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    ABSTRACT: The following new species of geotrupids and scarabs from Costa Rica and Panama are described: Athyreus gulesseriani new species, Ateuchus alutacius new species (the first recorded brachypterous Ateuchus species), Coprophanaeus geph-yra new species, Deltochilum acanthus new species, and Onthophagus turgidus new species. The following two species of Coprophanaeus are revalidated: C. kohlmanni Arnaud and C. uhleri Malý & Pokorný. Illustrations of the dorsal habitus of the new species are provided, as well as distribution maps for all species.
    Zootaxa 02/2012; 3193:28-52.
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    ABSTRACT: A historic analysis of the classification of Scarabaeoidea is presented. The analysis comprises a historic comparison of the type and number of characters of the genera and their grouping into higher-level taxa. The analysis is divided into three time periods, including a review of 70 publications on scarab systematics (beginning with Linnaeus). The first period (1735–1858) encompasses an era prior to the publication of Darwin's The Origin of Species, and it is, therefore, a classification process arguably free of any evolutionary influences. The second period (1859–1949) includes a comparative synthesis on the classification of the Scarabaeoidea based on faunistic and taxonomic works. The third period (1950–2006) summarizes and analyzes classifications influenced by phylogenetic theories and is based on monographs, faunal studies, keys for regional fauna, papers on comparative morphology, and studies specifically devoted to the understanding of evolutionary relationships and processes in the Scarabaeoidea. A large problem concerning the classification and phylogeny still remains because most studies do not consider all the diversity of the Scarabaeoidea in a single analysis.
    The Coleopterists Bulletin 01/2009; DOI:10.1649/0010-065X(2006)60[19:HOSC]2.0.CO;2
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    ABSTRACT: EARTH University is a small agronomic university with a theme of sustainability in eastern Costa Rica. Several natural and constructed wetlands on its campus are used for research, water quality improvement, and higher education. It has become an important location for research and teaching on humid tropical wetland ecology and management. A 112-ha flow-through Raphia taedigera (Arecaceae) forested wetland is being used for climate change research, focusing on carbon sequestration and methane generation. Methane emissions are measured seasonally and are comparable to rates in tropical wetlands published elsewhere. Carbon sequestration by the wetland appears to be substantially higher than similar flow-through temperate zone wetlands. Treatment wetlands are used on campus to improve water quality of effluents from an animal farm, a dairy plant, a landfill, and a banana paper plant. Water quality was substantially improved in all of these wetlands except the landfill leachate wetland. All of these campus wetlands have been integrated into the four-year education program of EARTH University and 22 undergraduate projects have been completed on wetlands over the past 14 years.
    Ecological Engineering 11/2008; 34(4-34):276-288. DOI:10.1016/j.ecoleng.2008.07.012

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