Publications

  • Saad HOWLADAR, Yassin AL-SODANY, Kadry ABDEL KHALIK
    Bothalia - African Biodiversity and Conservation 01/2015; 1. · 0.41 Impact Factor
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    Samia Heneidak, Kadry Abdel Khalik
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    ABSTRACT: To evaluate their diagnostic value in systematic studies, seed coat morphology for 16 taxa from 11 genera of Cucurbitaceae were examined using stereomicroscopy and scanning electron microscopy. The taxa included representatives of the tribes Benincaseae, Bryonieae, Coniandreae, and Luffeae in order to evaluate their diagnostic value in systematic studies. Macro- and micromorphological characters of their seeds are presented, including shape, color, size, surface, epidermal cell shape, anticlinal boundaries, and periclinal cell wall. The taxonomic and phylogenetic implications of seed coat micromorphology were compared with those of the available gross morphological and molecular data. Seed character analysis offered useful data for evaluating the taxonomy of Cucurbitaceae on both intrageneric and tribal levels. Monophyly of the tribes Bryonieae, Coniandreae, and Luffeae was supported. Moreover, these analyses supported previous biochemical and phylogenetic data, indicating that distinct lineages are present within the tribe Benincaseae, that this tribe is not monophyletic, and that the subtribe Benincasinae is highly polyphyletic. A key is provided for identifying the investigated taxa based on seed characters.
    Acta Botanica Brasilica 01/2015; 29(1):129-142. DOI:10.1590/0102-33062014abb3705 · 0.55 Impact Factor
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    Nasr M. Hassan, Kadry N. Abdel Khalik
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    ABSTRACT: Seed morphology of 29 taxa including 9 subgenera belonging to the genus Veronica was compared using scanning electron microscopy to assess their diagnostic value for systematic studies. Subgenus Beccabunga is the largest in this study represented by 9 taxa. Seed surface often varies from ridged reticulate or verrucose/granulate reticulate to rugose-reticulate. Seed color and size have limited taxonomic significance as their variation is uninformative. Thereagainst, seed shape and seed coat ornamentation present informative characters that can be used efficiently in distinguishing the studied taxa. Our seed features support the DNA sequence data in showing close relationships between V. biloba and V. campylopoda, and between V. fruticans and V. fruticulosa. The present study indicates that V. anagallis-aquatica, V. anagalloides, V. comosa, and V. catenata have more or less the same seed features, consequently, these four former species are most likely representing a single species as reported by earlier studies. Furthermore, V. polita and V. persica are very similar in terms of seed characteristics. Our results justify placement of V. peregina within subgenus Beccabunga and support the monophyly of the subgenus Veronica.
    Journal of Systematics and Evolution 03/2014; 52(2):215–230. DOI:10.1111/jse.12054 · 1.65 Impact Factor
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    Kadry Abdel Khalik, Magdy Abd El-Twab, Rasha Galal
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    ABSTRACT: Genetic diversity and phylogenetic analyses of 24 species, representing nine sections of the genus Galium (Rubiaceae), have been made using the Inter Simple Sequence Repeats (ISSR), Randomly Amplified Polymorphic DNA (RAPD), and combined ISSR and RAPD markers. Four ISSR primers and three RAPD primers generated 250 polymorphic amplified fragments. The results of this study showed that the level of genetic variation in Galium is relatively high. RAPD markers revealed a higher level of polymorphism (158 bands) than ISSR (92 bands). Clustering of genotypes within groups was not similar when RAPD and ISSR derived dendrograms were compared. Six clades can be recognized within Galium, which mostly corroborate, but also partly contradict, traditional groupings. UPGMA-based dendrogram showed a close relationship between members of section Leiogalium with G. verum and G. humifusum (sect. Galium), and G. angustifolium (sect. Lophogalium). Principal coordinated analysis, however, showed some minor differences with UPGMA-based dendrograms. The more apomorphic groups of Galium form the section Leiogalium clade including the perennial sections Galium, Lophogalium, Jubogalium, Hylaea and Leptogalium as well as the annual section Kolgyda. The remaining taxa of Galium are monophyletic.
    Biologia 03/2014; 69(3):300-310. DOI:10.2478/s11756-013-0314-z · 0.51 Impact Factor
  • Kadry Abdel Khalik
    Acta Botanica Brasilica 01/2014; 28(4). · 0.55 Impact Factor
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    Kadry Abdel Khalik, Iman Hussien Al Gohary
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    ABSTRACT: The seed morphology and seed proteins of 11 Vicia taxa from Egypt were studied. Macro- and micro-morphological characters, including seed shape, color, size, hilum shape and seed sculpture were described and analyzed. The secondary sculpture of the cell wall varies from papillose in nearly all Vicia species to lophate in V. hirsuta. Further, the seed proteins of studied taxa were investigated by SDS-PAGE and 39 different bands were scored. The similarity analysis based on the SDS-PAGE profile and seed morphology was found to be a useful characteristic for the discrimination of Vicia species both on the subgeneric and the sectional levels. The present study did not acknowledge V. tetrasperma and V. hirsuta as subgenus Ervum. As a result, division of Vicia into two subgenera i.e. Vicia and Cracca, is supported.
    Acta Scientiarum Biological Sciences 12/2013; 35(4):603-611. DOI:10.4025/actascibiolsci.v35i4.19345
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    Kadry ABDEL KHALIK a, Mohamed EL-SHEIKH c, Abeer EL-AIDAROUS a
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    ABSTRACT: Wadi Noman in Mecca is one of the most important wadis. It was included in the most important water sources where the spring and wells of Zobida were running. It was providing the holy places in Mecca with drinking water for visitors of Kaaba and Arafat regions. The present study provides an analysis of floristic composition, vegetation types as well as structure and species distribution of 20 sites, emphases on the environmental factors that affect the species distribution. A total of 126 species representing 39 families of the vascular plants are recorded. Fabaceae, Poaceae and Boraginaceae are the largest families. Therophytes and chamaephytes are the most frequent, indicating a typical desert life form spectrum. Floristic composition in the different geomorphologic landscape units showed differences in species richness. The highest species richness of (23 ssp. stand-1) is recorded in the wadi bed. The lowest species richness value (18 ssp. stand-1) is recorded in the wadi plateau and fissures. Chorological analysis revealed that 52% of the studied species are biregional, being native to the Saharo Arabian-Sudano Zambezian region. Four vegetation groups were identified after application of TWINISPAN, DCA and CCA programs and named after the characteristic species as follows: (I) 2 Aristolochia bracteolata-Cucumis prophetarum; (II) Calotropis procera-Acacia hamulosa-Caralluma russeliana; (III) Acacia abyssinica-Acacia hamulosa-Tephrosia desertorum and (IV) Argemone ochroleuca-Senna italica. These plants associations and speciation of Wadi Noman demonstrate significant variation in pH, EC, soil mineral contents and human impacts.
    Turkish Journal of Botany 01/2013; DOI:10.3906/bot-1209-56 · 1.60 Impact Factor
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    Kadry Abdel Khalik
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    ABSTRACT: Seed coat morphology of 15 species of Ipomoea L. was examined comparatively using 2 scanning and light microscopy methods in order to evaluate their diagnostic value for 3 systematic studies. Macro- micro morphological characters, including seed shape, color, 4 size, seed surface, epidermal cell shape, anticlinal boundaries, and periclinal cell wall 5 are presented. Descriptions of seed size, shape, color, surface and seed coat types are 6 summarized for the genus. Taxonomic phylogenetic implications of the seed coat micro 7 morphology are also discussed, in comparison with the available gross morphological 8 and molecular data. Results of the seed character analyses offer useful data evaluating 9 the taxonomy of Ipomoea both on the subgeneric and sectional level. Monophyly of 10 both section Erpipomoea Choisy and Eriospermum Hallier is not supported. A key for 11 the identification of the investigated taxa based on seed characters is provided.
    Turkish Journal of Botany 01/2013; 37(5). DOI:10.3906/bot-1302-56 · 1.60 Impact Factor
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    Abdel Kadry, Khalik, Gamal Osman, Waeil Al-Amoudi
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    ABSTRACT: To elucidate the genetic diversity of ten Ipomoea species, seed proteins were analyzed by SDS-PAGE and RAPD analysis. According to SDS-PAGE analysis, in total 51 different bands were identified across the studied species. The number of bands varies from 16 bands in Ipomoea cairica, to 3 bands in I. sinensis. The similarity analysis based on the SDS-PAGE profile turned out to be a useful character for the inequity of Ipomoea species both on the subgeneric and sectional level. Analysis of RAPD-PCR of DNA provided more precise information concerning relationships between Ipomoea sections than SDS-PAGE analysis. A remarkable result from this study was identifying a close relationship between Ipomoea purpurea of section Pharbitis (subgenus Ipomoea) and species of the subgenus Quamoclit. Further support comes from the molecular data of RAPD which indicate that Ipomoea cairica should be considered a well separated section which may be related to section Orthipomoea. Our results suggest that section Erpipomoea is not a monophyletic group, whereas species of section Orthipomoea form a single monophyletic section.
    Australian Journal of Crop Science 06/2012; 6:1088-1093. · 1.63 Impact Factor
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    K. Abdel Khalik, N. M. S. Hassan
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    ABSTRACT: Seed and trichome morphology of 13 taxa representing the genus Fagonia in Egypt have been studied using light (LM) and scanning electron microscopy (SEM). Macro- and micro-morphological characters, including seed shape, seed size, seed coat sculpture, trichome density and structure were studied to evaluate their systematic significance. Seed shape and trichome density were found to have only minor taxonomic value, but seed coat sculpturing and trichome structure proved to be valuable characters. These characters offer evidence to combine certain species, e.g. Fagonia kassasii and F. schimperi in F. bruguieri, Fagonia microphylla in F. scabra and Fagonia thebaica in F. arabica. However, F. sinaica, F. tristis, F. boveana and F. isotricha should be maintained as separate species. Generally, the different patterns of seed and trichome morphology are useful in delimitation of species within the genus Fagonia, but they could not be used to characterize groups of related species.
    Nordic Journal of Botany 02/2012; 30(1). DOI:10.1111/j.1756-1051.2011.01112.x · 0.60 Impact Factor
  • Kadry Abdel Khalik
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    ABSTRACT: To elucidate the genetic diversity of ten Ipomoea species, seed proteins were analyzed by SDS-PAGE and RAPD analysis. According to SDS-PAGE analysis, in total 51 different bands were identified across the studied species. The number of bands varies from 16 bands in Ipomoea cairica, to 3 bands in I. sinensis. The similarity analysis based on the SDS-PAGE profile turned out to be a useful character for the inequity of Ipomoea species both on the subgeneric and sectional level. Analysis of RAPD-PCR of DNA provided more precise information concerning relationships between Ipomoea sections than SDS-PAGE analysis. A remarkable result from this study was identifying a close relationship between Ipomoea purpurea of section Pharbitis (subgenus Ipomoea) and species of the subgenus Quamoclit. Further support comes from the molecular data of RAPD which indicate that Ipomoea cairica should be considered a well separated section which may be related to section Orthipomoea. Our results suggest that section Erpipomoea is not a monophyletic group, whereas species of section Orthipomoea form a single monophyletic section.
    Australian Journal of Crop Science 01/2012; 6(6):1088-1093. · 1.63 Impact Factor
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    K. Abdel Khalik and N. M. S. Hassan
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    ABSTRACT: Seed and trichome morphology of 13 taxa representing the genus Fagonia in Egypt have been studied using light (LM) and scanning electron microscopy (SEM). Macro- and micro-morphological characters, including seed shape, seed size, seed coat sculpture, trichome density and structure were studied to evaluate their systematic signifi cance. Seed shape and trichome density were found to have only minor taxonomic value, but seed coat sculpturing and trichome structure proved to be valuable characters. Th ese characters off er evidence to combine certain species, e.g. Fagonia kassasii and F. schimperi in F. bruguieri , Fagonia microphylla in F. scabra and Fagonia thebaica in F. arabica . However, F. sinaica , F. tristis, F. boveana and F. isotricha should be maintained as separate species. Generally, the diff erent patterns of seed and trichome morphology are useful in delimitation of species within the genus Fagonia , but they could not be used to characterize groups of related species.
    Nordic Journal of Botany 01/2012; 30:116-126. · 0.84 Impact Factor
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    Sami Rabei & Kadry Abdel Khalik
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    ABSTRACT: Three keys to 44 species belonging to the family Convolvulaceae in the flora of Egypt are prepared to facilitate the identification of those species, based on 72 characters. These keys were constructed using the DELTA key-generating programs. The first key was built using all characters and the second key was built using 56 characters (vegetative, floral and fruit characters). The first key is considered as main key and can be confirmed by the second key in the field. The third one was built using 16 characters (pollen grains, seed and cell wall characters) which are to be used to identify and confirm the specimen in the laboratory
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    Kadry N. Abdel khalik
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    ABSTRACT: A numerical taxonomic study of the family Zygophyllaceae from Egypt). A systematic study of 29 taxa belonging to 7 genera of subfamilies Balanitoideae, Zygophylloideae, Peganoideae, Tribuloideae, Seetzenioideae and Tetradiclidoideae of Zygophyllaceae from Egypt was carried out by means of numerical analysis based on sixty-one morphological characters, including vegetative parts, pollen grains and seeds. On the basis of UPGMA clustering and PCO analysis, six main groups are recognized. Representatives of these groups are clustered together based on characters with high factor loadings in the PCO analysis. Th e results indicate that Zygophyllaceae are heterogeneous, including Peganum harmala which has been proposed to belong in a separate family. Zygophylloideae, Tribuloideae, Tetradiclidoideae and Seetzenioideae are the most homogeneous groups. Sections and groups in both Tribulus and Fagonia seem artifi cial.
    Acta Botanica Brasilica 01/2012; 26(1):165-180. DOI:10.1590/S0102-33062012000100017 · 0.55 Impact Factor
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    A. A. El-Khatib, K. N. abdel-Khalek, Naglaa Usef
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    ABSTRACT: atmospheric pollution biomonitoring anatomy
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    Kadry Abdel Khalik Assoc. Prof. Dr
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    ABSTRACT: Pollen morphology of nine species belonging to four genera: Cephalaria, Pterocephalus, Scabiosa, and Lomelosia of Dipsacaceae in Egypt has been investigated by light microscope (LM) and scanning electron microscopy (SEM). Pollen shape, size, exine ornamentation, number and characters of apertures present powerful characters for distinguishing between species. The pollen grains were tricolpate and triporate. Their shapes vary from spheroidal to oblate-spheroidal. Regarding the position of apertures, three types can be recognized viz., circumaperturate, angulaperturate, and planaperturate. Concerning sculpturing of the exine in proximal face, two different pollen types can be distinguished viz., spinulate and gemmate types. The exine ornamentation was found useful to distinguish between closely related genera such as Scabiosa and Lomelosia. A diagnostic key is given for all studied taxa based on palynomorphological characters (© 2010 WILEY-VCH Verlag GmbH & Co. KGaA, Weinheim)
    Feddes Repertorium 09/2010; 121(3‐4):97 - 111. DOI:10.1002/fedr.201000004
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    kadry N. abdel Khalik
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    ABSTRACT: The seed morphology of nine taxa of Juncus from Egypt has been investigated using light and scanning electron microscopy, to determine the importance of seed coat features as taxonomic characters. Macro- and micromorphological characters, including seed shape, color, size, seed appendages, epidermal cell shape, anticlinal boundaries, and outer periclinal cell wall and secondary cell wall sculpture are presented. Four types of seed appendages are recognized: (i) seeds with two appendages; (ii) seeds without appendages; (iii) seeds with minutely a piculate at one end; and (iv) seeds with minutely a piculate at both ends. Two types of anticlinal cell wall boundaries, (i) raised-channeled, straight and (ii) raised, straight or sinuous, and three different shapes of outer periclinal cell wall are described: (i) flat; (ii) concave; and (iii) flat to slightly concave. The secondary sculpture of the cell wall varies from striate to microreticulate or reticulate, and smooth to finely folded. Seed characters provide useful data for formulating the taxonomy of Juncus both on the subgeneric and sectional level. A key for the identification of the investigated taxa based on seed characters is provided
    Journal of Systematics and Evolution 05/2010; 48(3):215-223. DOI:10.1111/j.1759-6831.2010.00075.x · 1.85 Impact Factor
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    Kadry N Abdel Khalik, Monier M Abd, El -Ghani, Ahmed El Kordy
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    ABSTRACT: On the basis of fifty morphological characters, including vegetative parts, flowers, fruits, seeds, pollen grains, and anatomical structure, a systematic study of 13 taxa belonging to genus Galium (Rubiaceae) from Egypt was conducted by means of numerical analysis. Four branches and clusters were distinguished. Representatives of these groups were clustered together according to characters with high factor loading in the principal coordinates analysis. The results showed congruence between the UPGMA clustering and principal coordinates analysis in suggesting four groups. There was some degree of similarity among the species of sect. Aparine (Kolgyda). The results indicated also that the sect. Leiogalium (G. mollugo) was a separate group, while Aparine (Kolgyda) was the most heterogeneous one.
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    ABSTRACT: Vegetation and soil were sampled in 74 stands representing habitat in the different phytogeographical regions of Egypt where Convolvulus species were recorded. Two Way Indicator Species Analysis (TWINSPAN) classified the vegetation into six groups. Each of them occupied a distinct type of habitats. Convolvulus species are assigned as indicator species for most of these groups. However, none of these species can be considered as a leading dominant in the surveyed stands. Certain species may exert a local dominance/co-dominance, or may be distinctly more important in a group of stands. Ordination of stands based on cover-abundance value of species and effects of edaphic factors on the spatial distribution of plants were investigated in the biplot of Detrended Canonical Correspondence Analysis (DCCA). Distribution of Convolvulus species was found to be highly correlated with edaphic factors. Among them, CaCO3, moisture content, SO4 and soil texture have significant effect on the distribution pattern.An 74 Standorten in verschiedenen phytogeographischen Regionen Ägyptens, von denen Convolvulus-Arten bekannt waren, wurden Vegetation und Boden beprobt. Mittels TWINSPAN (Two Way Indicator Species Analysis) wurde die Vegetation in sechs Gruppen klassifiziert. Jede von ihnen besetzt einen bestimmten Habitattyp. Convolvulus-Arten erwiesen sich für die meisten dieser Gruppen als Indikatoren. Jedoch kann keine dieser Arten für die untersuchten Habitate als führende Dominante angesehen werden. Manche Arten können als lokale Dominante/Ko-Dominante auftreten, oder sie sind in einer Gruppe von Standorten von großer Bedeutung. Die Position der Pflanzenstandorte basiert auf der Artendichte und der Rolle der edaphischen Faktoren in der rumlichen Verteilung der Pflanzen und wurde mittels DCCA (Detrended Canonical Correspondence Analysis) untersucht. Es wurde festgestellt, daß die Verteilung der Convolvulus-Arten sehr eng mit den edaphischen Faktoren korreliert ist. Von letzteren haben CaCO3, Feuchtegehalt, SO4 und Bodentextur den größten Einfluß auf das Verteilungsmuster.
    Feddes Repertorium 01/2008; 109(1‐2):175 - 185. DOI:10.1002/fedr.19981090119
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    Kadry Abdel Khalik
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    ABSTRACT: A systematic study of 44 taxa belonging to 10 genera of tribes Convolvuleae, Cuscuteae, Cresseae, Dichondreae, Ipomoeeae and Merremieae of the Convolvulaceae from Egypt was carried out by means of a phenetic analysis based on seventy one morphological characters, including vegetative parts, pollen grains and seeds. Based on UPGMA clustering and PCO analysis, five main groups are recognized: Convolvuleae, Cuscuteae, Cresseae, Dichondreae and a mixed group from Ipomoeeae and Merremieae. OUT’s of these groups are clustered together based on characters with high factor loading >0.60 in the PCO analysis. Our results indicate that the tribe Convolvuleae is heterogeneous and the Cuscuteae is homogeneous, and there is some degree of similarity between taxa of Merremieae and Ipomoeeae, and those Cresseae and Dichondreae.

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