatipennis Pic, 1917) syn. nov. et (= Aemyone silvanae Marcuzzi, 1994) syn. nov Aemyone crenata Champion, 1886 tombe en homonymie avec Goniadera crenata Perty, 1830 le nom est changé, devenant Goniadera (Aemyone) championi nom. nov.
 
A review of the species of the genus Meromyza Meigen, 1830 from Inner Mongolia is provided. The following 3 species are described as new to science: Meromyza acutata,M. congruens and M. neimengensis.
 
Three new species of the genus Suillia Robineau-Desvoidy, S. danielssoni sp. nov. (Ecuador), S. huggerti sp. nov. (Ecuador), and S. steyskali sp. nov. (Colombia) are described from the Neotropical Region. S. iniens (Giglio-Tos, 1893) is newly recorded from Honduras. A key to the known species of Suillia occurring in the Neotropical Region is provided.
 
The species of the genus Opatroides designated Brullé, 1832 are studied and a key to all species is provided. Lectotypes are designed for Penthicus curtulus Fairmaire, 1892, Penthicus frater Fairmaire, 1896, Penthicus vicinus Fairmaire, 1896 and Opatroidesangulatus Baudi, 1875. This taxon is considered to be a geographical subspecies from Syria and Iran of the nominal form, O. vicinus, described from India. Opatroides punctulatus Brullé, 1832 is confirmed to be a widely distributed species, consisting of three geographicalsubspecies: the nominal form O. punctulatus Brullé, described from Greece, occupying the coast of the Mediterranean Sea; the subspecies subcylindricus Ménétriès, 1849 occupying the Anatolian region, the Middle East, the Caucasus and the Central Asianregion; the subspecies lilligi ssp. nov. in the Niger, Burkina Faso, Mauritania and Sudan. The status of Opatroides hemistictus Gebien, 1920 from Namibia and South Africa, as a species from belonging to the genus Mesomorphus Seidlitz, 1893, is confirmed. Figuresof the habitus and aedeagus for all taxa are given for the first time.
 
Dictyophara europaea (L., 1767). (10) Head, pronotum and mesonotum, dorsal view; (11) head and pronotum, lateral view; (12) head, ventral view; (13) forewing; (14) pygofer and parameres of male, ventral view; (15) genitalia of male, lateral view; (16) pygofer and anal tube of male, dorsal view; (17) aedeagus, dorsal view; (18) aedeagus, lateral view; (19) aedeagus, ventral view.
Dictyophara nekkana Matsumura, 1940. (31) Head, pronotum and mesonotum, dorsal view; (32) head and pronotum, lateral view; (33) head, ventral view; (34) forewing; (35) hind wing; (36) pygofer and parameres of male, ventral view; (37) genitalia of male, lateral view; (38) pygofer and anal tube of male, dorsal view; (39) aedeagus, dorsal view; (40) aedeagus, lateral view; (41) aedeagus, ventral view.
Dictyophara pannonica (Germar, 1830). (42) Head, pronotum and mesonotum, dorsal view; (43) head and pronotum, lateral view; (44) head, ventral view; (45) forewing; (46) pygofer and parameres of male, ventral view; (47) genitalia of male, lateral view; (48) pygofer and anal tube of male, dorsal view; (49) aedeagus, dorsal view; (50) aedeagus, lateral view; (51) aedeagus, ventral view.
The dictyopharids planthopper genus Dictyophara Germar, 1833 from China is revised and restricted in Palaearctic Region of northern China. Only four recognized Dictyophara species are: D. europaea (Linnaeus, 1767), D. koreana Matsumura, 1915, D. nekkana Matsumura, 1940 and D. pannonica (Germar, 1830), which the former two are recorded from China for the first time. One new generic and three new specific synonyms are recognized: Dictyophara Germar, 1833 = Togaphora Matsumura, 1940 syn. nov.; D. koreana Matsumura, 1915 = Centomera manchurica Kato, 1933 syn. nov. = Togaphora hokuryonis Matsumura, 1940 syn. nov.; D. nekkana Matsumura, 1940 = Dictyophara kaszabi Dlabola, 1967 syn. nov. Photographs of the adults of all known species are presented. Descriptions of the genus and its included species are provided together with structural illustrations of the male genitalia. A key and a distribution map to the four species of Dictyophara from China are provided.
 
One species of the genus Argyra Macquart, 1834 from China is described as new to science. The species of the genus from China are keyed, and geographic distributio ns are updated and discussed.
 
Edéage de Tauroceras difformipes (Pic) (variation). 59-66. Ovipositeurs. (59) T. cornutum, (60) T. barclayi sp. nov., (61) T. angulatum, (62) T. aries, (63) T. evasprecherae sp. nov. (face dorsale), (64) T. evasprecherae sp. nov. (face ventrale), (65) T. lucifer sp. nov., (66) T. noelmali sp. nov. 67-68. Habitus de Tauroceras spp. (67) T. girardi sp. nov., (68) T. mulata Zayas (d'après Zayas, 1988).
Tauroceras cornutum (F.) (1) Avant-corps, (2–3) Protibias ♂, en vision dorsale et ventrale, (4) tête du ♂ vue de face, (5) avant-corps ♀, (6) tête de la ♀ vue de face, (7) édéage (Jamaïque), (8) édéage (Cuba). 9–14. Tauroceras barclayi sp. nov. (9) tête de la ♀ vue de face, (10) édéage, (11) avant-corps ♂, (12) protibia ♂, (13) tête du ♂ vue de face, (14) avant-corps de la ♀. 15–17. T. tichyi sp. nov. (15) tête du ♂ vue de face, (16) avant-corps du ♂, (17) édéage.  
Tauroceras cornutum (F.) (1) Avant-corps, (2-3) Protibias ♂, en vision dorsale et ventrale, (4) tête du ♂ vue de face, (5) avant-corps ♀, (6) tête de la ♀ vue de face, (7) édéage (Jamaïque), (8) édéage (Cuba). 9-14. Tauroceras barclayi sp. nov. (9) tête de la ♀ vue de face, (10) édéage, (11) avant-corps ♂, (12) protibia ♂, (13) tête du ♂ vue de face, (14) avant-corps de la ♀. 15-17. T. tichyi sp. nov. (15) tête du ♂ vue de face, (16) avant-corps du ♂, (17) édéage.
The type material of the species belonging to the genus Tauroceras Hope, 1840 and Tauroceropedus Pic, 1913 is studied and all the known taxa are revised. Tenebrio cornutum Fabricius, 1775 (genus Tauroceras) is redescribed on the basis of type material discovered in the Hunterian collection (1718-1783) of the Museum of Zoology, University of Glasgow, Scotland. The strange locus typicus "Smyrna" is found not to be Izmir in Turkey, but more probably a locality in the old scotish colony of Florida. Lectotypes are designated for the following taxa: Tenebrio aries Dalman, 1823, Tenebrio angulatum Perty, 1830, Tauroceras angulatum Pic, 1913, syn. nov. and T. angulatum var. nitidus Pic, 1913, syn. nov. Six new species are described: Tauroceras evasprecherae sp. nov. from Brazil, T. girardi sp. nov. from Central America, T. barclayi sp. nov. from Jamaica, T. tichyi sp. nov. from Ecuador, T. noelmali sp. nov. from Colombia and Bolivia, T. lucifer sp. nov. from Brazil, Colombia and Ecuador. The genus Tauroceropedus Pic, 1913 syn. nov. is considered as a junior synonym of Tauroceras Hope, 1840. The formerly unknown male of Tauroceras (= Tauroceropedus) rugosicollis (Pic, 1913) comb. nov. is described from French Guyana. Figures and keys to identify all species are given. The genus Tauroceras is maintained within the tribe Centronopini in the subfamily Tenebrioninae, sensu Doyen (1980).
 
O. fulvipennis: (29A, 31) 1 st instar larva; (29, 29B, C, 30, 30A–C, 32, 32A) mature larva. (29) Prothorax (I), mesothorax (II) and metathorax (III) in dorsal aspect (Cs – coeloconic sensillum); (29) microstructure of some part of mesothorax; (29A, 29B) coeloconic sensillum; (30) prothorax (I), mesothorax (II) and metathorax (III) in ventral aspect (Cc – coxal cavity, Cr – cervicosternum, Sat – atrophied spiracle, Sn – sternite, Sp – functional spiracles); (30A) microstructure of sternite; (30B, C) microstructure of cervicosternum; (31, 32, 32A) fore leg in anterior aspect (Fe – femur, Tb – tibia, Tu – tarsungulus, 1, 2, … – codes of setae).  
Ocypus fulvipennis: (1, 1A-E) Egg; (2, 4-10) 1 st larval instar; (3) mature larva. (1) General view, (1A) microstructure, (1B-D) microstructure of equatorial band (Eb) with aeropyles (Ap); (1E) distal end with tubercle (Tk); (2) general view; (3) general view; (4-10) setae of abdominal tergites.
The paper describes and illustrates the morphology of all preimaginal stages for Ocypus fulvipennis Erichson, 1840 including a detailed account of chaetotaxy. Diagnostic characters of egg, larva and pupa of this species are given. Morphological differences between the first (L1) and mature (L3) larval instars cover: chaetotaxy of head, profemur, protibia, tarsungulus, abdominal tergites, paratergites, sternites, parasternites and urogomphi; structure of antenna, maxillae and urogomphi; microstructure of abdominal tergites, proportions of the body parts, body colour and habitus. Some data on its distribution and biology in the field and laboratory conditions are also provided. All immature stages of O. fulvipennis were compared with those of other members of this genus. Instead of size, following distinguishing characters are provided for respective stages: egg - tubercle simple, without projection and equatorial band medium in width; larvae (L3) - teeth of nasale very sharp and well marked, apotome with sharply pointed apex, segment II of labial palp about 2.4 times longer than segment III; pupa - 12 setiform projections on pronotum, antennae distinctly protruding beyond apex of middle tibia, hind legs reaching ⅔ of length of 4th morphological segment.
 
Three larval instars of Nearctic Agathidium angulare Mannerheim, 1852, are described in detail and the first and third stage larvae are figured for the first time; morphology of the stages was analyzed and compared, with special emphasis on measurements, chaetotaxy and porotaxy of head, mouthparts, thorax, abdomen, leg and urogomphi. Preliminary discussion on larval characters of the genus Agathidium is provided.
 
Landscape of habitat of Nalassus bozdagus sp. nov., Bozdag ridge, 2300 m. 
Nalassus bozdagus sp. nov., general view. (1) Male; (2) female. 
Nalassus bozdagus sp. nov. (4-5) Pronotum of males; (6-7) pronotum of females; (8) aedeagus, ventral view; (9) aedeagus, lateral view; (10) gastral spicula. 
A new species Nalassus bozdagus sp. nov. is described from the Aegean Region of Turkey (Bozdag ridge). The new species differs from all known Nalassus in the following characters: the hair spot on first and second abdominal sternites of male is absent, temple grooves are absent, exosceleton very strongly sclerotized. New records of Nalassus Mulsant, 1854 from Turkey are provided. Two species new for the fauna of Turkey: Nalassus (Helopondrus) gloriosus (Faldermann, 1837) and N. (Helopocerodes) faldermanni (Faldermann, 1837).
 
The fourteen new species of Rhaconotus Ruthe, 1854 from South Asia are described and illustrated: Rh. affinis sp. nov., Rh. chinensis sp. nov., Rh. fujianus sp. nov. , Rh. hei sp. nov., Rh. heterotrichus sp. nov., Rh. intermedius sp. nov., Rh. ipodoryctoides sp. nov., Rh. iterabilis sp. nov., Rh. luteosetosus sp. nov., Rh. magnus sp. nov., Rh. oriens sp. nov., Rh. tergalis sp. nov., Rh. tianmushanus sp. nov., and Rh. yaoae sp. nov. The following species are recorded first time for the fauna of China: Rh. aciculatus Ruthe, Rh. menippus Nixon, Rh. nadezhdae (Tobias et Belokobylskij), and Rh. zarudnyi Belokobylskij. The key for determination of Chinese species of Rhaconotus is provided.
 
First and last instar larva and pupa of Laccoptera foveolata (Boheman, 1856) are described in detail along with notes on the life history. The beetles were observed to breed from June to August. The host plant of the insect is Ipomoea nil (Convolvulaceae), as the entire life cycle is completed on this plant. A small, oval ootheca containing a single egg is partially covered by fecal matter. There are five larval instars; the larvae and pupa are typical for the specialized cassidines. Imagines develop in about 35-40 days from the fertilized egg deposited in the ootheca.
 
Three new Apophylia species A. pesai, A. halberstadti and A. dembickyi spp. nov.) are described from Indian provinces Meghalaya, Assam and Arunachal Pradesh. Apophylia vietnamica Samoderzhenkov, 1988, is recorded for the first time in India. Male genitalia are figured for all species.
 
A new species, Irantha nigrina, is described in the subfamily Harpactorinae based on the specimen from China. The dorsal habitus, head, prothorax and other diagnostic morphological features are illustrated. The genus is reported for the first time to China. The reviewed diagnostic characters of the genus are presented.
 
Anurida riverina sp. nov. (1) Dorsal chaetotaxy; (2) antennal segment III-IV dorsally; (3) postantennal organ and eyes; (4) mandible; (5) maxilla; (6) labial chaetotaxy; (7) chaetotaxy of abdominal sterna; (8) tibiotarsus III.
Anurida riverina sp. nov. from river floodplains of Poland is described. Notes on distribution and systematic relationship are given.
 
The genus Diolenius Thorell is revised to include 14 species, 9 of them new to science. Diolenius carinifer Strand is synonymised with D. armatissimus Thorell. A female of D. armatissimus is described for the first time. Diolenius bicinctus Simonis considered nomen dubium. Four species: D. albomaculatus Thorell, D. bifasciatus Thorell, D. venustus Thorell and D. vittatus Thorell hither to included in Diolenius are transferred to Ohilimia. For D. phrynoides a neotype specimen is designated. All species of Diolenius sensu stricto are diagnosed, described and illustrated. A key to the species of Diolenius and distributional maps are provided. The genus is closely related to Chalcolecta Simon, 1884 and Ohilimia Strand, 1911 and its range is restricted to rain forests of New Guinea, New Britain, The Moluccas and some adjacent islands.
 
The genus Pellenes is newly recorded from Australia. Its relationships are discussed and diagnostic drawings, redescription and distributional data for Pellenes bitaeniata (Keyserling, 1882), the only known Australian representative of the genus, are given.
 
The larva of Enemothrombium bifoliosum (Canestrini, 1884) is described for the first time. Postlarval instars are redescribed and the neotype is designated. Diagnoses of adults, deutonymphs and larvae for Enemothrombium Berlese, 1910 and Valgothrombium Willmann, 1940 are provided. Parafeiderium culicoidium Vercammen-Grandjean et Cochrane, 1974 is transferred to Enemothrombium. Parafeiderium stuarti Baker, 1999 is transferred to Valgothrombium. Parafeiderium Vercammen-Grandjean et Cochrane, 1974 is considered a junior synonym of Enemothrombium Berlese, 1910. Lacinitrombium Southcott, 1994 and Furcotrombium Southcott, 1994 are synonymized with Valgothrombium. As a result, four new combinations arose: Enemothrombium culicoidium (Vercammen-Grandjean et Cochrane, 1974), comb. nov., Valgothrombium fluminis (Michener, 1946), comb. nov., Valgothrombium spasiscutum (Robaux, 1974), comb. nov. and Valgothrombium stuarti (Baker, 1999), comb. nov. Data about phenology, life cycle, development and parasitism of E. bifoliosum are reported. The phylogenetic position of Enemothrombium within Microtrombidiidae is discussed.
 
Archaeorthoptera elongata Brongniart, 1893: 433 (specimen MNHN A26167; see text for abbreviations; all photographs taken under polarized light; (A, B, D) dry-ethanol composites; (C, E, F) ethanol; (B–E): details as located on (A). (A) Habitus; (B) head; (C; arrows indicate spines) left mid-leg femur; (D; wide arrows indicate edges of the 2 nd tarsomere) right mid-leg tarsus; (E) right foreleg tibia; (F; wide arrows indicate the anterior wing margin) area between the anterior wing margin and RA, left forewing base.  
The discovery of new specimens of the Late Carboniferous species elongata Brongniat, 1893: 433, belonging to the taxon Archaeorthoptera nom. Béthoux and Nel, 2002, dis.-typ. Béthoux, 2007c (i.e. is more closely related to the Linnaean order Orthoptera than to any other recent insect order) is reported. The head and leg morphology of the species could be investigated thanks to the good preservation of the specimens, and new preparation. The tarsus is 5-segmented, supporting the view that this state is plesiomorphic for orthopterans s. s. Morphology of mandibules indicates a carnivorous food-habit. Fore- and mid-leg femora and tibiae were provided with long and spaced out spines, most probably arranged in two rows. These spines might have assisted the entrapment of preys. The relative length of leg segments, and the length of antennae, are reminiscent of the condition exhibited by phasmidans, suggesting a similar locomotion behavior.
 
Distribution of Megaphyllum uncinatum sp. nov. (black dots).
Megaphyllum uncinatum sp. nov. is described from both Israel and Cyprus. It is assigned, together with M. iranicum Golovatch, 1983, from Iran, to the previously monotypic subgenus Persebrachyiulus Golovatch, 1983 of the large Eastern Euro-Mediterranean genus Megaphyllum Verhoeff, 1894. A refined diagnosis of Persebrachyiulus is provided. Some zoogeographical comments are also presented, likely implying several cases of anthropochoric introduction to Cyprus from the mainland Middle East and possibly even vice versa.
 
Ceutorhynchus varius Rey, 1895, formerly considered to be a variety of C . hirtulus Germar, 1824, is revealed to be a valid species and redescribed. It lives on Arabidopsis thaliana (L.) Heynh. (Brassicaceae) and is recorded from France, Poland, Czech Republic, Slovakia, Austria and Bulgaria. Lectotypes are designated for C. varius Rey and C. hirtulus Germar.
 
The genus Phylacinus Fairmaire, 1896 (type species: Phylacinus asperipennis Fairmaire, 1896) is revised and illustrated. New species is described: Phylacinus ferreri sp. nov. Key for species determination is provided. Lectotype and paralectotype are designated for Phylacinus asperipennis Fairmaire, 1896. The genus represents of the Malagasy endemic fauna.
 
The diplotaxine genus Ceratogonia Kolbe, 1899 is revised. Currently, it includes only the species C. bicornuta (Kolbe, 1899), which is redescribed and has its male genitalia figured for the first time. Ceratogonia marshalli (Arrow, 1902) and C. bilaminiceps (Benderitter, 1920) are considered junior subjective synonyms of C. bicornuta. The synonymy of C. kraatzi (Dalla Torre, 1912) with Dichecephala ovata (Fåhraeus in Boheman, 1857) is confirmed. Lectotype and paralectotype designations are provided for following species: Apogonia (Ceratogonia) bicornuta, A. (C.) marshalli and A. ovata. Neotype is designated for A. (C.) kolbei Kraatz, 1899.
 
A revision of the Asian species of the genus Hypodoryctes Kokujev, 1900 is provided. Seven new species are described and figured: H. cantata Belokobylskij et Chen sp. nov. (Russian Far East, Korea, Japan), H. fuga Belokobylskij et Chen sp. nov. (Russian Far East, Korea, Japan, China, Vietnam), H. rapsodia Belokobylskij sp. nov. (Azerbaijan), H. rondo Belokobylskij et Chen sp. nov. (Vietnam, China), H. serenada Belokobylskij et Chen sp. nov. (China), H. symphonia Belokobylskij sp. nov. (Vietnam), H. tango Belokobylskij et Chen sp. nov. (China). Redescriptions of H. sibiricus Kokujev, 1900 (Palaearctic, Myanmar, Mexico, Costa Rica), H. bilobus (Shestakov, 1940) (Russian Far East, Korea, Japan, China) and H. torridus Papp, 1987 (Russian Far East, Korea, Japan, China) are given. A key to all species of the genus Hypodoryctes is provided.
 
Protaphorura ionescui sp. nov. is described from Romania. This species is closest to Protaphorura glebata (Gisin, 1952) which is redescribed, and P. fimata (Gisin, 1952). A key for Protaphorura with dorsal pseudocellar formula like 33/022/33333 is given.
 
The results of experimental rearing of Neotrombicula inopinata and Leptotrombidium russicum and of field studies aiming at finding the hitherto unknown habitats occupied by active postlarval forms are presented. Diagnoses of deutonymphs reared from field-collected larvae of both species are provided. Literature interpretation of deutonymph of N. inopinata is inconsistent with the characteristics of deutonymph of N. inopinata obtained from larvae by experimental rearing. Larvae of L. russicum and L. silvaticum can be separated only on the base of host spectrum. Considering the biology of the parasite and host species, it is likely that postlarval forms of bat-parasitizing species may be confined to tree and cave habitats, whereas those species that are known as parasites of rodents inhabit the soil habitats.
 
Hyllus treleaveni Peckham et Peckham, 1902 is redescribed. H. moestus Peckham et Peckham, 1903, H. bevisi Lessert, 1925 and H. marleyi Lessert, 1925 are recognised as junior synonyms of H. treleaveni. The new distributional data are mapped. Data on the species' natural history and observations from captive specimens are also presented.
 
Four Palearctic species of the genus Microchelonus Szépligeti, 1908 with very small apical metasomal aperture in males are reviewed. Both the male and the female of Microchelonus minifossa Tobias, 1986 and M. luzhetzkji (Tobias, 1966) are redescribed. Males of two species (females unknown), M. vickae sp. nov. and M. gracitis sp. nov., are described from Central Europe (Czech Republic).
 
Ecdyonurus nigrescens (Klapálek, 1908), mature larva, Brebeneskul stream, Chornohora Range, Ukrainian Carpathians: (20) fourth gill; (21) first gill; (22) variability of central part of posterior margin of fifth abdominal tergum.
The neotype of Ecdyonurus nigrescens (Klapálek, 1908) has been designated, basing on male imago reared from larva, collected in the Chornohora Range (the Ukrainian Carpathians). The male imago of this species is redescribed; male subimago, female imago and subimago, larvae and the structure of egg chorion are described and illustrated for the first time. Critical diagnostic characters distinguishing this species from other representatives of the Ecdyonurus helveticus species-group are discussed. The type locality is defined in accordance with the Article 76.3 and Recommendation 76A of ICZN as follows Ukraine: Zakarpattia Region, Carpathian Biosphere Reserve, the Chornohora Range, Polonyna Brebeneska district, mountain valley between SW slope of Gutyn Tomnatek Mt. and NW slope of Brebeneskul Mt., upper section of the Brebeneskul stream (left-bank tributary of the Hoverla stream, Tysa river-basin), 1450 m. a.s.l. The data on the original syntype series and type locality are discussed in detail. All known specimens of E. nigrescens collected by J. Dziędzielewicz between 1908 and 1910 are observed and reidentified. The data on distribution, ecology and life cycle of E. nigrescens are presented.
 
Juvenile stages of Punctoribates punctum (C. L. Koch, 1839) and P. hexagonus Berlese, 1908 are redescribed, those of P. sellnicki Willmann, 1928 are described for the first time, and both juveniles and adults of all species are illustrated. The juvenile stages of these species are similar in body shape and the number of setae on the prodorsum, gastronotum and anal, anogenital and epimeral regions, but differ in the shape and length of some prodorsal and gastronotal setae. Keys to the larvae and nymphs of these species are prepared.
 
Allothrombium meridionale Berlese, 1910 is redescribed based on type material originating from Italy and specimens collected in Mosel Valley, Germany. Characteristics of larvae, deutonymphs and adults are provided. Data on habitat specificity and phenology of the species as well as on developmental time of larvae are given. Morphological abnormalities have been observed under laboratory conditions in larvae which hatched from eggs not exposed to lower temperatures, contrary to larvae that hatched from eggs after chilling. Larvae of Allothrombium triticium Zhang, 1995 sensu Goldarazena and Zhang (1997), reported from Spain, are considered to represent A. meridionale. An identification key to species of Allothrombium, including central European members of the genus known from adults and world species known from larvae is provided. Phanolophus oedipodarum (Frauenfeld, 1868), is new to the fauna of Germany.
 
The genus Ohilimia Strand is revised to include O. scutellata (Kritscher, 1959) and O. albomaculata (Thorell, 1881) comb. nov. Both species are diagnosed, described and illustrated. A key to the species and map of distribution are given. Diolenius venustus Thorell, 1881, Diolenius bifasciatus Thorell, 1881 and Discocnemius coccineopilosus Simon, 1884 are newly synonymised with Ohilimia albomaculata (Thorell, 1881). Diolenius vittatus Thorell, 1881 and Discocnemius albocingulatus Simon, 1884 are transferred to Ohilimia, but considered nomina dubia. The genus is closely related to Diolenius Thorell, 1870 and Chalcolecta Simon, 1884. Its range is restricted to rain forests of NE Cape York Peninsula in Australia, New Guinea and The Moluccas (Ternate, Kai).
 
This paper deals with eight species belonging to the genus Prionchulus Cobb, 1916. P. spectabilis (Ditlevsen, 1911) and P. longus (Thorne, 1929) are redescribed on the basis of the type material. Six new species: P. kralli sp. nov., P. pinophilus sp. nov., P. polonicus sp. nov., P. pseudolongus sp. nov. P. septentrionalis sp. nov., and P. thornei sp. nov.are described and illustrated.
 
Agroiconota paraguayana: (23) male, (24) profile; A. aulatipennis: (25) male, (26) profile; A. lateripunctata: (27) male, (28) female, (29) profile; A. stupidula: (30) immaculate male, (31) maculate female, (32) profile; A. conflagrata: (33) male, (34) female, (35) profile; A. carlobrivioi: (36) male, (37) female, (38) profile; A. subtriangularis: (39) male, (40) female, (41) profile; A. atromaculata: (42) male, (43) female, (44) profile. 
Agroiconota judaica: (45) areolate male, (46) subvittate female, (47) immaculate female, (48) profile; A. atropunctata: (49) male, (50) female, (51) profile; A. punctipennis: (52) male, (53) female, (54) profile; A. gibbipennis: (55) male, (56) female, (57) profile; A. sanareensis: (58) male, (59) female, (60) profile; A. parellina: (61) male, (62) female, (63) profile; A. inedita: (64) male, (65) female, (66) profile; A. vilis: (67) male, (68) female, (69) profile. 
Agriconota atromaculata (Peru), A. atropunctata (Bolivia), A. arlobrivioi (Bolivia, Brazil, Peru), A. gibbipennis (Brazil), A. paraguayana (Paraguay), and A. sanareensis (Venezuela), new to the science, are described. New records of several species, colour photos of all species, and a key to the genus Agroiconot Spaeth, 1913 are given.
 
The morphology of the larva and nymphs of Pilogalumna crassiclava (Berlese, 1914) and the larva of P. ornatula Grandjean, 1956 is described for the first time, the morphology of nymphs of the latter species is redescribed, and adults of both species are illustrated. The juvenile stages of P. crassiclava are similar to those of P. ornatula in body shape and the number of setae, but differ by several morphological characters, like the shape of gastronotal seta dp in the larva and prodorsal seta le in the nymphs, intensity of reticulation of gastronotal shield of nymphs and body size. The morphology of juvenile stages of these species is also compared to that of P. tenuiclava (Berlese, 1908), and the tritonymph and adult of this species are illustrated. Keys to the larvae and nymphs of all these species are presented.
 
Prionchulus hygrophilus sp. nov. (1) Female anterior region; (2) female; (3) male; (4-6) female tails. (1, 2, 4-holotype female; 3-(Popul. I); 5, 6-paratype females). Scale bars: 1 = 10 µm; 2, 3, = 200 µm; 4-6 = 30 µm.
Prionchulus hygrophilus sp. nov. and P. bogdanowiczi sp. nov. are described based on the material collected from Ukraine and Taiwan.
 
The member of the African Ectateus group ("platynotoid" lineage), monotypic genus Nesopatrum Gebien, 1920 (type species: Opatrinus josephi Karsch, 1881), is revised and illustrated. Nesopatrum Gebien is the first time recorded as a genus of the tribe Platynotini due to the structure of strydulatory gula and "platynotoid" type of aedeagus. The following new synonym is proposed: Anchophthalmus densaticollis Fairmaire, 1887 (=Phallocentrion praelacinatum Koch, 1956). Key to the genera of Ectateus group and distribution are provided.
 
Last instar larva of Aethiopocassis rhodesiana (Spaeth, 1924) is described in detail. It is the only detailed description of a larva of an African member of the tribe Cassidini.
 
Megaselia anticheira sp. nov. from Oman and Yemen, M. epandquadrata sp. nov. from Yemen, M. gouteuxi sp. nov. from Ivory Coast, Oman and Yemen and M. nussbaumi sp. nov. from Israel and Yemen are described. A neotype from Israel is designated to replace the lost holotype of M. deltomera (Schmitz, 1924) from Croatia. The first Old World females of this complex are described and keys to the six Old World species are provided.
 
Top-cited authors
Jacek Szwedo
  • University of Gdansk
Adam STROIńSKI
  • Polish Academy of Sciences
Marcin Jan Kamiński
  • Polish Academy of Sciences
Łukasz Kaczmarek
  • Adam Mickiewicz University, Poznan
Magdalena Gawlak
  • Institut of Plant Protection - NRI, Poland, Poznań