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The twenty districts or dzongkhag of the kingdom of Bhutan. West: Chhukha (Ch), Haa (Ha), Paro (Pa), Samtse (St), Thimphu (Th). Central: Dagana (Da), Gasa (Ga), Punakha (Pu), Tsirang (Ts), Wangdue Phodrang (WP). South: Bumthang (Bu), Sarpang (Sp), Trongsa (To), Zhemgang (Zh). East: Lhuntse (Lh), Mongar (Mo), Pema Gatshel (PG), Samdrup Jongkhar (SJ), Trashigang (Ta), Trashi Yangtse (TY).

The twenty districts or dzongkhag of the kingdom of Bhutan. West: Chhukha (Ch), Haa (Ha), Paro (Pa), Samtse (St), Thimphu (Th). Central: Dagana (Da), Gasa (Ga), Punakha (Pu), Tsirang (Ts), Wangdue Phodrang (WP). South: Bumthang (Bu), Sarpang (Sp), Trongsa (To), Zhemgang (Zh). East: Lhuntse (Lh), Mongar (Mo), Pema Gatshel (PG), Samdrup Jongkhar (SJ), Trashigang (Ta), Trashi Yangtse (TY).

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Article
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An annotated distributional checklist of the Coccinellidae of Bhutan is presented, enumerating 91 species, 17 of which are here recorded for the first time: Shirozuella tibetina Wang, Ge & Ren, 2012, Jauravia limbata (Motschulsky, 1858), Scymnus bourdilloni (Kapur, 1958), Illeis confusa Timberlake, 1943, Calvia albida Bielawski, 1972, Harmonia expa...

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Context 1
... natural vegetation includes subtropical broadleaved forests, temperate forests, conifer forests, dry grasslands, and alpine shrublands and meadows. Forest coverage is still extensive occupying about 70% of the country's surface area (National Soil Services Centre 2011). The country is divided into 20 administrative districts, called dzongkhag ( Fig. 1). ...
Context 2
... records taken from the literature, reference is given for the first occurrence only. Secondary sources without precise records, e.g. catalogues, are only included if no earlier publications could be traced. As far as possible, an effort was made to place the collecting localities in one of the twenty dzongkhag (districts) currently recognized (Fig. 1). Dzongkhag in bold face indicate new records. Species hitherto not known from outside the territory of Bhutan are considered endemic and labelled ...
Context 3
... & O. Vorst (CNR). . By far the largest number of species (42) has been reported from Chhukha dzongkhag in the southwest of the country, followed by Wangdue Phodrang (central) with 21 species and Thimphu (west) with 18 species (Table 1, Fig. 1). From two of the 20 dzongkhag, Gasa and Lhuntse, no coccinellids have been reported at all. It is certain that the large regional differences observed here are mainly artefacts caused by various levels of collecting effort made in different parts of the country. This is further corroborated by the fact that the western part of the ...
Context 4
... natural vegetation includes subtropical broadleaved forests, temperate forests, conifer forests, dry grasslands, and alpine shrublands and meadows. Forest coverage is still extensive occupying about 70% of the country's surface area (National Soil Services Centre 2011). The country is divided into 20 administrative districts, called dzongkhag ( Fig. 1). ...
Context 5
... records taken from the literature, reference is given for the first occurrence only. Secondary sources without precise records, e.g. catalogues, are only included if no earlier publications could be traced. As far as possible, an effort was made to place the collecting localities in one of the twenty dzongkhag (districts) currently recognized (Fig. 1). Dzongkhag in bold face indicate new records. Species hitherto not known from outside the territory of Bhutan are considered endemic and labelled ...
Context 6
... & O. Vorst (CNR). . By far the largest number of species (42) has been reported from Chhukha dzongkhag in the southwest of the country, followed by Wangdue Phodrang (central) with 21 species and Thimphu (west) with 18 species (Table 1, Fig. 1). From two of the 20 dzongkhag, Gasa and Lhuntse, no coccinellids have been reported at all. It is certain that the large regional differences observed here are mainly artefacts caused by various levels of collecting effort made in different parts of the country. This is further corroborated by the fact that the western part of the ...

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... Samples were observed and collected within a five-meter width on either side of the transect line, as well as within five-meters in front (Mayur et al., 2013). To ensure their preservation, all samples were initially stored in the field using 70-100% ethanol and were later identified using literature but not limited to Bielawski (1979) ;Poorani, (2002); Dorji et al (2017) ;Dorji, et al. (2019). Currently, all the specimens studied are stationed in the collection of College of Natural Resources, Royal University of Bhutan. ...
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... Samples were observed and collected within a five-meter width on either side of the transect line, as well as within five-meters in front (Mayur et al., 2013). To ensure their preservation, all samples were initially stored in the field using 70-100% ethanol and were later identified using literature but not limited to Bielawski (1979) ;Poorani, (2002); Dorji et al (2017) ;Dorji, et al. (2019). Currently, all the specimens studied are stationed in the collection of College of Natural Resources, Royal University of Bhutan. ...
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... Distribution: Cambodia (Kampong Speu and Kampot pro.), Australia, Bangladesh, Bhutan, China, India, Indonesia, Japan, Nepal, Sri Lanka and Thailand (Chunram & Sasaji, 1980;Poorani, 2003;Dorji et al., 2019). ...
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... Distribution: Cambodia (Kampong Speu pro.) NEW RECORD, Australia, Borneo, Bhutan, India, Indonesia, Philippines and Thailand (Chunram & Sasaji, 1980;Poorani, 2003;Dorji et al., 2019). Description: Body semi-oval. ...
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