(10) Tolania sp., late-instar nymph; (11) Nodonica bispinigera, holotype male, left lateral view. (12) Nicomia sp., metathoracic leg; (13-14) Euwalkeria rubrica: (13) mesothoracic leg; (14) metathoracic leg; (15-16) male abdomen, dorsal view: (15) Nicomia buccina, holotype male, genital capsule removed; (16) Holdgatiella chiloensis, paratype male; (17) Nodonica bispinigera, holotype, aedeagus, lateral view. (cu) Cucullate setal row; (pp) posterior pronotal process, (sp) trochanter spine, (tp) tergal pits.

(10) Tolania sp., late-instar nymph; (11) Nodonica bispinigera, holotype male, left lateral view. (12) Nicomia sp., metathoracic leg; (13-14) Euwalkeria rubrica: (13) mesothoracic leg; (14) metathoracic leg; (15-16) male abdomen, dorsal view: (15) Nicomia buccina, holotype male, genital capsule removed; (16) Holdgatiella chiloensis, paratype male; (17) Nodonica bispinigera, holotype, aedeagus, lateral view. (cu) Cucullate setal row; (pp) posterior pronotal process, (sp) trochanter spine, (tp) tergal pits.

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Morphological characters of adults of the treehopper subfamily Nicomiinae Haupt, 1929 (Hemiptera, Membracidae) including seven genera (Eudonica gen. nov.; Euwalkeria Goding, 1926; Holdgatiella Evans, 1962; Nicomia Stål, 1858; Nodonica Dietrich, McKamey& Deitz, 2001; Stalomia gen. nov.; and Tolania Stål, 1858) and 22 species (16 new) are described a...

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Context 1
... these species, unlike many membracids, are solitary as adults. Tolania apparently lack parental care (DIETRICH et al. 2001), another common behavior in Membracidae. The nymphs are known only for Tolania and Nodonica (DIETRICH et al. 2001). Tolania nymphs are unusual in being relatively active in comparison to most membracid nymphs. Tolania nymphs (Fig. 10) have been observed running along the branches of their host plants with the tips of their abdomens held erect (Dietrich, ...
Context 2
... setae on the metathoracic tibia, and the male abdomen ovoid in dorsal view with the genital capsule retracted into the abdomen. (Fig. 15); furcasternum with anterior lobes or anterior and posterior lobes (Fig. 78) (Fig. 7); pronotum yellow anteriorly, pale green posteriorly with dorsal black macula; aedeagus with pair of dorsal, serrate flanges (Fig. 106) (Fig. 7); second valvulae with dorsal margin more or less evenly arcuate throughout (Fig. 122) .......................... N. pulchella Albertson, sp. ...
Context 3
... without anterior carina dorsally; lat- eral plate (Fig. 99) length more than half length of subgenital plate, slender, more or less evenly tapered, apex rounded; stern- ite IX anterior margin convex; subgenital plate lobes incom- pletely fused, with sub-basal constriction, apical two-thirds with lateral margins parallel, uniformly sclerotized without distinct fenestra. Aedeagus (Figs 100-101) posterior arm straight; shaft tubular; with sub-apical triangular process dorsally; apex nar- row, processes absent; gonopore on ventral preapical surface. Connective anterior margin straight; posterior apex truncate. ...
Context 4
... anterior margin straight; posterior apex truncate. Style (Figs 100-101) apodeme with apex curved slightly medi- ally; shank straight; apex acute. ...
Context 5
... with short, laterally directed anterior lobe and large, broad posterior lobe with apical process directed laterally. Abdomen. Terga with conspicuous pits; sternite III without medial keel. Male. Sternite VIII posterior margin with weak keeled projec- tion. Pygofer without anterior carina dorsally, with conspicu- ous pits; lateral plate robust (Fig. 102) Nicomia serrata Albertson, sp. sp. sp. sp. sp. nov nov nov nov nov. . . . . Figs 74-75, 79, 105-107, 123-124 Type locality. 18.1 km E Campinas field station, km 60 N Manaus, Amazonas, Brazil ...
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... spine I large and recurved. Furcasternum with short anterior lobes. Abdomen. Terga with conspicuous pits; sternite III with weak medial keel; pygofer with conspicuous pits dorsally. Male. Sternite VIII with anterior ca- rina; posterior margin with well developed keeled projection. Pygofer with anterior carina dorsally (Fig. 75); lateral plate ( Fig. 105) short, digitiform, strongly curved medially, only just vis- ible in dorsal view; sternite IX anterior margin emarginate me- dially; subgenital plate lobes completely fused, with sub-basal constriction, apex broad and rounded, with densely sclerotized lateral ridge (with setae) and median basal fenestra (glabrous). Aedeagus (Figs ...
Context 7
... with anterior carina dorsally (Fig. 75); lateral plate ( Fig. 105) short, digitiform, strongly curved medially, only just vis- ible in dorsal view; sternite IX anterior margin emarginate me- dially; subgenital plate lobes completely fused, with sub-basal constriction, apex broad and rounded, with densely sclerotized lateral ridge (with setae) and median basal fenestra (glabrous). Aedeagus (Figs 106-107) posterior arm straight; shaft tubular; apex narrow, processes absent; two serrate flanges extending lengthwise along dorsal shaft; gonopore on ventral preapical surface. Connective anterior margin strongly emarginate, api- ces diverging; posterior apex truncate. ...
Context 8
... anterior margin strongly emarginate, api- ces diverging; posterior apex truncate. Style (Figs 106-107) apodeme short, extended anterolaterad; shank elongate, apex bent and blade-like. Female. ...
Context 9
... The name 'serrata' is Latin for "serrate" and refers to the serrate dorsal flange on the aedeagus. Pygofer without anterior carina dorsally; lateral plate digitiform (Fig. 108); sternite IX anterior margin straight; subgenital plate lobes incompletely fused, with subbasal constriction, apical two- thirds tapering, uniformly sclerotized without distinct fenestra. Aedeagus (Figs 109-110) posterior arm straight; shaft broad in lateral view and strongly compressed in ventral view; apex nar- row, processes absent; ...
Context 10
... without anterior carina dorsally; lateral plate digitiform (Fig. 108); sternite IX anterior margin straight; subgenital plate lobes incompletely fused, with subbasal constriction, apical two- thirds tapering, uniformly sclerotized without distinct fenestra. Aedeagus (Figs 109-110) posterior arm straight; shaft broad in lateral view and strongly compressed in ventral view; apex nar- row, processes absent; serrate flange surrounding apex and ex- tending partially along distal shaft; gonopore on ventral son Research NIC-0623) [UFPC]; Santa Catarina: Coleção Cam- pos Seabra, December 1953, A. Maller (Albertson Research NIC- 0622 female) [UFPC]; Santa Catarina: Mafra, December 1930, A. Maller (Albertson Research NIC-0349 female) [NCSU]; 1 fe- male [unknown South American locality] (Albertson Research NIC-0347 female) [NCSU]. Distribution. ...

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... Relationships among the treehoppers were mostly consistent and well supported between analyses, with the exception of the relationships among the three treehopper families. Although members of Melizoderidae (represented here by Llanquihuea pilosa) have been considered 'primitive' treehoppers and have been recovered as sister to Aetalionidae + Membracidae or Membracidae in previous studies (Dietrich & Dietz, 1993;Dietrich, 2002;Cryan & Urban, 2012), all of our analyses placed Llanquihuea sister to Holdgatiella, another endemic Chilean genus, currently placed in the membracid subfamily Nicomiinae (Albertson & Dietrich, 2005). Several of our analyses placed this clade within Membracidae with high support. ...
Article
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... Studies of male genitalia of United States species (Caldwell 1949, Dennis 1952, Kopp and Yonke 1979 has had mixed results for separating taxa with best results among males. Kopp and Yonke (1979) and Caldwell (1949) showed male genitalia a useful taxonomic tool in the tribe Ceresini, Albertson and Dietrich (2004) in the subfamily Nicomiinae, and Albertson and Dietrich (2006) in the genus Tolania. Dennis (1952) illustrated both male and female genitalia of 74 species from Wisconsin with usefulness among some species but uniqueness for general species identification was not shown. ...
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... Reviews of life-history data on solitary, aggregation, and subsocial behavior, ant-attendance, and host-plant associations (Wood 1984, Lin et al. 2004, Wallace and Deitz 2004, Albertson and Dietrich 2005, Godoy et al. 2006, Wallace 2011 underscore the need to fill gaps in our knowledge of the life histories of many membracid taxa. Behavioral observations (Table 19.2) are limited for most taxa, and we found no definitive data on the Centronodinae (Fig. 19.6a), ...
... Summary of available data on life-history patterns in the Membracidae(Wood 1984;Lin et al. 2004;Wallace and Deitz 2004;Albertson and Dietrich 2005;Godoy et al. 2006;Wallace 2011; O. Evangelista, personal communication on Heteronotinae). ...
Chapter
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... nov. will run to Euwalkeria in the generic key of Nicomiinae of Albertson & Dietrich (2005). They superficially resemble representatives of Euwalkeria in the reticulation of the forewings, but are easily distinguished by the male genitalia. ...
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... Recent revisions of the membracid subfamilies Stegaspidinae (Cryan and Deitz 1999), Centrotinae (Wallace and Deitz 2004), and Nicomiinae (Albertson and Dietrich 2005, 2006) have greatly facilitated identification of genera and species in these diverse, plesiomorphic treehopper lineages. These revisions have increased the numbers of known species in various groups by as much as 80% (Albertson and Dietrich 2005, 2006), indicating that the extant treehopper fauna is considerably more speciose than indicated by the number of described species, which now stands at approximately 3,200. ...
... Recent revisions of the membracid subfamilies Stegaspidinae (Cryan and Deitz 1999), Centrotinae (Wallace and Deitz 2004), and Nicomiinae (Albertson and Dietrich 2005, 2006) have greatly facilitated identification of genera and species in these diverse, plesiomorphic treehopper lineages. These revisions have increased the numbers of known species in various groups by as much as 80% (Albertson and Dietrich 2005, 2006), indicating that the extant treehopper fauna is considerably more speciose than indicated by the number of described species, which now stands at approximately 3,200. Further improvements in knowledge of the Neotropical treehopper fauna are needed to help resolve relationships among the major membracid lineages which, despite several recent morphology-and DNA sequence-based phylogenetic analyses (Dietrich and Deitz 1993, Dietrich et al. 2001, Cryan et al. 2000, 2003, Lin et al. 2004), remain poorly understood. ...
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The new Neotropical treehopper genera Smergotomia, based on S. clairae, new species, from Ecuador, and Braxtonota, based on B. enigmata, new species, from Puerto Rico are described and illustrated. Despite lacking a posterior pronotal process, Smergotomia appears to be most closely related to Smerdalea Fowler based on the forewing venation and male genitalia, but the subfamily and tribal placement of these two genera is uncertain. Braxtonota appears to be related to the two membracid tribes that are endemic to the Caribbean, but lacks the extra hind femoral cucullate setae diagnostic for Monobelini and the large teeth on the second valvulae diagnostic for Nessorhinini. It is provisionally placed in Monobe- lini. Smerdalea veracruzensis, new species from Mexico, with a pronotum considerably less ornate than its congeners, is also described and illustrated and a key to species of Smerdalea is provided.