Male genitalia of Hypolycaena narada sp. nov. (paratype, NCBS-PY983). A : lateral view, with the aedeagus removed. B : right valva, lateral view from outside. C : ventral view of the fused valvae. D : aedeagus, lateral view. All separated parts of the male genitalia (B-D) are proportional to A. The scale bar is approximate. Dissection and images by Dipendra Nath Basu. 

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Context 1

... Voucher code NCBS-PY976. Fig. 1, Table 2. . The type locality is near the village of Bodhisatta, Changlang District, Arunachal Pradesh, NE India (N 27°33 ′ , E 96°23 ′ ). Collected by Krushnamegh Kunte, 14 March 2015. Preserved dry, pinned, deposited in the Research Collections Facility at the National Centre for Biological Sciences, Bengaluru (=Bangalore), India. Paratypes: (1) NCBS-PY977. Fig. 1, Table 2. ♂ . Collection data same as the holotype, except collected on 8 March 2015. (2) NCBS- PY978. Fig. 1, Table 2. ♂ . Collection data same as the holotype, except collected on 8 March 2015. (3) NCBS-PY979. Table 2. ♂ . Collection data same as the holotype. (4) NCBS-PY980. Table 2. ♂ . Collection data same as the holotype, except collected on 11 March 2015. (5) NCBS-PY981. Table 2. ♂ . Collection data same as the holotype, except collected on 11 March 2015. (6) NCBS-PY982. Table 2. ♂ . Collection data same as the holotype, except collected on 11 March 2015 and preserved in ethanol for molecular work. (7) NCBS-PY983. Fig. 2 ♂ genitalia, Table 2. ♂ . Collection data same as the holotype, except collected on 14 March 2015 and preserved in ethanol for molecular work. Description: Holotype: Forewing length 15mm. Upper side : Both wings dark and slightly shining purple-blue. Forewing border sooty-grey, narrow, just a thread at tornus, gradually dilating to 2mm at apex and continuing along the costa almost up to the base. This border sometimes appears pale grey-brown in a side light (Fig. 3B). A dark, diffused androconial discal patch, spanning from the base of v2 to v7 and from cell-end outward for approx. 4mm. Hindwing black tails, ending in white tips, at v1 (approx. 6mm long) and v2 (approx. 4mm long), wing margin wavy at v3 and v4. The abdominal fold and costal margin in space 8 dark grey-brown. Dispersed but prominent white scalation in the submarginal terminal area, which is condensed into submarginal white bars in spaces 1b, 1c and 2. A tornal, large, white-crowned black spot adjacent to the white bar in 1b. A black marginal line running the length of the termen, and cilia FW and HW pale grey. HW basal half and space 1c covered in ochreous, hair-like long scales, which are especially visible in a side light. Underside : Both wings pale grey, shining silvery-grey in bright light. Forewing cell-end bar defined on both sides by inner black and outer white lines. Discal band narrow, uneven but contiguous, ochreous in the middle, black- and then white-edged on both the sides, pointed and vanishing near v1, ending in a black spot at v10. A faint grey-brown, wavy, post-discal line from v1 to v6. Hindwing cell-end bars similar to the FW, but less well-marked. Two prominent, oblong, white-ringed black spots along the costal margin, of which the basal spot is before the cell-end bar, and the outer spot is beyond the discal band. The discal band narrow, ochreous in the middle, black- and then white-edged on both the sides, and highly broken. It is composed of an oblique line in 1b, a pair of disconnected oblique lines in 1c converging towards each other in the middle, and then a series of largely conjoined spots in spaces 2 to 6, which is dislocated at v4. A tornal, large, white-crowned black spot in 1b, lined towards v1 by sparse, iridescent blue scales. Another large, orange-crowned black spot in space 2. This is followed by grey-centred white rings in spaces 3 to 6, which are inwardly defined by dark grey. A black marginal line running the length of the termen of both wings, with cilia bright pale grey. Head, thorax and abdomen overall similar to other Hypolycaena . Frons white. Labial palps white at the base, black near the tip. Eyes black. Antennae black, narrowly ringed white; club black, tipped reddish similar to Hypolycaena othona Hewitson, [1865]. Proboscis ochreous. Thorax white below, purple or purple-brown above. Legs broadly ringed black and white. Abdomen dark brown to almost black above, white below. Paratypes, and intraspecific variation: All the paratypes are largely similar to the holotype, including in the overall colour pattern and forewing length (Fig. 1, Table 2). Variation is seen primarily in the nature of the discal bands on the underside, and the extent of white scalation near the terminal margin on the upper side of hindwing. In the holotype, the discal band on the forewing underside is straight and almost touches v1. This discal band is similarly straight and almost touching v1 in all the paratypes except in NCBS-PY977, in which the discal band is bent inwards at v3, and ends in the middle of space 1c. On the hindwing underside, the discal spot in 3 is not elongated in the holotype and all the paratypes, except in NCBS-PY977, in which it is elongated and points towards the wing base. In the holotype, there are dispersed but prominent white scales in the submarginal terminal area of the hindwing, which are condensed into submarginal white bars in spaces 1b, 1c and 2. Similar white scales and submarginal white bars are present in paratypes NCBS-PY980 and NCBS-PY981. In paratypes NCBS-PY977, NCBS- PY978 and NCBS-PY979, the dispersed white scales are nearly absent, and the submarginal bars in 1a, 1b and 2 are very narrow and therefore somewhat indistinct (this is especially the case in NCBS- PY978). Male genitalia: The male genitalia of the paratype (NCBS- PY983; Fig. 2) are of similar form but quite distinct from the male genitalia of H. kina celastroides Corbet, 1938 and H. tora ...

Context 2

... Voucher code NCBS-PY976. Fig. 1, Table 2. . The type locality is near the village of Bodhisatta, Changlang District, Arunachal Pradesh, NE India (N 27°33 ′ , E 96°23 ′ ). Collected by Krushnamegh Kunte, 14 March 2015. Preserved dry, pinned, deposited in the Research Collections Facility at the National Centre for Biological Sciences, Bengaluru (=Bangalore), India. Paratypes: (1) NCBS-PY977. Fig. 1, Table 2. ♂ . Collection data same as the holotype, except collected on 8 March 2015. (2) NCBS- PY978. Fig. 1, Table 2. ♂ . Collection data same as the holotype, except collected on 8 March 2015. (3) NCBS-PY979. Table 2. ♂ . Collection data same as the holotype. (4) NCBS-PY980. Table 2. ♂ . Collection data same as the holotype, except collected on 11 March 2015. (5) NCBS-PY981. Table 2. ♂ . Collection data same as the holotype, except collected on 11 March 2015. (6) NCBS-PY982. Table 2. ♂ . Collection data same as the holotype, except collected on 11 March 2015 and preserved in ethanol for molecular work. (7) NCBS-PY983. Fig. 2 ♂ genitalia, Table 2. ♂ . Collection data same as the holotype, except collected on 14 March 2015 and preserved in ethanol for molecular work. Description: Holotype: Forewing length 15mm. Upper side : Both wings dark and slightly shining purple-blue. Forewing border sooty-grey, narrow, just a thread at tornus, gradually dilating to 2mm at apex and continuing along the costa almost up to the base. This border sometimes appears pale grey-brown in a side light (Fig. 3B). A dark, diffused androconial discal patch, spanning from the base of v2 to v7 and from cell-end outward for approx. 4mm. Hindwing black tails, ending in white tips, at v1 (approx. 6mm long) and v2 (approx. 4mm long), wing margin wavy at v3 and v4. The abdominal fold and costal margin in space 8 dark grey-brown. Dispersed but prominent white scalation in the submarginal terminal area, which is condensed into submarginal white bars in spaces 1b, 1c and 2. A tornal, large, white-crowned black spot adjacent to the white bar in 1b. A black marginal line running the length of the termen, and cilia FW and HW pale grey. HW basal half and space 1c covered in ochreous, hair-like long scales, which are especially visible in a side light. Underside : Both wings pale grey, shining silvery-grey in bright light. Forewing cell-end bar defined on both sides by inner black and outer white lines. Discal band narrow, uneven but contiguous, ochreous in the middle, black- and then white-edged on both the sides, pointed and vanishing near v1, ending in a black spot at v10. A faint grey-brown, wavy, post-discal line from v1 to v6. Hindwing cell-end bars similar to the FW, but less well-marked. Two prominent, oblong, white-ringed black spots along the costal margin, of which the basal spot is before the cell-end bar, and the outer spot is beyond the discal band. The discal band narrow, ochreous in the middle, black- and then white-edged on both the sides, and highly broken. It is composed of an oblique line in 1b, a pair of disconnected oblique lines in 1c converging towards each other in the middle, and then a series of largely conjoined spots in spaces 2 to 6, which is dislocated at v4. A tornal, large, white-crowned black spot in 1b, lined towards v1 by sparse, iridescent blue scales. Another large, orange-crowned black spot in space 2. This is followed by grey-centred white rings in spaces 3 to 6, which are inwardly defined by dark grey. A black marginal line running the length of the termen of both wings, with cilia bright pale grey. Head, thorax and abdomen overall similar to other Hypolycaena . Frons white. Labial palps white at the base, black near the tip. Eyes black. Antennae black, narrowly ringed white; club black, tipped reddish similar to Hypolycaena othona Hewitson, [1865]. Proboscis ochreous. Thorax white below, purple or purple-brown above. Legs broadly ringed black and white. Abdomen dark brown to almost black above, white below. Paratypes, and intraspecific variation: All the paratypes are largely similar to the holotype, including in the overall colour pattern and forewing length (Fig. 1, Table 2). Variation is seen primarily in the nature of the discal bands on the underside, and the extent of white scalation near the terminal margin on the upper side of hindwing. In the holotype, the discal band on the forewing underside is straight and almost touches v1. This discal band is similarly straight and almost touching v1 in all the paratypes except in NCBS-PY977, in which the discal band is bent inwards at v3, and ends in the middle of space 1c. On the hindwing underside, the discal spot in 3 is not elongated in the holotype and all the paratypes, except in NCBS-PY977, in which it is elongated and points towards the wing base. In the holotype, there are dispersed but prominent white scales in the submarginal terminal area of the hindwing, which are condensed into submarginal white bars in spaces 1b, 1c and 2. Similar white scales and submarginal white bars are present in paratypes NCBS-PY980 and NCBS-PY981. In paratypes NCBS-PY977, NCBS- PY978 and NCBS-PY979, the dispersed white scales are nearly absent, and the submarginal bars in 1a, 1b and 2 are very narrow and therefore somewhat indistinct (this is especially the case in NCBS- PY978). Male genitalia: The male genitalia of the paratype (NCBS- PY983; Fig. 2) are of similar form but quite distinct from the male genitalia of H. kina celastroides Corbet, 1938 and H. tora ...