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The twenty districts or dzongkhag of the kingdom of Bhutan. West: Chhukha (Ch), Haa (Ha), Paro (Pa), Samtse (St), Thimphu (Th). Central: Dagana (Da), Gasa (Ga), Punakha (Pu), Tsirang (Ts), Wangdue Phodrang (WP). South: Bumthang (Bu), Sarpang (Sp), Trongsa (To), Zhemgang (Zh). East: Lhuntse (Lh), Mongar (Mo), Pema Gatshel (PG), Samdrup Jongkhar (SJ), Trashigang (Ta), Trashi Yangtse (TY).

The twenty districts or dzongkhag of the kingdom of Bhutan. West: Chhukha (Ch), Haa (Ha), Paro (Pa), Samtse (St), Thimphu (Th). Central: Dagana (Da), Gasa (Ga), Punakha (Pu), Tsirang (Ts), Wangdue Phodrang (WP). South: Bumthang (Bu), Sarpang (Sp), Trongsa (To), Zhemgang (Zh). East: Lhuntse (Lh), Mongar (Mo), Pema Gatshel (PG), Samdrup Jongkhar (SJ), Trashigang (Ta), Trashi Yangtse (TY).

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Article
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An annotated distributional checklist of the Coccinellidae of Bhutan is presented, enumerating 91 species, 17 of which are here recorded for the first time: Shirozuella tibetina Wang, Ge & Ren, 2012, Jauravia limbata (Motschulsky, 1858), Scymnus bourdilloni (Kapur, 1958), Illeis confusa Timberlake, 1943, Calvia albida Bielawski, 1972, Harmonia expa...

Contexts in source publication

Context 1
... natural vegetation includes subtropical broadleaved forests, temperate forests, conifer forests, dry grasslands, and alpine shrublands and meadows. Forest coverage is still extensive occupying about 70% of the country's surface area (National Soil Services Centre 2011). The country is divided into 20 administrative districts, called dzongkhag ( Fig. 1). ...
Context 2
... records taken from the literature, reference is given for the first occurrence only. Secondary sources without precise records, e.g. catalogues, are only included if no earlier publications could be traced. As far as possible, an effort was made to place the collecting localities in one of the twenty dzongkhag (districts) currently recognized (Fig. 1). Dzongkhag in bold face indicate new records. Species hitherto not known from outside the territory of Bhutan are considered endemic and labelled ...
Context 3
... & O. Vorst (CNR). . By far the largest number of species (42) has been reported from Chhukha dzongkhag in the southwest of the country, followed by Wangdue Phodrang (central) with 21 species and Thimphu (west) with 18 species (Table 1, Fig. 1). From two of the 20 dzongkhag, Gasa and Lhuntse, no coccinellids have been reported at all. It is certain that the large regional differences observed here are mainly artefacts caused by various levels of collecting effort made in different parts of the country. This is further corroborated by the fact that the western part of the ...
Context 4
... natural vegetation includes subtropical broadleaved forests, temperate forests, conifer forests, dry grasslands, and alpine shrublands and meadows. Forest coverage is still extensive occupying about 70% of the country's surface area (National Soil Services Centre 2011). The country is divided into 20 administrative districts, called dzongkhag ( Fig. 1). ...
Context 5
... records taken from the literature, reference is given for the first occurrence only. Secondary sources without precise records, e.g. catalogues, are only included if no earlier publications could be traced. As far as possible, an effort was made to place the collecting localities in one of the twenty dzongkhag (districts) currently recognized (Fig. 1). Dzongkhag in bold face indicate new records. Species hitherto not known from outside the territory of Bhutan are considered endemic and labelled ...
Context 6
... & O. Vorst (CNR). . By far the largest number of species (42) has been reported from Chhukha dzongkhag in the southwest of the country, followed by Wangdue Phodrang (central) with 21 species and Thimphu (west) with 18 species (Table 1, Fig. 1). From two of the 20 dzongkhag, Gasa and Lhuntse, no coccinellids have been reported at all. It is certain that the large regional differences observed here are mainly artefacts caused by various levels of collecting effort made in different parts of the country. This is further corroborated by the fact that the western part of the ...

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... Ahmad et al., (2017) reported 9 species from district Sargodha, Pakistan. Dorji et al. (2019) reported 91 species of coccinellids and out of them 17 were recorded for the first time from Bhutan. Poorani (2002) have described 400 species of ladybird beetles from Indian subregion. ...
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