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Most important (in my view) is my idea that mutation rate is influenced by genetic heterozygosity. If true, sections of classical population genetic theory would need revisiting. One finding is that African humans have diverged more from our common ancestor than non-Africans. I also study tuberculosis susceptibility in British cattle, microsatellite evolution, the role of heterozygosity in animal fitness, human population history and marine mammal breeding behaviour and population structure.

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