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Lygosoma quadrupes from Cat Tien National Park, Dong Nai Province, Southern Vietnam. Photograph: W. Van Devender.
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We review the supple skinks (Lygosoma Hardwicke & Gray, 1827) known to occur in Indochina (Cambodia, Laos and Vietnam). Here, the existence of seven species is documented: L. angeli, L. boehmei, L. bowringii, L. corpulentum, L. haroldyoungi, L. isodactylum (type of Squamicilia Mittleman, 1952), and L. quadrupes (type of Lygosoma). Based on morpholo...
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... in preservative. Dorsum light brown; dorsum, flanks, and tail with indistinct dark longitu- dinal lines along the edges of the scales; ventral sur- face cream; supralabials rather dark, sutures lighter. Smith (1935) reported that the juveniles bear a black- ish colouration with fine longitudinal golden lines. For colouration in life see Fig. 7. (Bobrov 1992(Bobrov , 1995Ziegler et al. 2006;Nguyen et al. 2009). Our records are new for Lam Dong and Tay Ninh provinces. For distribution in Indochina see Fig. 8F. Outside of Indochina, this species is known from Thailand, China, Western Malaysia, Indonesia, and the Philippines (Boulenger 1887;Smith 1935;Taylor 1963;Manthey & ...
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Citations
... However, relatively few reports document the diets of skinks, especially predation on other reptiles (Attum et al. 2004;Civantos et al. 2013;Patil et al. 2021). The semifossorial scincid genus Riopa is mainly associated with leaf litter, rocks, rotting logs, and other cover in semiarid regions, grasslands, and urban areas, where they are known to feed on arthropods, annelids, snails, and other small invertebrates (Geissler et al. 2011;Freitas et al. 2020;Bhilala et.al. 2021). ...
... Phân bố: Ở Việt Nam loài này được ghi nhận ở các tỉnh từ Hải Dương, Bắc Giang ở phía bắc, vào đến thành phố Hồ Chí Minh và Kiên Giang ở phía Nam [1], [16]. Trên thế giới, loài này ghi nhận ở Mi-an-ma, Campuchia, Lào, Thái Lan, Ma-lai-xi-a, Singapore, Indonexia, Băng-la-đét, Trung Quốc, Ấn Độ, Phi-líp-pin, Ô-xtrây-li-a [1]. ...
... Lygosoma bowringii (Günther, 1864) (Hình 1.a) Mẫu vật nghiên cứu (n = 2): QNU DC TL.39, 55 (♀) thu vào tháng 5/2021. Mẫu vật có đặc điểm hình thái phù hợp với mô tả của Vassilieva và cộng sự (2016)[14], Geissler và cộng sự (2011)[16] ...
Nghiên cứu này đánh giá mức độ đa dạng thành phần loài bò sát ở huyện Đức Cơ, tỉnh Gia Lai, một khu vực còn ít được biết đến về mặt đa dạng sinh học. Dựa vào kết quả khảo sát thực địa và phân tích đặc điểm hình thái mẫu vật tại huyện Đức Cơ trong năm 2020 và 2021, kết hợp với số liệu đã công bố trước đây, chúng tôi đã cập nhật danh sách thành phần loài bò sát của tỉnh Gia Lai là 117 loài thuộc 23 họ, 2 bộ. Đã ghi nhận bổ sung 06 loài bò sát cho tỉnh Gia Lai, bao gồm: 01 loài thằn lằn Lygosoma bowringii; 02 loài rắn Dendrelaphis subocularis, Lycodon subcinctus và 03 loài rùa Heosemys grandis, Siebenrockiella crassicollis, Indotestudo elongata. Về giá trị bảo tồn, ghi nhận 26 loài có giá trị bảo tồn (chiếm 22,2% tổng số loài bò sát của Tỉnh) bao gồm: 16 loài có tên trong Sách Đỏ Việt Nam (2007); 19 loài có tên trong Danh lục Đỏ thế giới (IUCN, 2023); 01 loài có tên trong Phụ lục I, Nghị định 64/2019/NĐ-CP của Chính phủ; 02 loài có tên trong nhóm IB và 12 loài có tên trong nhóm IIB, Nghị định 84/2021/NĐ-CP của Chính phủ.
... Though the species is known to occur in a wide range of habitats, including dipterocarp forests and agricultural lands, any encounter with a L. haroldyoungi seems to be a rather unusual event, even for local people or farmers. One such rare fi nding of a "Naga with legs" in Laos was reported in an ethnozoological remark by Geissler et al. (2011). Currently the species is only known to occur in Thailand and Laos. ...
... Based on this material, we were able to identify all four specimens as L. haroldyoungi on account of their body shape and characteristic colour patt ern. They match the holotype in the following diagnostic traits (Taylor, 1962;Grossmann, 2007;Geissler et Cambodian Journal of Natural History 2022 (2) 74-77 First records of Lygosoma haroldyoungi al., 2011): body extremely elongated, snake-like; limbs short, pentadactyle (can be seen in the specimen in Fig. 2); dorsal surface of head dark brown, sharply bordered by the light dorsum colouration behind the parietals; ground colouration of dorsum light yellowish-brown (the specimen depicted in Fig. 2 shows a rather greyishbrown ground colouration); dorsum bearing dark brown or blackish crossbands which are narrowing towards the fl anks, forming a reticulated patt ern (in the specimen depicted in Fig. 2 the anterior crossbands are widely fused, forming an irregular paravertebral band on the anterior part of trunk); original parts of tail bearing dark brown crossbands, alternating with thin bands in grey, whitish or bluish colouration; regenerated parts of tail dark brown, scatt ered with irregular light spots. Interestingly, one specimen from Kulen Promtep Wildlife Sanctuary (the larger specimen depicted in Fig. 2) is bearing a completely original, not regenerated tail. ...
Based on a total of four specimens found at two localities in central and northern Cambodia, Lygosoma haroldyoungi is reported for the first time in the country. These new records extend the known range of the species by approximately 350 km towards the southeast. Besides, the four modes of locomotion observed in these individuals are also described.
... The genus Lygosoma Hardwicke & Gray, 1827 is generally known as writhing skink or supple skink, which formerly contained more than 30 species distributed across Africa, India, Southeast Asia, the western and southern Philippines, as well as Christmas Island (Australia) (Cogger 2014;Datta-Roy et al. 2014;Geissler et al. 2011;Heitz et al. 2016;Siler et al. 2018). Recently, Freitas et al. (2019) revised the taxonomy of the genus Lygosoma sensu lato by using a molecular phylogeny, which indicated that Lygosoma sensu stricto is restricted to Southeast Asia only. ...
... The type species of Lygosoma, L. quadrupes (Linnaeus, 1766) was previously known as a widely distributed species, from Indochina, Thailand, southern China (including Hong Kong), Western Malaysia, Indonesia, and the Philippines (Geissler et al. 2011). Until recently, Siler et al. (2018) redescribed true Lygosoma quadrupes (Linnaeus, 1766) and restricted its geographic distribution to the island of Java in Indonesia, whereas the Indochina specimens were placed in a newly described species, namely L. siamense Siler, Heitz, Davis, Freitas, Aowphol, Termprayoon & Grismer, 2018 based on historical collections from Thailand. ...
... Thus, the solely mentioned specimen of L. quadrupes from China can hardly be traced. Although most authors also recorded the distribution of L. quadrupes with inclusion of "South China" as well as "Hongkong", seemingly directly followed Boulenger (1887), no further Chinese specimens were reported to our best knowledge (Geissler et al. 2011;Smith 1935;Taylor 1963). Most recently, Siler et al. (2018) restricted true L. quadrupes to Java, Indonesia only, and revised specimens from Thailand to their newly described species, i.e., L. siamense. ...
The genus Lygosoma Hardwicke & Gray, 1827 is a group of semi-fossorial supple skink from Southeast Asia, the members of which are still understudied in Indochina. Lysosoma quadrupes (Linnaeus, 1766) was once recorded from China based on historical literature; until recent its Indochina specimens were revised and placed into a newly described species, i.e., L. siamense Siler, Heitz, Davis, Freitas, Aowphol, Termprayoon & Grismer, 2018. However, Lygosoma quadrupes had long been unknown from China but the historical record had not been confirmed. Here we report a vouchered record for L. siamense from China for the first time based on four specimens collected in Sanya, Hainan, China. Furthermore, the taxonomic history of Lygosoma from China is also reviewed.
... Morphologically, species of Subdoluseps differ from other skinks by their small size (snout-vent length up to 70.0 mm), moderately elongate trunk, short limbs, 26-34 midbody scale rows, 50-69 paravertebral scale rows, lower eyelid scaly or with a transparent disc, supranasal scales in contact medially or not in contact medially, prefrontals not in contact medially, frontoparietal single or paired, parietals in contact behind interparietal, enlarged nuchal scales present or absent, palatine bones with posteriomedially projecting processes, and pterygoids rounded along posterior edge . Vietnam has a single species of Subdoluseps (S. bowringii), which occurs widely throughout the country (Nguyen et al. 2009;Geissler et al. 2011;Uetz et al. 2020). ...
... For comparison, all species in the Lygosoma s.l. in Asia were used. Morphological characters used for the comparison were taken from the literature Geissler et al. 2011;Geissler et al. 2012;Grismer et al. 2019;Heitz et al. 2016;Karin et al. 2018;Seetharamaraju et al. 2009;Sharma 1976;Sharma 1977;Siler et al. 2018;Smith 1916;Smith 1921;Smith 1935;Smith 1937;Taylor 1950;Taylor 1962;Werner 1910;and Ziegler et al. 2007). Further, additional morphological data of the two closest species, S. bowringii and S. frontoparietale, were obtained from specimens in the Institute of Tropical Biology Collection of Zoology (ITBCZ), Ho Chi Minh City, Vietnam; the Field Museum (FMNH), Chicago, USA; and Zoological Museum of Kasetsart University (ZMKU), Thailand (Appendix 1). ...
... The new species differs from S. bowringii by having a single frontoparietal scale (vs. paired) (Geissler et al. 2011;Smith 1935; this study), tail length approximately equal to SVL (vs. tail longer than SVL) (mean TL/SVL = 1.04 vs. 1.27 [Smith 1935; this study]), and fewer subcaudal scales (64-74 vs. 76-81) (this study); from S. frontoparietale by having a tail length approximately equal to SVL (mean TL/SVL = 1.04 vs. tail longer than SVL, mean TL/SVL = 1.24 [Taylor 1962; this study, Table 4]), fewer subcaudal scales (64-74 vs. 78-80) [Taylor 1962; this study]), fewer axilla-groin scales (34-37 vs. 39-42) (this study), four distinct longitudinal black stripes on dorsum (vs. ...
We describe a new species of the genus Subdoluseps Freitas, Datta-Roy, Karanth, Grismer & Siler from a coastal area in southern Vietnam. Subdoluseps vietnamensis sp. nov. is characterized by the following morphological characters: medium size in adults (snout-vent length up to 48.7 mm); tail length/snout-vent length ratio 1.04; toes not reaching finger when limbs adpressed; 27–30 midbody scale rows, smooth; 55–57 paravertebral scales; 55–62 ventral scale rows; 64–74 subcaudal scales; frontoparietal scale single; four supraoculars; prefrontals not in contact with one another; two loreal scales; seven supralabials; ear-opening with two lobules on the anterior border; smooth lamellae beneath finger III 9 or 10 and toe IV 12–15; six enlarged precloacal scales; and four distinct black stripes on dorsum. The new species differs genetically from its closest congeners, S. bowringii (Günther) and S. frontoparietale (Taylor), by uncorrected p-distances of 10.0% and 9.5%, respectively in ND1 sequences, and clusters into the same matriline with these two congeners on the phylogenetic trees.
... Nach Pauwels et al. (2008) zählt dieser Laubskink zu den seltensten und am schlechtesten untersuchten Arten der Herpetofauna Thailands. Selbst in der umfassenden Arbeit über die Gattung Lygosoma in Indochina (Geissler et al. 2011) ist relativ wenig über die Art zu finden. Fehlende Lebendabbildungen in der Literatur bis in jüngere Zeit (z. ...
... Im Nordosten Thailands ist die Dämmerung nur sehr kurz, also wurde das aktive Exemplar während der Dunkelheit gefangen. Weiterhin übersahen sie und Geissler et al. (2011) Hinweise auf eine gewisse Gewandtheit der Art im Klettern, da der Holptypus erfolgreich aus dem ihm zugedachten Behältnis ausbrechen konnte (Taylor 1962). Anonymus (2011) berichtet, dass der Skink in Gefangenschaft an heißen Tagen gerne im Wasser schwimmt. ...
In the following the history of discovery of the taxon and its mention in the herpetological literature with data on distribution and detailed information on the biotope of a specimen
can be made. On the basis of different specimens, notes on pholidosis, drawing and colouring made by Harold Young's Laubskink. About the way of life of the species in the terrarium is reported, whereby experience of almost eighteen years of care of a specimen is referred to.
Key words: Squamata: Sauria: Scincidae: Lygosominae: Lygosoma haroldyoungi (Taylor, 1962): natural habitat; lepidosis; colouration and pattern; captive care; ethology; modes of locomotion; Southeast Asia: Thailand, Laos.
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... We chose meristic and mensural characters based on Heitz et al. (2016) and Geissler et al. (2011): snout-vent length (SVL: distance from tip of snout to vent), axilla-groin distance (AGD: distance between posterior edge of forelimb insertion and anterior edge of hind-limb insertion), total length (TotL: distance from tip of snout to tip of tail), midbody width (MBW: measured from lateral surface to opposing lateral edge at midpoint of axilla-groin region), midbody depth (MBD: measured from ventral surface to dorsal surface at midpoint of axilla-groin region), tail length (TL: measured from posterior margin of vent to tip of tail), tail width (TW: measured at widest section of tail posterior to hemipene bulge), tail depth (TD: measured from ventral to dorsal surface of tail at the same point as TW), head length (HL: measured from tip of snout to posterior margin of retroarticular process), head width (HW: widest measure of head width at jaw articulations), head depth (HD: measured from ventral to dorsal surface of head at jaw articulations), eye diameter (ED: at widest point), eye-narial distance (END: from anterior margin of eye to posterior margin of nares), snout length (SNL: from anterior margin of eye to tip showing known provincial distribution of Lygosoma siamensis based on vouchered museum specimens (black and grey provinces), with focal provincial names shown for reference. Close-up view of provinces in grey shown within box A. (Bottom left) Hypothesized relationships among species of the Lygosoma quadrupes complex illustrated by the maximum clade credibility tree resulting from Bayesian analyses of 16S. ...
... Boulenger, 1887:340;de Rooij, 1915:225;Taylor, 1922:233. Lygosoma quadrupes Cochran, 1930:16;Smith, 1935Smith, :290, 1937 Bobrov, 1995:16;Manthey and Grossmann, 1997:267;Cox et al., 1998:115;Gaulke, 1999:273;Honda et al., 2000:453;Bauer and Gü nther, 2006:17;Ziegler et al., 2007:399;Wagner et al., 2009:2;Chan-ard et al., 2010:132;Das, 2010:240;Geissler et al., 2011Geissler et al., :1169Geissler et al., , 2012Pyron et al., 2013:14;Datta-Roy et al., 2014:165;Heitz et al., 2016:352. ...
... Interestingly, many recent discoveries and descriptions of scincid diversity throughout the region have been of smaller and more secretive species, most often semifossorial or leaf-litter specialists (Grismer et al., 2009(Grismer et al., , 2014(Grismer et al., , 2017Grismer, 2011). In fact, across Indochina, numerous taxonomic problems pertaining to burrowing genera continue to present (e.g., Isopachys, Larutia, Leptoseps, Lygosoma; Geissler et al., 2011Geissler et al., , 2012Grismer, 2011;Linkem et al., 2011;Datta-Roy et al., 2014). Semifossorial lizards in general represent an ecomorph class of reptiles that remains poorly understood on our planet, largely attributable to the logistical difficulty in locating and studying populations in the wild (Siler et al., 2011(Siler et al., , 2012Davis et al., 2014;Grismer et al., 2016Grismer et al., , 2017. ...
Based on molecular and morphological data sets, we describe a new species of scincid lizard of the genus Lygosoma from Indochina, and redescribe true Lygosoma quadrupes (Linnaeus, 1766). The new species is small and slender, and represents the third member of the L. quadrupes complex, increasing the diversity of Lygosoma species recognized in Southeast Asia to 24. Based on the reevaluation of vouchered specimens from the type locality of L. quadrupes sensu Linnaeus (1766), the recognized geographic distribution of true L. quadrupes is restricted to the island of Java in Indonesia. With 10 species of Lygosoma recognized in Thailand, the country possesses considerable species-level diversity of these enigmatic, semifossorial skinks. In addition to being one of the smallest species in the genus, the new species can be distinguished from all congeners by features of its external morphology, including having small relative limb lengths, longer trunk length, and greater numbers of axilla-groin and paravertebral scale rows. Phylogenetic analyses support three divergent lineages corresponding to recognized and newly described members of the L. quadrupes complex. The descriptions underscore the need for continued and comprehensive biodiversity survey work throughout much of Southeast Asia, particularly in Indochina, where scincid diversity remains poorly understood.
... Lygosoma albopunctatum is one of only a few skinks to have a wide distribution from peninsular India to Indochina into South East Asia (Jerdon 1853;Boulenger 1887Boulenger , 1890Smith 1935;Das 2010 [part], Geissler et al. 2011). An editorial footnote mentioned its occurrence in Mergui in Myanmar near the Siamese Peninsula (Jerdon 1853 Later, Boulenger (1887Boulenger ( , 1890 and Smith (1935) mentioned the distributional range of L. albopunctatum to be southern and central India, Bengal, Assam, Burma and the Malay peninsula, specifiying precise localities such as Madras, Denkanikottah Hills [near Hosur, Tamil Nadu, India], lower Godavary Valley, Bilaspur, Bengal and Penang. ...
The White-spotted Skink Lygosoma albopunctatum, a rarely recorded lizard, has been re-sighted from its type locality: Madras in the Coromandel Coastal Plains. Morphological details and field notes on the findings are elaborated. Since many past surveys in and around Madras, and in southern India generally, did not record this species, lack of consensus about its existence in southern India had developed, leading to published misconceptions about its distribution. These are highlighted and corrected herein. The species is also illustrated in life based on topotypical examples.
... The genus Lygosoma has a complicated taxonomic history, beginning with its original description of L. quadrupes (Linnaeus 1766), and including recent debate over whether they belong to Lygosoma vs. the genus Riopa (Geissler et al. 2011). The validity of the latter assignment, and what genera occur in Southeast Asia, remain open questions (Greer 1977;Moravec and Böhme 2008;Pauwels et al. 2008;Nguyẽn et al. 2009;Das 2010;Geissler et al. 2011). ...
... The genus Lygosoma has a complicated taxonomic history, beginning with its original description of L. quadrupes (Linnaeus 1766), and including recent debate over whether they belong to Lygosoma vs. the genus Riopa (Geissler et al. 2011). The validity of the latter assignment, and what genera occur in Southeast Asia, remain open questions (Greer 1977;Moravec and Böhme 2008;Pauwels et al. 2008;Nguyẽn et al. 2009;Das 2010;Geissler et al. 2011). Currently, 29 species of Lygosoma are recognized, with 17 present in Southeast Asia (Geissler et al. 2011;Uetz and Hošek 2016). ...
... The validity of the latter assignment, and what genera occur in Southeast Asia, remain open questions (Greer 1977;Moravec and Böhme 2008;Pauwels et al. 2008;Nguyẽn et al. 2009;Das 2010;Geissler et al. 2011). Currently, 29 species of Lygosoma are recognized, with 17 present in Southeast Asia (Geissler et al. 2011;Uetz and Hošek 2016). Several species possess widespread distributions spanning recognized faunal demarcations (Geissler et al. 2011). ...
The Palawan Pleistocene Aggregate Island Complex, a western Philippines island group, possesses a complex geologic history and numerous endemic vertebrates. On the basis of multiple unique morphological features, bolstered by genetic data supporting divergent lineages, we describe the first Philippine endemic species of Lygosoma from Palawan and Cuyo islands. This new semifossorial species is phenotypically similar to Lygosoma quadrupes, a widespread Southeast Asian scincid lizard. We recognize the Palawan lineage as a new species, the second member of the L. quadrupes species complex, and comment on its ecology and conservation status. This new species increases species-level diversity in the genus to 31, including 19 species distributed across Southeast Asia.
... 属(时磊, 个人通讯), 因此阿赖山裂睑蜥更名为阿 赖山泛蜥(Ablepharus alaicus)。 变更4属。原Eumeces属被拆分, 分布于东亚(包 括 中 国 ) 与 北 美 的 "Eumeces" 和 Neoseps 合 并 为 Plestiodon属 (Griffith et al., 2000;Schmitz et al., 2004;Brandley et al., 2005Brandley et al., , 2011Brandley et al., , 2012Pyron et al., 2013), 该属继续沿用中文属名"石龙子属", 属下种 中文名不变; 原Riopa属东南亚的一支全部归入蝘 蜓属(Lygosoma) (Geissler et al., 2011(Geissler et al., , 2012Pyron et al., 2013), 原中文种名"侏蜥"不变; 原Mabuya属亚 洲一支全部归入Eutropis 属 (Ota & Huang, 2000;Mausfeld et al., 2002;Pyron et al., 2013), 继续沿用 中文属名"南蜥属", 属下种中文名不变; 蛇蜥科中, 《中国动物志》 中脆蛇属(Ophisaurus)变动为脆蛇蜥 属(Dopasia) (Conrad et al., 2011;Nguyen et al., 2011;Pyron et al., 2013) (Ota et al., 1995;Rösler et al., 2011;Yang et al., 2012)、海南壁虎(G. similignum) (Ota et al., 1995;Rösler et al., 2011) 、 文 县 壁 虎 (G. ...
... 属(时磊, 个人通讯), 因此阿赖山裂睑蜥更名为阿 赖山泛蜥(Ablepharus alaicus)。 变更4属。原Eumeces属被拆分, 分布于东亚(包 括 中 国 ) 与 北 美 的 "Eumeces" 和 Neoseps 合 并 为 Plestiodon属 (Griffith et al., 2000;Schmitz et al., 2004;Brandley et al., 2005Brandley et al., , 2011Brandley et al., , 2012Pyron et al., 2013), 该属继续沿用中文属名"石龙子属", 属下种 中文名不变; 原Riopa属东南亚的一支全部归入蝘 蜓属(Lygosoma) (Geissler et al., 2011(Geissler et al., , 2012Pyron et al., 2013), 原中文种名"侏蜥"不变; 原Mabuya属亚 洲一支全部归入Eutropis 属 (Ota & Huang, 2000;Mausfeld et al., 2002;Pyron et al., 2013), 继续沿用 中文属名"南蜥属", 属下种中文名不变; 蛇蜥科中, 《中国动物志》 中脆蛇属(Ophisaurus)变动为脆蛇蜥 属(Dopasia) (Conrad et al., 2011;Nguyen et al., 2011;Pyron et al., 2013) (Ota et al., 1995;Rösler et al., 2011;Yang et al., 2012)、海南壁虎(G. similignum) (Ota et al., 1995;Rösler et al., 2011) 、 文 县 壁 虎 (G. ...
... .,地龟科Geoemydidae龟科 Emydidae水龟亚科 BatagurinaeShaffer et al., 1997; 张孟闻等, 1998; Krenz et al., 2005;Zhang et al., 2008; Barley et al., 2010; Thomson & Shaffer, 2010; Dijk et al., 2014 球趾虎科 SphaerodactylidaeGamble et al., 2008Gamble et al., , 2011Gamble et al., , 2012 Pyron et al., 2013 Xenodermatidae Vidal et al., 2007Wiens et al., 2008;Zaher et al., 2009; Pyron et al., 2011 Pyron et al., , 2013 Wallach et al., 2014 //www.biodiversity-science.net/CN/article/downloadArticleFile.do?attachType=PDF&id=10021 附录 3 属、种级分类阶元调整 Taxonomic adjustment for genera and speciesFritz & Havaš, 2007;Fritz et al., 2010; 周婷 和李丕鹏, 2013; Le et al., 2014 Le et al., 唐业忠, 1997Yang P et al., 2011; 周婷和李 丕鹏, 2013; Le et al., 2014 Le et al., 周工健等, 1991 陈合格等, 2006;Fritz et al., 2010; 周婷和李丕鹏, 2013; Le et al., Iverson et al., 1989; Barth et al., 2002 Barth et al., , 2004Feldman & Parham, 2004; Shen et al., 2013Griffith et al., 2000 Hikida et al., 2001; Schmitz et al., 2004; Brandley et al., 2005 Brandley et al., , 2011 Brandley et al., , 2012 Pyron et al., 2013Geissler et al., 2011 Pyron et al., 2013 Ota & Huang, 2000 Mausfeld et al., 2002; Pyron et al., 2013 Arnold, 1997 Lin et al., 2002; Lue & Lin, 2008; 向高世等, 2009; Pyron et al., 2013 鹿野草蜥 蔡波, 王跃招, 陈跃英, 李家堂. 中国爬行纲动物分类厘定. ...
